Grey-capped Warbler
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The grey-capped warbler (''Eminia lepida'') is a species of
bird Birds are a group of warm-blooded vertebrates constituting the class (biology), class Aves (), characterised by feathers, toothless beaked jaws, the Oviparity, laying of Eggshell, hard-shelled eggs, a high Metabolism, metabolic rate, a fou ...
in the family
Cisticolidae The family Cisticolidae is a group of about 160 warblers, small passerine birds found mainly in warmer southern regions of the Old World. They were formerly included within the Old World warbler family Sylviidae. This family probably originated ...
. It is
monotypic In biology, a monotypic taxon is a taxonomic group (taxon) that contains only one immediately subordinate taxon. A monotypic species is one that does not include subspecies or smaller, infraspecific taxa. In the case of genera, the term "unisp ...
(only species) in the genus ''Eminia''. The grey-capped warbler is found in
Burundi Burundi, officially the Republic of Burundi, is a landlocked country in East Africa. It is located in the Great Rift Valley at the junction between the African Great Lakes region and Southeast Africa, with a population of over 14 million peop ...
,
Democratic Republic of the Congo The Democratic Republic of the Congo (DRC), also known as the DR Congo, Congo-Kinshasa, or simply the Congo (the last ambiguously also referring to the neighbouring Republic of the Congo), is a country in Central Africa. By land area, it is t ...
,
Kenya Kenya, officially the Republic of Kenya, is a country located in East Africa. With an estimated population of more than 52.4 million as of mid-2024, Kenya is the 27th-most-populous country in the world and the 7th most populous in Africa. ...
,
Rwanda Rwanda, officially the Republic of Rwanda, is a landlocked country in the Great Rift Valley of East Africa, where the African Great Lakes region and Southeast Africa converge. Located a few degrees south of the Equator, Rwanda is bordered by ...
,
South Sudan South Sudan (), officially the Republic of South Sudan, is a landlocked country in East Africa. It is bordered on the north by Sudan; on the east by Ethiopia; on the south by the Democratic Republic of the Congo, Uganda and Kenya; and on the ...
,
Tanzania Tanzania, officially the United Republic of Tanzania, is a country in East Africa within the African Great Lakes region. It is bordered by Uganda to the northwest; Kenya to the northeast; the Indian Ocean to the east; Mozambique and Malawi to t ...
, and
Uganda Uganda, officially the Republic of Uganda, is a landlocked country in East Africa. It is bordered to the east by Kenya, to the north by South Sudan, to the west by the Democratic Republic of the Congo, to the south-west by Rwanda, and to the ...
. It is a large, chunky, thin-tailed-warbler with a distinctive grey cap, a black band around its head, and a chestnut throat wrapping its neck. Grey-capped warblers maintain a diet of insects and other invertebrates, including caterpillars, moths, grasshoppers, and mantids.


Identification

The grey-capped warbler is roughly 15 cm in length, and weighs 16–24 grams. It is typically chunky and thin-tailed with a distinctive grey cap, a black band around the head, and a chestnut throat with rump feathers that are long and lax. Its cheek, neck, and underparts are grey and it is tinged olive-green on the flanks and under tail-coverts. Grey-capped warblers have a chestnut vent patch and upperparts. The tail and the median and greater upper wing-coverts are olive-green. The lesser coverts and underwing-coverts are chestnut. Flight-feathers are dark brown (edged olive-green). The irises are reddish-brown, the bill and mouth are black, and the legs are pinkish-brown. Identification for sexes is similar. Younger grey-capped warblers are duller than adults and have a smaller, paler throat patch and brown eyes. Recently fledged warblers have a prominent yellow gape and shorter tail.


Habitat

The natural
habitat In ecology, habitat refers to the array of resources, biotic factors that are present in an area, such as to support the survival and reproduction of a particular species. A species' habitat can be seen as the physical manifestation of its ...
of the grey-capped warbler is
subtropical The subtropical zones or subtropics are geographical zone, geographical and Köppen climate classification, climate zones immediately to the Northern Hemisphere, north and Southern Hemisphere, south of the tropics. Geographically part of the Ge ...
or moist
tropical The tropics are the regions of Earth surrounding the equator, where the sun may shine directly overhead. This contrasts with the temperate or polar regions of Earth, where the Sun can never be directly overhead. This is because of Earth's ax ...
shrubland Shrubland, scrubland, scrub, brush, or bush is a plant community characterized by vegetation dominance (ecology), dominated by shrubs, often also including grasses, herbaceous plant, herbs, and geophytes. Shrubland may either occur naturally o ...
. It is found in dense undergrowth, scrub, and creepers, usually in damp areas along forest edges, seeps, or riparian strips, as well as around well-planted gardens. It is confined to areas with annual rainfall exceeding 500 mm. It usually dwells in areas at altitudes from 800 m to 2,500 m in Kenya. It is regarded as a sedentary bird.


