Grey Partridge
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The grey partridge (''Perdix perdix'') is a bird in the pheasant family
Phasianidae Phasianidae is a family (biology), family of heavy, ground-living birds, which includes pheasants, grouse, partridges, junglefowl, chickens, Turkey bird, turkeys, Old World quail, and peafowl. The family includes many of the most popular Game (hu ...
of the order
Galliformes Galliformes is an order (biology), order of heavy-bodied ground-feeding birds that includes turkey (bird), turkeys, chickens, Old World quail, quail, and other landfowl. Gallinaceous birds, as they are called, are important in their ecosystems ...
, gallinaceous birds. The scientific name is the
Latin Latin ( or ) is a classical language belonging to the Italic languages, Italic branch of the Indo-European languages. Latin was originally spoken by the Latins (Italic tribe), Latins in Latium (now known as Lazio), the lower Tiber area aroun ...
for "partridge".


Taxonomy

The grey partridge formally described in 1758 by the Swedish naturalist
Carl Linnaeus Carl Linnaeus (23 May 1707 – 10 January 1778), also known after ennoblement in 1761 as Carl von Linné,#Blunt, Blunt (2004), p. 171. was a Swedish biologist and physician who formalised binomial nomenclature, the modern system of naming o ...
in the tenth edition of his ''
Systema Naturae ' (originally in Latin written ' with the Orthographic ligature, ligature æ) is one of the major works of the Sweden, Swedish botanist, zoologist and physician Carl Linnaeus (1707–1778) and introduced the Linnaean taxonomy. Although the syste ...
'' under the
binomial name In taxonomy, binomial nomenclature ("two-term naming system"), also called binary nomenclature, is a formal system of naming species of living things by giving each a name composed of two parts, both of which use Latin grammatical forms, altho ...
''Tetrao perdix''. Linnaeus specified the type locality as Europe but this has been restricted to Sweden. The word ''perdix'' is
Latin Latin ( or ) is a classical language belonging to the Italic languages, Italic branch of the Indo-European languages. Latin was originally spoken by the Latins (Italic tribe), Latins in Latium (now known as Lazio), the lower Tiber area aroun ...
meaning "partridge", from
Ancient Greek Ancient Greek (, ; ) includes the forms of the Greek language used in ancient Greece and the classical antiquity, ancient world from around 1500 BC to 300 BC. It is often roughly divided into the following periods: Mycenaean Greek (), Greek ...
περδιξ/''perdix'' meaning "partridge". The grey partridge together with the Daurian partridge and the Tibetan partridge are now placed in the genus '' Perdix'' that was introduced in 1760 by the French zoologist
Mathurin Jacques Brisson Mathurin Jacques Brisson (; 30 April 1723 – 23 June 1806) was a French zoologist and natural philosophy, natural philosopher. Brisson was born on 30 April 1723 at Fontenay-le-Comte in the Vendée department of western France. Note that page 14 ...
.


