Grazing (behavior)
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Grazing is a method of
feeding Eating (also known as consuming) is the ingestion of food. In biology, this is typically done to provide a heterotrophic organism with energy and nutrients and to allow for growth. Animals and other heterotrophs must eat in order to survive – ...
in which a
herbivore A herbivore is an animal anatomically and physiologically evolved to feed on plants, especially upon vascular tissues such as foliage, fruits or seeds, as the main component of its diet. These more broadly also encompass animals that eat ...
feeds on low-growing
plant Plants are the eukaryotes that form the Kingdom (biology), kingdom Plantae; they are predominantly Photosynthesis, photosynthetic. This means that they obtain their energy from sunlight, using chloroplasts derived from endosymbiosis with c ...
s such as
grass Poaceae ( ), also called Gramineae ( ), is a large and nearly ubiquitous family (biology), family of monocotyledonous flowering plants commonly known as grasses. It includes the cereal grasses, bamboos, the grasses of natural grassland and spe ...
es or other multicellular organisms, such as
algae Algae ( , ; : alga ) is an informal term for any organisms of a large and diverse group of photosynthesis, photosynthetic organisms that are not plants, and includes species from multiple distinct clades. Such organisms range from unicellular ...
. Many
species A species () is often defined as the largest group of organisms in which any two individuals of the appropriate sexes or mating types can produce fertile offspring, typically by sexual reproduction. It is the basic unit of Taxonomy (biology), ...
of animals can be said to be grazers, from large animals such as
hippopotamuses The hippopotamus (''Hippopotamus amphibius;'' ; : hippopotamuses), often shortened to hippo (: hippos), further qualified as the common hippopotamus, Nile hippopotamus and river hippopotamus, is a large semiaquatic Mammal, mammal native to su ...
to small aquatic
snails A snail is a shelled gastropod. The name is most often applied to land snails, terrestrial molluscs, terrestrial pulmonate gastropod molluscs. However, the common name ''snail'' is also used for most of the members of the molluscan class Gas ...
. Grazing
behaviour Behavior (American English) or behaviour (British English) is the range of actions of Individual, individuals, organisms, systems or Artificial intelligence, artificial entities in some environment. These systems can include other systems or or ...
is a type of feeding strategy within the
ecology Ecology () is the natural science of the relationships among living organisms and their Natural environment, environment. Ecology considers organisms at the individual, population, community (ecology), community, ecosystem, and biosphere lev ...
of a species. Specific grazing strategies include graminivory (eating grasses);
coprophagy Coprophagia ( ) or coprophagy ( ) is the consumption of feces. The word is derived from the Ancient Greek "feces" and "to eat". Coprophagy refers to many kinds of feces-eating, including eating feces of other species (heterospecifics), of o ...
(producing part-digested pellets which are reingested);
pseudoruminant Pseudoruminant is a classification of animals based on their digestive tract differing from the ruminants. Hippopotamidae, Hippopotami and camels are ungulate mammals with a three-chambered stomach (ruminants have a four-chambered stomach) while Eq ...
(having a multi-chambered stomach but not
chewing the cud Ruminants are herbivorous grazing or browsing artiodactyls belonging to the suborder Ruminantia that are able to acquire nutrients from plant-based food by fermenting it in a specialized stomach prior to digestion, principally through microbia ...
); and grazing on plants other than grass, such as on
marine algae Marine is an adjective meaning of or pertaining to the sea or ocean. Marine or marines may refer to: Ocean * Maritime (disambiguation) * Marine art * Marine biology * Marine current power * Marine debris * Marine energy * Marine habitats * M ...
. Grazing's ecological effects can include redistributing nutrients, keeping grasslands open or favouring a particular species over another.