Diet and Foraging

Its diet includes insects and other invertebrates, including caterpillars and moths (
Lepidoptera Lepidoptera ( ) or lepidopterans is an order (biology), order of winged insects which includes butterflies and moths. About 180,000 species of the Lepidoptera have been described, representing 10% of the total described species of living organ ...
), grasshoppers (
Orthoptera Orthoptera () is an order of insects that comprises the grasshoppers, locusts, and crickets, including closely related insects, such as the bush crickets or katydids and wētā. The order is subdivided into two suborders: Caelifera – gras ...
), mantids (
Mantodea Mantises are an Order (biology), order (Mantodea) of insects that contains List of mantis genera and species, over 2,400 species in about 460 genera in 33 families. The largest family is the Mantidae ("mantids") ...
), spiders (
Araneae Spiders ( order Araneae) are air-breathing arthropods that have eight limbs, chelicerae with fangs generally able to inject venom, and spinnerets that extrude silk. They are the largest order of arachnids and rank seventh in total species ...
), and millipedes (
Diplopoda Millipedes (originating from the Latin , "thousand", and , "foot") are a group of arthropods that are characterised by having two pairs of jointed legs on most body segments; they are known scientifically as the class Diplopoda, the name derive ...
). It forages by gleaning from foliage and bark, actively searching among dead, curled-up leaves, crevices in bark, and holes in twigs. It is secretive and lurks among dense vegetation, and may easily be overlooked if not singing. It is joined by a large variety of frugivorous and insectivorous species attracted to stands of fruiting
Euclea divinorum ''Euclea divinorum'', called diamond leaf, diamond-leaved euclea, magic guarri, and toothbrush tree, is a species of flowering plant in the genus '' Euclea'', native to eastern and southern Africa. A shrub or small tree, it has many uses in Afric ...
trees in Tanzania.


Sounds and Vocal Behavior

Male songs usually come from an exposed perch, mainly during the breeding season, making a loud, varied series of trills and whistles, lasting 6–20 seconds, typically containing several repeated elements uttered every 30 seconds. The female utters a dry trill with up to 7 notes per second, usually during the male song in a duet. The male also utters trilling calls, alarms with a soft “pree”, or louder scolding calls when agitated.


Breeding

The grey-capped warbler breeds during rains: Mar-Jun and Oct–Nov in Sudan and DR Congo, Apr-May in Uganda, and May–Aug and Nov–Jan in Kenya. It attempts multiple broods. Possibly monogamous, partners apparently remain together throughout the year, being solitary and territorial. When singing together, partners sit close together or hop around each other, with throat feathers raised and their tail fanned and erect. Once the duet ends, the male bends forward, holding his back, rump, and flank feathers raised in the manner of a puff back (
Dryoscopus ''Dryoscopus'' is a genus of bird in the Malaconotidae or bushshrike family. Its members are known as puffbacks. The six species, all of fairly uniform appearance and habits, are Indigenous (ecology), native to various parts of sub-Saharan Africa ...
). Nests are built by both sexes, with the male sometimes calling while the female builds. Construction of a nest usually takes 2 weeks. The nest is an untidy, ball-shaped structure, with a side entrance and protruding porch (up to 8 cm long) and platform (2·5 cm), made of long strips of fibrous vegetation and leaves, woven together and lined with moss, spider webs, rootlets, feathers and plant down, suspended from a thin branch or creeper inside dense vegetation, sometimes slung between two saplings, usually, 1–3 m (rarely 5 m) up, often over water, where it resembles debris lodged in the overhanging branch. The nest is sometimes re-used in successive seasons; one record of a weaver (
Ploceidae Ploceidae is a family of small passerine birds, many of which are called weavers, weaverbirds, weaver finches, or bishops. These names come from the nests of intricately woven vegetation created by birds in this family. In most recent classificat ...
) nest being used after entrance modified and long, trailing vegetation added to enhance camouflage. The clutch includes 2 or 3 eggs incubated mainly by the female for 12–13 days. Chicks are fed by both sexes, the male often passing food to the female and she then feeds the nestlings. Both sexes also remove feces, dropping them some distance away from the nest. The fledging period is 16 days with the adults accompanying the fledglings, calling loudly if approached. Breeding success is low, averaging only 0·27 broods/pair/year in Kenya (multiple attempts during a protracted breeding season) with some streamside nests destroyed by floodwaters.


References

*Ryan, Peter (2006). Family Cisticolidae (Cisticolas and allies). pp. 378–492 in del Hoyo J., Elliott A. & Christie D.A. (2006) ''
Handbook of the Birds of the World The ''Handbook of the Birds of the World'' (HBW) is a multi-volume series produced by the Spanish publishing house Lynx Edicions in partnership with BirdLife International. It is the first handbook to cover every known living species of bird. ...
''. Volume 11. ''Old World Flycatchers to Old World Warblers'' Lynx Edicions, Barcelona * Nguembock B.; Fjeldsa J.; Tillier A.; Pasquet E. (2007): A phylogeny for the Cisticolidae (Aves: Passeriformes) based on nuclear and mitochondrial DNA sequence data, and a re-interpretation of unique nest-building specialization. ''Molecular Phylogenetics and Evolution'' 42: 272–286. *Ryan, P. (2020, March 4). Gray-capped Warbler (Eminia Lepida). Retrieved October 16, 2020, from https://birdsoftheworld.org/bow/species/gycwar3/cur/introduction *Canto. (1881). Retrieved October 16, 2020, from https://www.xeno-canto.org/species/Eminia-lepida *Grey-capped Warbler- Eminia Lepida. (2003). Retrieved October 16, 2020, from https://avibase.bsc-eoc.org/species.jsp?avibaseid=31E5F1E9921CF416 {{Taxonbar, from=Q2512467 grey-capped warbler Birds of East Africa Grey-capped warbler Taxonomy articles created by Polbot