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Grullo (pronounced GREW-yo) or grulla is a
color Color (or colour in English in the Commonwealth of Nations, Commonwealth English; American and British English spelling differences#-our, -or, see spelling differences) is the visual perception based on the electromagnetic spectrum. Though co ...
of
horse The horse (''Equus ferus caballus'') is a domesticated, one-toed, hoofed mammal. It belongs to the taxonomic family Equidae and is one of two extant subspecies of ''Equus ferus''. The horse has evolved over the past 45 to 55 mi ...
s in the dun family, characterized by tan-gray or mouse-colored hairs on the body, often with shoulder and dorsal stripes and black barring on the lower legs. The genotype for grulla horses is a black base with dun dilution. In this coloration, each individual hair is mouse-colored, unlike a roan, which is composed of a mixture of dark and light hairs. The several shades of grulla are informally referred to with a variety of terms, including black dun, blue dun, slate grulla, silver grulla or light grulla, silver dun, or lobo dun. Silver grulla may also refer to a grulla horse with silver dapple, regardless of shade. In terms of
equine coat color genetics Equine coat color genetics determine a horse's coat color. Many colors are possible, but all variations are produced by changes in only a few genes. bay horse, Bay is the most common color of horse, followed by black and chestnut. A change at the ...
, all of these shades are based on the
dun gene The dun gene is a dilution gene that affects both red and black pigments in the equine coat color, coat color of a horse. The dun gene lightens most of the body while leaving the mane (horse), mane, tail, legs, and primitive markings the shade of ...
acting as a dilution gene over the
black Black is a color that results from the absence or complete absorption of visible light. It is an achromatic color, without chroma, like white and grey. It is often used symbolically or figuratively to represent darkness.Eva Heller, ''P ...
gene. Because the grulla color is not due to the
gray Grey (more frequent in British English) or gray (more frequent in American English) is an intermediate color between black and white. It is a neutral or achromatic color, meaning that it has no chroma. It is the color of a cloud-covered s ...
gene, a grulla horse remains the same basic color from birth, though some minor shade variation may occur from summer to winter coats. If a grulla also carries the gray gene, it will be born a mouse tan-gray shade, usually with bold primitive markings, but then lighten and eventually develop a white hair coat with age. Because black is less common in general than bay or chestnut, grulla is likewise less common than red duns or bay (classic or zebra) duns. For example, only 0.7% of quarter horses registered each year with the
AQHA The American Quarter Horse Association (AQHA), based in Amarillo, Texas, United States, is an international organization dedicated to the preservation, improvement and record-keeping of the American Quarter Horse. The association sanctions many ...
are grulla. The most obvious ways to tell whether a horse is grulla are not only the gray or tan-gray body color, but also its
primitive markings Primitive markings are a group of equine coat color, hair coat markings and qualities seen in several equine species, including horses, donkey, donkeys, and asses. In horses, they are associated with primitive breeds, though not limited to such ...
, which include some or all of the following: dark face, cobwebbing around the eyes and forehead, dark mottling on the body, leg barring (sometimes called tiger striping), dark ear tips and edging, dark ear barring, dark shadowing of the neck, dark dorsal and transverse striping, and light guard hairs bordering a dark mane and tail.


Terminology

The word "grulla" in Spanish means crane. (The original Spanish noun is pronounced in American Spanish and in
Peninsular Spanish Peninsular Spanish (), also known as the Spanish of Spain (), European Spanish (), or Iberian Spanish (), is the set of varieties of the Spanish language spoken in Peninsular Spain. This construct is often framed in opposition to varieties from ...
.) Though the Spanish word for "crane" is always "grulla" regardless of gender, some people call male horses "grullo" and female horses "grulla", including in Spanish. Grulla is also called mouse dun or blue dun. In the Norwegian Fjord horse, the coat is called ''grĂ¥'' (meaning "gray").


Distribution

The
tarpan The tarpan (''Equus ferus ferus'') was a free-ranging horse population of the Eurasian steppe from the 18th to the 20th century. What qualifies as a tarpan is subject to debate; it is unclear whether tarpans were genuine wild horses, feral domest ...
(''Equus ferus ferus'') was a relative of the domestic horse that became
extinct Extinction is the termination of an organism by the death of its Endling, last member. A taxon may become Functional extinction, functionally extinct before the death of its last member if it loses the capacity to Reproduction, reproduce and ...
in the nineteenth century, and which appears to have had grulla coloration. The tarpan has been considered a true
wild horse The wild horse (''Equus ferus'') is a species of the genus Equus (genus), ''Equus'', which includes as subspecies the modern domestication of the horse, domesticated horse (''Equus ferus caballus'') as well as the Endangered species, endangered ...
, an undomesticated relative or ancestor of the domestic horse. However, some authorities in the early twentieth century held the opinion that most equines called tarpans were actually domestic or
feral horse A feral horse is a free-roaming horse of domesticated stock. As such, a feral horse is not a wild animal in the sense of an animal without domesticated ancestors. However, some populations of feral horses are managed as wildlife, and thes ...
s, not a separate species. Several breeds with the grulla color have been developed in efforts to recreate (" breed back") the tarpan. These breeds include the Heck horse and
Konik The Konik or Polish Konik, , is a Polish list of horse breeds, breed of small horse or pony. There are semi-feral populations in some regions. They are usually dun gene, mouse dun or primitive markings, striped dun. The Bilgoray, , of south-e ...
. One of the first experiments in this regard was published in 1906 by James Cossar Ewart, who obtained a "tarpan-like" horse by crossing a Shetland mare and a black Welsh pony.


References


Sources and external links




Grulla Color Genetics and Photos
* * http://www.equusite.com/articles/basics/colors/colorsGrullo.shtml
Dun Zygosity test
''from'' Veterinary Genetics Laboratory, School of Veterinary Medicine, University of California, Davis. Web site accessed January 13, 2008.
"Introduction to Coat Color Genetics"
''from'' Veterinary Genetics Laboratory, School of Veterinary Medicine, University of California, Davis. Web Site accessed January 12, 2008 {{Equine coat colors https://www.spanishdict.com/translate/grullo Horse coat colors