Ecology

Many small selective herbivores follow larger grazers which skim off the highest, tough growth of grasses, exposing tender shoots. For terrestrial animals, grazing is normally distinguished from
browsing Browsing is a kind of orienting strategy. It is supposed to identify something of relevance for the browsing organism. In context of humans, it is a metaphor taken from the animal kingdom. It is used, for example, about people browsing open sh ...
in that grazing is eating grass or
forb A forb or phorb is a herbaceous flowering plant that is not a graminoid (grass, sedge, or rush). The term is used in botany and in vegetation ecology especially in relation to grasslands and understory. Typically, these are eudicots without woo ...
s, whereas browsing is eating woody twigs and
leaves A leaf (: leaves) is a principal appendage of the stem of a vascular plant, usually borne laterally above ground and specialized for photosynthesis. Leaves are collectively called foliage, as in "autumn foliage", while the leaves, stem, ...
from
tree In botany, a tree is a perennial plant with an elongated stem, or trunk, usually supporting branches and leaves. In some usages, the definition of a tree may be narrower, e.g., including only woody plants with secondary growth, only ...
s and
shrub A shrub or bush is a small to medium-sized perennial woody plant. Unlike herbaceous plants, shrubs have persistent woody stems above the ground. Shrubs can be either deciduous or evergreen. They are distinguished from trees by their multiple ...
s. Grazing differs from
predation Predation is a biological interaction in which one organism, the predator, kills and eats another organism, its prey. It is one of a family of common List of feeding behaviours, feeding behaviours that includes parasitism and micropredation ...
because the organism being grazed upon may not be killed. It differs from
parasitism Parasitism is a close relationship between species, where one organism, the parasite, lives (at least some of the time) on or inside another organism, the host, causing it some harm, and is adapted structurally to this way of life. The en ...
because the two organisms live together in a constant state of physical externality (i.e., low intimacy).
Water animal An aquatic animal is any animal, whether vertebrate or invertebrate, that lives in a body of water for all or most of its lifetime. Aquatic animals generally conduct gas exchange in water by extracting dissolved oxygen via specialised respirato ...
s that feed by rasping algae and other micro-organisms from stones are called ''grazers–scrapers''.


Graminivory

Graminivory is a form of grazing involving feeding primarily on grass (specifically "true" grasses in the
Poaceae Poaceae ( ), also called Gramineae ( ), is a large and nearly ubiquitous family of monocotyledonous flowering plants commonly known as grasses. It includes the cereal grasses, bamboos, the grasses of natural grassland and species cultivate ...
).
Horse The horse (''Equus ferus caballus'') is a domesticated, one-toed, hoofed mammal. It belongs to the taxonomic family Equidae and is one of two extant subspecies of ''Equus ferus''. The horse has evolved over the past 45 to 55 mi ...
s,
cattle Cattle (''Bos taurus'') are large, domesticated, bovid ungulates widely kept as livestock. They are prominent modern members of the subfamily Bovinae and the most widespread species of the genus '' Bos''. Mature female cattle are calle ...
,
capybara The capybara or greater capybara (''Hydrochoerus hydrochaeris'') is the largest living rodent, native to South America. It is a member of the genus '' Hydrochoerus''. The only other extant member is the lesser capybara (''Hydrochoerus isthmi ...
,
hippopotamus The hippopotamus (''Hippopotamus amphibius;'' ; : hippopotamuses), often shortened to hippo (: hippos), further qualified as the common hippopotamus, Nile hippopotamus and river hippopotamus, is a large semiaquatic mammal native to sub-Sahar ...
es,
grasshopper Grasshoppers are a group of insects belonging to the suborder Caelifera. They are amongst what are possibly the most ancient living groups of chewing herbivorous insects, dating back to the early Triassic around 250 million years ago. Grassh ...
s,
geese A goose (: geese) is a bird of any of several waterfowl species in the family Anatidae. This group comprises the genera '' Anser'' (grey geese and white geese) and ''Branta'' (black geese). Some members of the Tadorninae subfamily (e.g., Egyp ...
, and
giant panda The giant panda (''Ailuropoda melanoleuca''), also known as the panda bear or simply panda, is a bear species endemic to China. It is characterised by its white animal coat, coat with black patches around the eyes, ears, legs and shoulders. ...
s are graminivores. Giant pandas (''Ailuropoda melanoleuca'') are obligate
bamboo Bamboos are a diverse group of mostly evergreen perennial plant, perennial flowering plants making up the subfamily (biology), subfamily Bambusoideae of the grass family Poaceae. Giant bamboos are the largest members of the grass family, in th ...
grazers, 99% of their diet consisting of sub-alpine bamboo species.


