Great Helen
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''Papilio iswara'', the great Helen, is a species of large
swallowtail butterfly Swallowtail butterflies are large, colorful butterflies in the family Papilionidae, and include over 550 species. Though the majority are tropical, members of the family inhabit every continent except Antarctica. The family includes the larg ...
found in parts of
Southeast Asia Southeast Asia is the geographical United Nations geoscheme for Asia#South-eastern Asia, southeastern region of Asia, consisting of the regions that are situated south of China, east of the Indian subcontinent, and northwest of the Mainland Au ...
.


Description

P. iswara. Very similar to '' P. helenus''. Palpi entirely white; the white area of the hindwing consists of 4 patches, of which the 3rd is the largest; the area in the male smaller beneath than above, with the exception of the 4th spot, which is larger beneath, between it and the red anal eye-spot in the female two more white spots, of which at least the upper one is also distinct in the mostly only 2 red submarginal spots present, both large, usually forming a ring-spot with the marginal spots, before the anal submarginal spot no red discal spot as in ''P. helenus'', but always 3 blue lunules. The disc of the upper surface of the forewing in the with pilose hairs. The female paler than the male, the red submarginal spots larger on both surfaces. The genitalia similar to those of ''P. helenus'', the harpe of the more spoon-shaped and the infra-anal processes shorter. Like ''P. helenus'' occurring in wooded districts, principally in the hills. The early stages not known. Distributed from South Tenasserim to Sumatra and Borneo; only locally plentiful. -— ''iswara'' White (21 c). The white area of the hindwing posteriorly broad; the yellow-red spots of the underside of the hindwing large, the anterior ring-spot also in the male entire, the black pupils of both the eye-spots in the female small. Lower Tenasserim, Malay Peninsula, Penang, Sumatra, Banka. The specimens from the last two islands approximate to the next form. —''araspes'' Fldr. On an average smaller than ''iswara'', but the largest specimens much larger than the smallest ''iswara''. The white area of the hindwing mostly not so broad as in ''iswara'', posteriorly more pointed; the red spots of the hindwing beneath and the 4th white discal patch smaller. One of the two females in the Tring Museum from Lawas, North Borneo, on the underside of the hindwing anteriorly 3 thin red submarginal lunules, before the 1. median a fine red longitudinal curved mark and before the tail traces of another submarginal spot, so that the complete row of submarginal lunules is at least indicated. Borneo and Natuna Islands; our two specimens from Bungaran, Natuna, approximate a little to ''iswar''- Karl Jordan in Seitz.Seitz, A. Band 9: Abt. 2, Die exotischen Großschmetterlinge, Die indo-australischen Tagfalter, 1927, 1197 Seiten 177 Tafeln


Status

''Papilio iswara'' is not uncommon and not
threatened A threatened species is any species (including animals, plants and fungi) which is vulnerable to extinction in the near future. Species that are threatened are sometimes characterised by the population dynamics measure of ''critical depensatio ...
.


Subspecies

*''Papilio iswara iswara'' (southern Burma to Singapore, Sumatra, Bangka) *''Papilio iswara araspes'' C. & R. Felder, 1859 (northern Borneo, Natuna Island)


Taxonomy

''Papilio iswara'' is a member of the ''helenus''
species group In biology, a species complex is a group of closely related organisms that are so similar in appearance and other features that the boundaries between them are often unclear. The taxa in the complex may be able to hybridize readily with each oth ...
. The members of this
clade In biology, a clade (), also known as a Monophyly, monophyletic group or natural group, is a group of organisms that is composed of a common ancestor and all of its descendants. Clades are the fundamental unit of cladistics, a modern approach t ...
are: *'' Papilio helenus'' Linnaeus, 1758 *''Papilio iswara'' White, 1842 *'' Papilio iswaroides'' Fruhstorfer, 1898 *''
Papilio nephelus ''Papilio nephelus'' is a species of swallowtail butterfly belonging to the family Papilionidae. Subspecies include ''P. n. chaon'', the yellow Helen, and ''P. n. sunatus'', the black and white Helen. Description ''Papilio nephelus'' has a win ...
'' Boisduval, 1836 *'' Papilio nubilus'' Staudinger, 1895 *'' Papilio sataspes'' C. & R. Felder, 1865


Type material

The
holotype A holotype (Latin: ''holotypus'') is a single physical example (or illustration) of an organism used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of s ...
is conserved in the
Natural History Museum, London The Natural History Museum in London is a museum that exhibits a vast range of specimens from various segments of natural history. It is one of three major museums on Exhibition Road in South Kensington, the others being the Science Museum (Lo ...
.


References

* White, A., 1842, Notice of two New Species of ''Papilio'' from Penang, presented to the British Museum by Sir. Wm. Norris. ''
Entomologist Entomology (from Ancient Greek ἔντομον (''éntomon''), meaning "insect", and -logy from λόγος (''lógos''), meaning "study") is the branch of zoology that focuses on insects. Those who study entomology are known as entomologists. In ...
'' 1:280

*


External links


Butterflies in Indo-China
Papilio, iswara Butterflies described in 1842 Butterflies of Singapore Butterflies of Borneo Butterflies of Indochina Taxa named by Adam White (zoologist) Lepidoptera of Sumatra Butterflies of Malaysia {{Papilionidae-stub