''Granastrapotherium'' is an extinct

genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...

ungulate Ungulates ( ) are members of the diverse clade Euungulata ("true ungulates"), which primarily consists of large mammals with Hoof, hooves. Once part of the clade "Ungulata" along with the clade Paenungulata, "Ungulata" has since been determined ...

mammals, described from remains found in rocks of the Honda Group in the Tatacoa Desert, in the

Colombia Colombia, officially the Republic of Colombia, is a country primarily located in South America with Insular region of Colombia, insular regions in North America. The Colombian mainland is bordered by the Caribbean Sea to the north, Venezuel ...

n departments of Huila and Tolima, at the

Miocene The Miocene ( ) is the first epoch (geology), geological epoch of the Neogene Period and extends from about (Ma). The Miocene was named by Scottish geologist Charles Lyell; the name comes from the Greek words (', "less") and (', "new") and mea ...

fossil site

La Venta La Venta is a pre-Columbian archaeological site of the Olmec civilization located in the present-day Mexican state of Tabasco. Some of the artifacts have been moved to the museum "Parque - Museo de La Venta", which is in nearby Villaherm ...

. The only species formally recognized is ''Granastrapotherium snorki'' (from

Spanish Spanish might refer to: * Items from or related to Spain: **Spaniards are a nation and ethnic group indigenous to Spain **Spanish language, spoken in Spain and many countries in the Americas **Spanish cuisine **Spanish history **Spanish culture ...

, ''gran'', "great"; ''

Astrapotherium ''Astrapotherium'' ("lightning beast") is an extinct genus of large Astrapotheria, astrapotherian ungulate native to South America during the early-middle Miocene. It is the best known member of the group. The type species. ''A. magnus'' have bee ...

'', "lightning beast"; and snorkel, breathing tube, in reference to the trunk). Remains found in

Bolivia Bolivia, officially the Plurinational State of Bolivia, is a landlocked country located in central South America. The country features diverse geography, including vast Amazonian plains, tropical lowlands, mountains, the Gran Chaco Province, w ...

and

Peru Peru, officially the Republic of Peru, is a country in western South America. It is bordered in the north by Ecuador and Colombia, in the east by Brazil, in the southeast by Bolivia, in the south by Chile, and in the south and west by the Pac ...

, seem to belong to ''Granastrapotherium'' or a very similar animal.

Description

This

astrapothere Astrapotheria is an extinct Order (biology), order of South American and Antarctic ungulate, hoofed mammals that existed from the late Paleocene to the Middle Miocene, ."The uruguaytheriine Astrapotheriidae from the rich middle Miocene Honda Grou ...

differs from its coeval, the uruguaytheriine astrapotheriid ''

Xenastrapotherium ''Xenastrapotherium'' is an extinct genus of astrapothere, a type of hoofed herbivorous mammal, native to South America, which lived in the Middle to Late Miocene period, typically during the Laventan stage. It is a member of the family Astrap ...

'' by their larger size, between 3 and 4 tonnes, with

tusk Tusks are elongated, continuously growing front teeth that protrude well beyond the mouth of certain mammal species. They are most commonly canine tooth, canine teeth, as with Narwhal, narwhals, chevrotains, musk deer, water deer, muntjac, pigs, ...

s about one meter long, making it one of the largest representatives of Astrapotheria, only surpassed by some species of ''

Parastrapotherium ''Parastrapotherium'' is an extinct genus of South American land mammal that existed from the Late Oligocene (Deseadan SALMA) to the Early Miocene (Colhuehuapian SALMA). The genus includes some of the largest and smallest known astrapotherians, ...

''. Other differences include the presence of only one

premolar The premolars, also called premolar Tooth (human), teeth, or bicuspids, are transitional teeth located between the Canine tooth, canine and Molar (tooth), molar teeth. In humans, there are two premolars per dental terminology#Quadrant, quadrant in ...

, the lack of incisors in both jaws and the disposition of the

canine tooth In mammalian oral anatomy, the canine teeth, also called cuspids, dogteeth, eye teeth, vampire teeth, or fangs, are the relatively long, pointed teeth. In the context of the upper jaw, they are also known as '' fangs''. They can appear more f ...

, which are very large and horizontal, which reminds much less of those of

hippo The hippopotamus (''Hippopotamus amphibius;'' ; : hippopotamuses), often shortened to hippo (: hippos), further qualified as the common hippopotamus, Nile hippopotamus and river hippopotamus, is a large semiaquatic Mammal, mammal native to su ...

s and more of the tusks of some ancient relatives of elephants (such as ''

Palaeomastodon ''Palaeomastodon'' ("ancient mastodon") is an extinct genus of basal proboscideans from the Oligocene of North Africa. The first specimen discovered was recovered from strata belonging to the Fayum fossil deposits of Egypt. It was described and ...

''), although the tusks in

elephant Elephants are the largest living land animals. Three living species are currently recognised: the African bush elephant ('' Loxodonta africana''), the African forest elephant (''L. cyclotis''), and the Asian elephant ('' Elephas maximus ...

s and their relatives are not formed by the canines but the incisors. Similarly, the large nostrils appear extremely withdrawn on the skull, so this creature had to have a larger trunk than other astrapotheres. Most likely, like elephants, this animal used its muscular

proboscis A proboscis () is an elongated appendage from the head of an animal, either a vertebrate or an invertebrate. In invertebrates, the term usually refers to tubular arthropod mouthparts, mouthparts used for feeding and sucking. In vertebrates, a pr ...

together with its tusks to cut leaves off trees and shrubs.Johnson, Steven C. & Madden, Richard H. 1997. Uruguaytheriinae Astrapotheres of Tropical South America. Chapter 22 in “Vertebrate Paleontology in the Neotropics. The Miocene Fauna of La Venta, Colombia”. Edited by Richard F. Kay, Richard H. Madden, Richard L. Cifelli, and John J. Flynn. Smithsonian Institution Press. Washington and London.

Phylogeny

Cladogram A cladogram (from Greek language, Greek ''clados'' "branch" and ''gramma'' "character") is a diagram used in cladistics to show relations among organisms. A cladogram is not, however, an Phylogenetic tree, evolutionary tree because it does not s ...

based in the

phylogenetic analysis In biology, phylogenetics () is the study of the evolutionary history of life using observable characteristics of organisms (or genes), which is known as phylogenetic inference. It infers the relationship among organisms based on empirical data ...

published by Vallejo-Pareja ''et al''., 2015, showing the position of ''Granastrapotherium'':

References

External links

Website of the Berkeley University about the fauna of La Venta, including ''Granastrapotherium''

"Snorki the giant's friends and relatives" in the blog Tetrapod Zoology

{{Taxonbar, from1=Q2888784, from2=Q28563695 Astrapotheriidae Miocene mammals of South America Laventan Neogene Colombia Fossils of Colombia Honda Group, Colombia Fossil taxa described in 1997 Prehistoric placental genera