Cecotrophy

For
lagomorphs The lagomorphs () are the members of the taxonomic order Lagomorpha, of which there are two living families: the Leporidae (rabbits and hares) and the Ochotonidae (pikas). There are 110 recent species of lagomorph, of which 109 species in twelve ...
(
rabbit Rabbits are small mammals in the family Leporidae (which also includes the hares), which is in the order Lagomorpha (which also includes pikas). They are familiar throughout the world as a small herbivore, a prey animal, a domesticated ...
s,
hare Hares and jackrabbits are mammals belonging to the genus ''Lepus''. They are herbivores and live Solitary animal, solitarily or in pairs. They nest in slight depressions called forms, and their young are precociality, able to fend for themselves ...
s,
pika A pika ( , or ) is a small, mountain-dwelling mammal native to Asia and North America. With short limbs, a very round body, an even coat of fur, and no external tail, they resemble their close relative the rabbit, but with short, rounded ears. ...
s), easily digestible food is processed in the gastrointestinal tract & expelled as regular feces. But to get nutrients out of hard-to-digest fiber, lagomorphs ferment fiber in the cecum (in the GI tract) and then expel the contents as
cecotrope Cecotropes (also caecotropes, cecotrophs, caecotrophs, cecal pellets, soft feces, or night feces) are a nutrient-filled package created in the gastrointestinal (GI) tract that is expelled and eaten by many animals (such as rabbits, guinea pigs, ...
s, which are reingested (
cecotrophy Cecotropes (also caecotropes, cecotrophs, caecotrophs, cecal pellets, soft feces, or night feces) are a nutrient-filled package created in the gastrointestinal (GI) tract that is expelled and eaten by many animals (such as rabbits, guinea pigs, mi ...
). The cecotropes are then absorbed in the small intestine to utilize the nutrients. This process is different from cows chewing their cud but with similar results.
Capybara The capybara or greater capybara (''Hydrochoerus hydrochaeris'') is the largest living rodent, native to South America. It is a member of the genus '' Hydrochoerus''. The only other extant member is the lesser capybara (''Hydrochoerus isthmi ...
(''Hydrochoerus hydrochaeris'') are herbivores that graze mainly on grasses and aquatic plants,Capybara Facts.
Smithsonian National Zoological Park The National Zoological Park, commonly known as the National Zoo, is one of the oldest zoos in the United States. The zoo is part of the Smithsonian Institution and does not charge admission. Founded in 1889, its mission is to "provide engaging ...
. Retrieved on December 16, 2007.
PDF
/ref> as well as
fruit In botany, a fruit is the seed-bearing structure in flowering plants (angiosperms) that is formed from the ovary after flowering. Fruits are the means by which angiosperms disseminate their seeds. Edible fruits in particular have long propaga ...
and tree
bark Bark may refer to: Common meanings * Bark (botany), an outer layer of a woody plant such as a tree or stick * Bark (sound), a vocalization of some animals (which is commonly the dog) Arts and entertainment * ''Bark'' (Jefferson Airplane album), ...
.Capybara.
Palm Beach Zoo. Retrieved on December 17, 2007.
As with other grazers, they can be very selective, feeding on the leaves of one species and disregarding other species surrounding it. They eat a greater variety of plants during the dry season, as fewer plants are available. While they eat grass during the wet season, they have to switch to more abundant
reeds Reed or Reeds may refer to: Science, technology, biology, and medicine * Reed bird (disambiguation) * Reed pen, writing implement in use since ancient times * Reed (plant), one of several tall, grass-like wetland plants of the order Poales * Re ...
during the dry season. The capybara's jaw hinge is not perpendicular; hence, it chews food by grinding back-and-forth rather than side-to-side.Capybara. ''Hydrochaeris hydrochaeris''
San Francisco Zoo
Like lagomorphs, capybara create, expel & eat
cecotrope Cecotropes (also caecotropes, cecotrophs, caecotrophs, cecal pellets, soft feces, or night feces) are a nutrient-filled package created in the gastrointestinal (GI) tract that is expelled and eaten by many animals (such as rabbits, guinea pigs, ...
s (
cecotrophy Cecotropes (also caecotropes, cecotrophs, caecotrophs, cecal pellets, soft feces, or night feces) are a nutrient-filled package created in the gastrointestinal (GI) tract that is expelled and eaten by many animals (such as rabbits, guinea pigs, mi ...
) to get more nutrition from their food. They may also regurgitate food to masticate again, similar to cud-chewing by a cow. As with other rodents, the front teeth of capybara grow continually to compensate for the constant wear from eating grasses.Bristol Zoo Gardens (UK) ''Capybara''
. Bristolzoo.org.uk. Retrieved on 2011-12-07.
Their cheek teeth also grow continuously.