Subspecies

Eight
subspecies In Taxonomy (biology), biological classification, subspecies (: subspecies) is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (Morphology (biology), morpholog ...
are recognised by the IOC World Bird List, though the differences are clinal, and not all are accepted by other authorities; the HBW/BirdLife International list only accepts six subspecies; the differences noted below: * ''P. p. perdix'' (
Linnaeus Carl Linnaeus (23 May 1707 – 10 January 1778), also known after ennoblement in 1761 as Carl von Linné,#Blunt, Blunt (2004), p. 171. was a Swedish biologist and physician who formalised binomial nomenclature, the modern system of naming o ...
, 1758)
. Nominate, found in the
British Isles The British Isles are an archipelago in the Atlantic Ocean, North Atlantic Ocean off the north-western coast of continental Europe, consisting of the islands of Great Britain, Ireland, the Isle of Man, the Inner Hebrides, Inner and Outer Hebr ...
and southern
Scandinavia Scandinavia is a subregion#Europe, subregion of northern Europe, with strong historical, cultural, and linguistic ties between its constituent peoples. ''Scandinavia'' most commonly refers to Denmark, Norway, and Sweden. It can sometimes also ...
to
Italy Italy, officially the Italian Republic, is a country in Southern Europe, Southern and Western Europe, Western Europe. It consists of Italian Peninsula, a peninsula that extends into the Mediterranean Sea, with the Alps on its northern land b ...
and the
Balkans The Balkans ( , ), corresponding partially with the Balkan Peninsula, is a geographical area in southeastern Europe with various geographical and historical definitions. The region takes its name from the Balkan Mountains that stretch throug ...
. * ''P. p. armoricana'' ( Hartert, 1917). Southwestern
France France, officially the French Republic, is a country located primarily in Western Europe. Overseas France, Its overseas regions and territories include French Guiana in South America, Saint Pierre and Miquelon in the Atlantic Ocean#North Atlan ...
. * ''P. p. sphagnetorum'' ( Altum, 1894). On peaty soils in the northern part of the
Netherlands , Terminology of the Low Countries, informally Holland, is a country in Northwestern Europe, with Caribbean Netherlands, overseas territories in the Caribbean. It is the largest of the four constituent countries of the Kingdom of the Nether ...
and northwest
Germany Germany, officially the Federal Republic of Germany, is a country in Central Europe. It lies between the Baltic Sea and the North Sea to the north and the Alps to the south. Its sixteen States of Germany, constituent states have a total popu ...
. Barely distinct from ''P. p. perdix'' and included in it by HBW/BLI. * ''P. p. hispaniensis'' ( Reichenow, 1892). Central
Pyrenees The Pyrenees are a mountain range straddling the border of France and Spain. They extend nearly from their union with the Cantabrian Mountains to Cap de Creus on the Mediterranean coast, reaching a maximum elevation of at the peak of Aneto. ...
to northeast
Portugal Portugal, officially the Portuguese Republic, is a country on the Iberian Peninsula in Southwestern Europe. Featuring Cabo da Roca, the westernmost point in continental Europe, Portugal borders Spain to its north and east, with which it share ...
* ''P. p. italica'' (Hartert, 1917). Included in ''P. p. perdix'' by HBW/BLI. Formerly extinct in the wild, now reintroduced. * ''P. p. lucida'' ( Altum, 1894).
Finland Finland, officially the Republic of Finland, is a Nordic country in Northern Europe. It borders Sweden to the northwest, Norway to the north, and Russia to the east, with the Gulf of Bothnia to the west and the Gulf of Finland to the south, ...
east to the
Ural Mountains The Ural Mountains ( ),; , ; , or simply the Urals, are a mountain range in Eurasia that runs north–south mostly through Russia, from the coast of the Arctic Ocean to the river Ural (river), Ural and northwestern Kazakhstan.
and south to
Black Sea The Black Sea is a marginal sea, marginal Mediterranean sea (oceanography), mediterranean sea lying between Europe and Asia, east of the Balkans, south of the East European Plain, west of the Caucasus, and north of Anatolia. It is bound ...
and northern
Caucasus The Caucasus () or Caucasia (), is a region spanning Eastern Europe and Western Asia. It is situated between the Black Sea and the Caspian Sea, comprising parts of Southern Russia, Georgia, Armenia, and Azerbaijan. The Caucasus Mountains, i ...
. * ''P. p. canescens'' ( Burturlin, 1906).
Turkey Turkey, officially the Republic of Türkiye, is a country mainly located in Anatolia in West Asia, with a relatively small part called East Thrace in Southeast Europe. It borders the Black Sea to the north; Georgia (country), Georgia, Armen ...
east to the
South Caucasus The South Caucasus, also known as Transcaucasia or the Transcaucasus, is a geographical region on the border of Eastern Europe and West Asia, straddling the southern Caucasus Mountains. The South Caucasus roughly corresponds to modern Armenia, ...
and northwest
Iran Iran, officially the Islamic Republic of Iran (IRI) and also known as Persia, is a country in West Asia. It borders Iraq to the west, Turkey, Azerbaijan, and Armenia to the northwest, the Caspian Sea to the north, Turkmenistan to the nort ...
. * ''P. p. robusta'' ( Homeyer and Tancré, 1883). The
Ural Mountains The Ural Mountains ( ),; , ; , or simply the Urals, are a mountain range in Eurasia that runs north–south mostly through Russia, from the coast of the Arctic Ocean to the river Ural (river), Ural and northwestern Kazakhstan.
to southwestern
Siberia Siberia ( ; , ) is an extensive geographical region comprising all of North Asia, from the Ural Mountains in the west to the Pacific Ocean in the east. It has formed a part of the sovereign territory of Russia and its predecessor states ...
and northwestern
China China, officially the People's Republic of China (PRC), is a country in East Asia. With population of China, a population exceeding 1.4 billion, it is the list of countries by population (United Nations), second-most populous country after ...
. The largest subspecies.


