Graphania Averilla
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''Ichneutica averilla'' is a
moth Moths are a group of insects that includes all members of the order Lepidoptera that are not Butterfly, butterflies. They were previously classified as suborder Heterocera, but the group is Paraphyly, paraphyletic with respect to butterflies (s ...
of the family
Noctuidae The Noctuidae, commonly known as owlet moths, cutworms or armyworms, are a family (biology), family of moths. Taxonomically, they are considered the most controversial family in the superfamily Noctuoidea because many of the clades are constantly ...
. It is
endemic Endemism is the state of a species being found only in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also foun ...
to
New Zealand New Zealand () is an island country in the southwestern Pacific Ocean. It consists of two main landmasses—the North Island () and the South Island ()—and List of islands of New Zealand, over 600 smaller islands. It is the List of isla ...
. This species is found in the
North Island The North Island ( , 'the fish of Māui', historically New Ulster) is one of the two main islands of New Zealand, islands of New Zealand, separated from the larger but less populous South Island by Cook Strait. With an area of , it is the List ...
at
Mount Taranaki Mount Taranaki (), officially Taranaki Maunga and also known as Mount Egmont, is a dormant stratovolcano in the Taranaki region on the west coast of New Zealand's North Island. At , it is the second highest mountain in the North Island, afte ...
but is widespread throughout the
South Island The South Island ( , 'the waters of Pounamu, Greenstone') is the largest of the three major islands of New Zealand by surface area, the others being the smaller but more populous North Island and Stewart Island. It is bordered to the north by ...
and
Stewart Island Stewart Island (, ' glowing skies', officially Stewart Island / Rakiura, formerly New Leinster) is New Zealand's third-largest island, located south of the South Island, across Foveaux Strait. It is a roughly triangular island with a la ...
. It prefers mountainous habitat but can be found down to sea level in the southern parts of the South Island. Adults of the species are on the wing between November and March. Larvae likely exist on a variety of herbaceous plants but have been recorded as feeding on species within the genus ''
Plantago ''Plantago'' is a genus of about 200 species of flowering plants in the family Plantaginaceae, commonly called plantains or fleaworts. The common name plantain is shared with the unrelated cooking plantain. Most are herbaceous plants, though a ...
.'' This species is sometimes confused with ''I. mutas'' but can be distinguished from the latter on the basis of forewing colour (''I. averilla'' has a forewing shaded a uniform red-ochreous colour) as well as the absence of or an indistinct antemedian forewing line.


Taxonomy

''Ichneutica averilla'' was first described by
George Hudson George Hudson (probably 10 March 1800 – 14 December 1871) was an English railway financier and politician who, because he controlled a significant part of the Railway Mania, railway network in the 1840s, became known as "The Railway King"—a ...
in 1921 under the name ''Melanchra averill''. Hudson used the two female specimens of the moth collected by
Averil Lysaght Averil Margaret Lysaght (14 April 1905 – 21 August 1981) was a New Zealand biologist, science historian and artist, best known for her scholarly work on Joseph Banks. Early life Lysaght was born in Mokoia, Taranaki, New Zealand on 14 April 1 ...
in December 1920 on
Mount Taranaki Mount Taranaki (), officially Taranaki Maunga and also known as Mount Egmont, is a dormant stratovolcano in the Taranaki region on the west coast of New Zealand's North Island. At , it is the second highest mountain in the North Island, afte ...
. The
lectotype In biology, a type is a particular specimen (or in some cases a group of specimens) of an organism to which the scientific name of that organism is formally associated. In other words, a type is an example that serves to anchor or centralizes ...
specimen of this species is held at the
Museum of New Zealand Te Papa Tongarewa The Museum of New Zealand Te Papa Tongarewa is New Zealand's national museum and is located in Wellington. Usually known as Te Papa ( Māori for ' the treasure box'), it opened in 1998 after the merging of the National Museum of New Zealand ...
. In 1924
Alfred Philpott Alfred Philpott (15 December 1870 – 24 July 1930) was a New Zealand museum curator, entomologist and writer. He was born in Tysoe, Warwickshire Warwickshire (; abbreviated Warks) is a Ceremonial counties of England, ceremonial county in ...
, thinking this was a new species, described this moth under the name ''Melanchra furtiva'' from specimens collected around Mount Arthur as well as in the mountainous areas near
Lake Wakatipu Lake Wakatipu () is an inland lake (finger lake) in the South Island of New Zealand. It is in the southwest corner of the Otago region, near its boundary with Southland, New Zealand, Southland. ''Lake Wakatipu'' comes from the original Māori l ...
. In 1988
John S. Dugdale John Stewart Dugdale (5 April 1934 – 4 September 2020) was a New Zealand entomologist known for his contributions to knowledge about New Zealand lepidoptera, as well as tachinid flies and cicadas The cicadas () are a superfamily, the ...
placed this species within the genus ''Graphania'' and recognised ''Melanchra furtiva'' as a new synonym. In 2019
Robert Hoare The name Robert is an ancient Germanic given name, from Proto-Germanic "fame" and "bright" (''Hrōþiberhtaz''). Compare Old Dutch ''Robrecht'' and Old High German ''Hrodebert'' (a compound of '' Hruod'' () "fame, glory, honour, praise, reno ...
undertook a major review of New Zealand
Noctuidae The Noctuidae, commonly known as owlet moths, cutworms or armyworms, are a family (biology), family of moths. Taxonomically, they are considered the most controversial family in the superfamily Noctuoidea because many of the clades are constantly ...
. During this review the genus ''Ichneutica'' was greatly expanded and the genus ''Graphania'' was subsumed into that genus as a synonym. As a result of this review, this species is now known as ''Ichneutica averilla.''