Pseudoruminant

The hippopotamus is a large, semi-aquatic mammal inhabiting rivers, lakes, and mangrove swamps. During the day, they remain cool by staying in the water or mud; reproduction and childbirth occur in water. They emerge at dusk to graze on grasses. While hippopotamuses rest near each other in the water, grazing is solitary. Their
incisor Incisors (from Latin ''incidere'', "to cut") are the front teeth present in most mammals. They are located in the premaxilla above and on the mandible below. Humans have a total of eight (two on each side, top and bottom). Opossums have 18, wher ...
s can be as long as and the canines (tusks) up to ; however, the canines and incisors are used for combat, and play no role in feeding. Hippos rely on their broad, horny lips to grasp and pull grasses which are then ground by the
molars The molars or molar teeth are large, flat tooth, teeth at the back of the mouth. They are more developed in mammal, mammals. They are used primarily to comminution, grind food during mastication, chewing. The name ''molar'' derives from Latin, '' ...
. The hippo is considered to be a
pseudoruminant Pseudoruminant is a classification of animals based on their digestive tract differing from the ruminants. Hippopotamidae, Hippopotami and camels are ungulate mammals with a three-chambered stomach (ruminants have a four-chambered stomach) while Eq ...
; it has a complex three- or four-chambered stomach but does not "chew cud".


Non-grass grazing

Although grazing is typically associated with
mammal A mammal () is a vertebrate animal of the Class (biology), class Mammalia (). Mammals are characterised by the presence of milk-producing mammary glands for feeding their young, a broad neocortex region of the brain, fur or hair, and three ...
s feeding on
grassland A grassland is an area where the vegetation is dominance (ecology), dominated by grasses (Poaceae). However, sedge (Cyperaceae) and rush (Juncaceae) can also be found along with variable proportions of legumes such as clover, and other Herbaceo ...
s,
ecologist Ecology () is the natural science of the relationships among living organisms and their environment. Ecology considers organisms at the individual, population, community, ecosystem, and biosphere levels. Ecology overlaps with the closely re ...
s sometimes use the word in a broader sense to include any organism that feeds on any other species without ending the life of the
prey Predation is a biological interaction in which one organism, the predator, kills and eats another organism, its prey. It is one of a family of common feeding behaviours that includes parasitism and micropredation (which usually do not ki ...
organism. Use of the term "grazing" varies further; for example, a
marine biologist Marine biology is the scientific study of the biology of marine life, organisms that inhabit the sea. Given that in biology many phyla, families and genera have some species that live in the sea and others that live on land, marine biology clas ...
may describe herbivorous
sea urchin Sea urchins or urchins () are echinoderms in the class (biology), class Echinoidea. About 950 species live on the seabed, inhabiting all oceans and depth zones from the intertidal zone to deep seas of . They typically have a globular body cove ...
s that feed on kelp as ''grazers'', even when they kill the organism by cutting the plant at the base.
Malacologist Malacology, from Ancient Greek μαλακός (''malakós''), meaning "soft", and λόγος (''lógos''), meaning "study", is the branch of invertebrate zoology that deals with the study of the Mollusca (molluscs or mollusks), the second-largest ...
s sometimes apply the word to aquatic snails that feed by consuming the microscopic film of
alga Algae ( , ; : alga ) is an informal term for any organisms of a large and diverse group of photosynthetic organisms that are not plants, and includes species from multiple distinct clades. Such organisms range from unicellular microalgae, suc ...
e, diatoms and detritus—a biofilm—that covers the substrate (biology), substrate and other surfaces underwater. In marine ecosystems, grazing by mesograzers such as some crustaceans maintains habitat structure by preventing algal overgrowth, especially in coral reefs.


Benefits


Environmental

Grazer urine and feces "recycle nitrogen, phosphorus, potassium and other plant nutrients and return them to the soil"."Benefits of Grazing Cattle on the Prairie". Native Habitat Organization. Retrieved 1 Dec 2008
Grazing can allow for the accumulation of organic matter which may help to combat soil erosion. This acts as nutrition for insects and organisms found within the soil. These organisms "aid in carbon sequestration and water filtration".


Biodiversity

When grass is grazed, dead litter grass is reduced which is advantageous for birds such as waterfowl. Grazing can increase biodiversity. Without grazing, many of the same grasses grow, for example Bromus, brome and Poa, bluegrass, consequently producing a monoculture.


References


External links

{{Authority control Ethology Herbivory Land use