Description

The grey partridge is a rotund bird, brown-backed, with grey flanks and chest. The belly is white, usually marked with a large chestnut-brown horse-shoe mark in males, and also in many females. Hens lay up to twenty eggs in a ground nest. The nest is usually in the margin of a cereal field, most commonly winter wheat. Measurements: * Length: * Wingspan: * Weight (''P. p. perdix''): * Weight (''P. p. robusta''): Males and females are the same size, and very similar in plumage, though the females tend to be slightly duller, and have a smaller dark belly patch. Svensson, L., Mullarney, K., & Zetterström, D. (2022) '' Collins Bird Guide'', ed. 3. , pages 58–59 In the hand, the so-called "cross of Lorraine" on the tertiary coverts of females are marked with two transverse bars, as opposed to the one in males. These are present after around 16 weeks of age when the birds have moulted into adult plumage. Young grey partridges are mostly yellow-brown and lack the distinctive face and underpart markings. The song is a harsh, high-pitched ''kieerr-ik''. When disturbed, like most gamebirds, it flies a short distance on rounded wings, often calling ''rick rick rick'' as it rises. They are a seed-eating species, but the young in particular take
insect Insects (from Latin ') are Hexapoda, hexapod invertebrates of the class (biology), class Insecta. They are the largest group within the arthropod phylum. Insects have a chitinous exoskeleton, a three-part body (Insect morphology#Head, head, ...
s as an essential
protein Proteins are large biomolecules and macromolecules that comprise one or more long chains of amino acid residue (biochemistry), residues. Proteins perform a vast array of functions within organisms, including Enzyme catalysis, catalysing metab ...
supply. During the first 10 days of life, the young can only digest insects. The parents lead their chicks to the edges of
cereal A cereal is a grass cultivated for its edible grain. Cereals are the world's largest crops, and are therefore staple foods. They include rice, wheat, rye, oats, barley, millet, and maize ( Corn). Edible grains from other plant families, ...
fields, where they can forage for insects.