Description

Hudson described the larvae of this species as follows: Hudson, in his original description, described the female adult of the species as follows: ''I. averilla'' is a medium-sized moth. The adult male of the species has a wingspan of between 33 and 41 mm whereas the female has a wingspan of between 36 and 41 mm. The forewings of this species are pinkish brown coloured, tinged with grey and have dark markings with a notable streak running lengthwise from the forewing base. There is also a distinctive v-shaped dark mark on the posterior corner of the wing. The hind wings of this species are greyish brown in colour. This species can be confused with ''I. mutans'' but can be distinguished on forewing colour as ''I. averilla'' has a ground colour of uniform red-ochreous shade where as ''I. mutans'' is more brown with blackish hints. Also the antemedian forewing line is normally distinct in ''I. mutans'' where as in ''I. averilla'' it is absent or just a hint of pale scaling is present.


Distribution

This species is endemic to New Zealand. It is found in the North Island, but only in the Mount Taranaki area. This species is widespread throughout the South Island, where it has been collected in such areas as on Mount Arthur, around
Ben Lomond Ben Lomond (, ), , is a mountain in the Scottish Highlands. Situated on the eastern shore of Loch Lomond, it is the most southerly of the Munros. Ben Lomond lies within the Ben Lomond National Memorial Park and the Loch Lomond and The Trossach ...
, Elfin Bay, Lake Luna and Lake MacKenzie in
Otago Otago (, ; ) is a regions of New Zealand, region of New Zealand located in the southern half of the South Island and administered by the Otago Regional Council. It has an area of approximately , making it the country's second largest local go ...
, and at the Cass Basin in the
Waimakariri River The Waimakariri River is one of the largest rivers in Canterbury, on the eastern coast of New Zealand's South Island. It flows for in a generally southeastward direction from the Southern Alps across the Canterbury Plains to the Pacific Ocean. ...
catchment. ''I. averilla'' is also present on Stewart Island.


Habitat

''I. averilla'' inhabits mountainous country. However it can be found down to sea level altitudes in the southern parts of the South Island, such as in the
Catlins The Catlins (sometimes referred to as The Catlins Coast) comprise an area in the southeastern corner of the South Island of New Zealand. The area lies between Balclutha, New Zealand, Balclutha and Invercargill, straddling the boundary between ...
.


Behaviour

The adult moths can be found on the wing from November to March.


Life history and host plants

After emerging the larvae eat their egg shell. The larvae of ''I. averilla'' have been found to feed on plants in the genus ''
Plantago ''Plantago'' is a genus of about 200 species of flowering plants in the family Plantaginaceae, commonly called plantains or fleaworts. The common name plantain is shared with the unrelated cooking plantain. Most are herbaceous plants, though a ...
'' and it is likely the larvae of this species feeds on a variety of plant hosts.


References

{{Taxonbar, from=Q104173024 Hadeninae Moths of New Zealand Endemic fauna of New Zealand Moths described in 1921 Taxa named by George Hudson Endemic moths of New Zealand