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''Graphium leonidas'', the veined swordtail, veined swallowtail or common graphium, is a species of butterfly in the family Papilionidae, found in
Sub-Saharan Africa Sub-Saharan Africa is, geographically, the area and regions of the continent of Africa that lies south of the Sahara. These include West Africa, East Africa, Central Africa, and Southern Africa. Geopolitically, in addition to the List of sov ...
.''Graphium''
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Description

For terms see External morphology of Lepidoptera The wingspan is 75–80 mm in males and 75–85 mm in females. Both wings beneath without distinct dark longitudinal streaks on the folds between the veins and without dark longitudinal streaks in the cell of the hindwing. Both wings with submarginal spots. The discal spot of cellule 1 b of the forewing never reaches the base of the cellule. Hindwing usually with free discal spots in cellules 2—5 and with a broad basal transverse band, which covers the greater part of the cell and the base of cellules 1 c, 2, 6 and 7. Forewing always with discal spots in cellules 1 b, 3, 7 and 8, and usually also with small ones in 2, 4, 5 and 6; the cell mostly with three spots. — ''leonidas'' F. (= ''similis'' Cr.) (7 d). All the spots light blue (male) or bluish white (female); forewing without discal spot in cellule 1 a; the submarginal spots of both wings large; the discal spot of cellule 4 of the forewing is proximally rounded and does not reach the base of the cellule. Sierra Leone, Delagoa Bay, British East Africa. In ab. (?) ''interniplaga'' Auriv. the forewing has a large discal onidale. spot in cellule 1 a, otherwise it is similar to the preceding form. German East Africa, ab. (?) ''onidale'' Suf. has the light basal part of the hindwing sprinkled with black scales, the veins in that part are margined with black and the spots in cellules 1 c and 2 are small or entirely wanting; otherwise similar to the form ''leonidas''. German East Africa: Muanza. — ''brasidas'' Fldr. (7 d). Here the spots on the upper surface are small, white or yellowish white; the submarginal spots dot-like. No discal spot in cellule 1 a of the forewing. Angola to Natal. — ''umanus'' R. & J. differs from all the preceding in that the light basal transverse band of the hindwing only extends as far as the base of vein 2. Abyssinia. — ''pelopidas'' Oberth. The spots of the forewing light bluish, on an average somewhat larger than in leonidas, the discal spot in cellule 4 reaches the base of the cellule, the basal transverse band of the hindwing is very broad and almost pure white, but the submarginal spots are small, dot-like, not larger than the discal spots. A little known form as yet, perhaps a separate species, although it agrees very closely with ''leonidas'' in the markings. German East Africa and Pemba Island.


Biology

Has continuous broods, peaking from October to April. The larvae feed on '' Popowia caffra'', ''
Annona ''Annona'' (from Taíno ''annon'') is a genus of flowering plants in the pawpaw/sugar apple family, Annonaceae. It is the second largest genus in the family after ''Guatteria'', containing approximately 166
'', ''
Monanthotaxis ''Monanthotaxis'' is an Afrotropical The Afrotropical realm is one of Earth's eight biogeographic realms. It includes Africa south of the Sahara Desert, the majority of the Arabian Peninsula, the island of Madagascar, southern Iran and extre ...
'', '' Uvaria'', ''
Monanthotaxis caffra ''Monanthotaxis'' is an Afrotropical plant genus with some 56 species, belonging to the Annonaceae. They are native to the tropics and subtropics of southeastern Africa and Madagascar Madagascar (; mg, Madagasikara, ), officially the Rep ...
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Annona senegalensis ''Annona senegalensis'', commonly known as African custard-apple, wild custard apple, wild soursop, ( Mandinka language), and ( Wolof language) is a species of flowering plant in the custard apple family, Annonaceae. The specific epithet, ''s ...
'', ''
Landolphia ugandensis ''Landolphia'' is a genus of flowering plants in the family Apocynaceae first described as a genus in 1806. They take the form of vines that scramble over host trees. ''Landolphia'' is native to tropical Africa. Characteristics There are about f ...
'', '' L. buchannani'', ''
Annickia chlorantha ''Annickia chlorantha'' is a tree that grows up to 25 meters tall, it belongs to the Annonaceae family. An important tree used in traditional medical practices for the treatment of malaria and various diseases in Nigeria and Cameroon; oil extra ...
'', ''
Friesodielsia obovata ''Friesodielsia'' is a genus of flowering plants in the custard apple and soursop family Annonaceae, with all species found in the Old World, mostly in the tropics. A molecular study shows that ''Friesodielsia'' should be more narrowly circumscr ...
'', '' Uvaria'' and ''
Artabotrys ''Artabotrys'' is a genus of plants in the Annonaceae family. There are over 100 species in the Old World tropics, with 31 species in Africa. It is part of the custard apple family (Annonaceae). All species are small trees or shrubs with a tenden ...
'' species.


Subspecies

*''Graphium leonidas leonidas'' (Sub-Saharan Africa) *''Graphium leonidas pelopidas'' (Oberthür, 1879) (Tanzania: islands of Unguja and Pemba) *''Graphium leonidas santamarthae'' (Joicey & Talbot, 1927) (São Tomé and Príncipe: Principe) *''Graphium leonidas thomasius'' (Le Cerf, 1924) (São Tomé and Príncipe: Sao Tome) *''Graphium leonidas zanzibaricus'' Kielland, 1990Kielland, J. 1990 in Kielland, J. 1990. ''Butterflies of Tanzania.'' Hill House, Melbourne and London: 1-363. Tanzania


Taxonomy

It is the nominal member of the ''leonidas''-group of closely related and similar species (''Graphium leonidas'', '' Graphium levassori'', '' Graphium cyrnus'').


Images

External images
from Royal Museum of Central Africa.


References

*Carcasson, R.H. 1960 "The Swallowtail Butterflies of East Africa (Lepidoptera, Papilionidae)". ''Journal of the East Africa Natural History Society'
pdf
Key to East Africa members of the species group, diagnostic and other notes and figures. (Permission to host granted by The East Africa Natural History Society leonidas Butterflies of Africa Butterflies described in 1793 Taxa named by Johan Christian Fabricius {{Papilionidae-stub