Distribution

Widespread and common throughout much of its range, the grey partridge is evaluated as "of Least Concern" on the
IUCN Red List The International Union for Conservation of Nature (IUCN) Red List of Threatened Species, also known as the IUCN Red List or Red Data Book, founded in 1964, is an inventory of the global conservation status and extinction risk of biological ...
of Threatened Species. However, it has suffered a serious decline in the UK, and in 2015 appeared on the "Birds of Conservation Concern" Red List. This partridge breeds on farmland across most of
Europe Europe is a continent located entirely in the Northern Hemisphere and mostly in the Eastern Hemisphere. It is bordered by the Arctic Ocean to the north, the Atlantic Ocean to the west, the Mediterranean Sea to the south, and Asia to the east ...
and across the western
Palearctic The Palearctic or Palaearctic is a biogeographic realm of the Earth, the largest of eight. Confined almost entirely to the Eastern Hemisphere, it stretches across Europe and Asia, north of the foothills of the Himalayas, and North Africa. Th ...
as far as southwestern Siberia; it is a non-migratory terrestrial species, and forms flocks of up to 30 outside of the breeding season. It has been introduced widely as a
gamebird Galliformes is an order of heavy-bodied ground-feeding birds that includes turkeys, chickens, quail, and other landfowl. Gallinaceous birds, as they are called, are important in their ecosystems as seed dispersers and predators, and are ofte ...
into
Canada Canada is a country in North America. Its Provinces and territories of Canada, ten provinces and three territories extend from the Atlantic Ocean to the Pacific Ocean and northward into the Arctic Ocean, making it the world's List of coun ...
,
United States The United States of America (USA), also known as the United States (U.S.) or America, is a country primarily located in North America. It is a federal republic of 50 U.S. state, states and a federal capital district, Washington, D.C. The 48 ...
,
South Africa South Africa, officially the Republic of South Africa (RSA), is the Southern Africa, southernmost country in Africa. Its Provinces of South Africa, nine provinces are bounded to the south by of coastline that stretches along the Atlantic O ...
,
Australia Australia, officially the Commonwealth of Australia, is a country comprising mainland Australia, the mainland of the Australia (continent), Australian continent, the island of Tasmania and list of islands of Australia, numerous smaller isl ...
and
New Zealand New Zealand () is an island country in the southwestern Pacific Ocean. It consists of two main landmasses—the North Island () and the South Island ()—and List of islands of New Zealand, over 600 smaller islands. It is the List of isla ...
. A popular gamebird in large areas of North America, it is sometimes known there as "Hungarian partridge" or just "hun".


Status and conservation

Though common and not threatened, it is declining in numbers in some areas of intensive cultivation such as the
United Kingdom The United Kingdom of Great Britain and Northern Ireland, commonly known as the United Kingdom (UK) or Britain, is a country in Northwestern Europe, off the coast of European mainland, the continental mainland. It comprises England, Scotlan ...
, due to a loss of breeding habitat and
insecticides Insecticides are pesticides used to kill insects. They include ovicides and larvicides used against insect eggs and larvae, respectively. The major use of insecticides is in agriculture, but they are also used in home and garden settings, in ...
harming insect numbers, an important food source for the species. Their numbers have fallen in these areas by as much as 85% in the last 25 years. Efforts are being made in the
United Kingdom The United Kingdom of Great Britain and Northern Ireland, commonly known as the United Kingdom (UK) or Britain, is a country in Northwestern Europe, off the coast of European mainland, the continental mainland. It comprises England, Scotlan ...
by organizations such as the Game & Wildlife Conservation Trust to halt this decline by creating conservation headlands. In 1995, it was nominated a Biodiversity action plan (BAP) species. In
Ireland Ireland (, ; ; Ulster Scots dialect, Ulster-Scots: ) is an island in the North Atlantic Ocean, in Northwestern Europe. Geopolitically, the island is divided between the Republic of Ireland (officially Names of the Irish state, named Irelan ...
, it is now virtually confined to the Lough Boora reserve in
County Offaly County Offaly (; ) is a Counties of Ireland, county in Republic of Ireland, Ireland. It is part of the Eastern and Midland Region and the Provinces of Ireland, province of Leinster. It is named after the Ancient Ireland, ancient Kingdom of Uí ...
where a recent conservation project has succeeded in boosting its numbers to around 900, raising hopes that it may be reintroduced to the rest of Ireland.


References


External links

*
Game & Wildlife Conservation Trust - Grey Partridge
*

– Cornell Lab of Ornithology
Ageing and sexing (PDF; 2.6 MB) by Javier Blasco-Zumeta & Gerd-Michael Heinze


{{Authority control Game birds grey partridge Birds of Europe Birds of Central Asia Birds of North America grey partridge Taxa named by Carl Linnaeus