Granary Weevil
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The wheat weevil (''Sitophilus granarius''), also known as the grain weevil or granary weevil, is an insect that feeds on cereal grains, and is a common pest in many places. It can cause significant damage to harvested stored grains, and may drastically decrease crop yields. The females lay many eggs, and the larvae eat the inside of the grain kernels.


Identification

Adult wheat weevils are about long, with elongated snouts and chewing mouth parts. Depending on the grain kernels, the size of the weevil varies. In small grains, such as millet or grain sorghum, they are small in size, but in maize (corn), they are larger. The adults are a reddish-brown colour and lack distinguishing marks. Adult wheat weevils are not capable of flight. Larvae are legless, humpbacked, and are white with a tan head. Weevils in the pupal stage have snouts similar to those of adults.


Natural history


Life cycle

Female wheat weevils lay between 36 and 254 eggs; usually, one egg is deposited in each grain kernel. All larval stages and the pupal stage occur within the grain. The larvae feed inside the grain until pupation, after which they bore a hole out of the grain and emerge. They are rarely seen outside of the grain kernel. The life cycle takes about five weeks in the summer, but may take up to 20 weeks in cooler temperatures. Adults can live up to eight months after emerging. File:Pupa of Sitophilus granarius (L.) inside a wheat kernel (cropped).jpg,
Pupa A pupa (; : pupae) is the life stage of some insects undergoing transformation between immature and mature stages. Insects that go through a pupal stage are holometabolous: they go through four distinct stages in their life cycle, the stages th ...
inside a
wheat Wheat is a group of wild and crop domestication, domesticated Poaceae, grasses of the genus ''Triticum'' (). They are Agriculture, cultivated for their cereal grains, which are staple foods around the world. Well-known Taxonomy of wheat, whe ...
kernel File:Sitophilus granarius (Linné, 1758).jpg, Adult, dorsal view


Behavior

When threatened or disturbed, adult wheat weevils will pull their legs close to their bodies and feign death. Female weevils can tell if a grain kernel has had an egg laid in it by another weevil. They avoid laying another egg in this grain. Females chew a hole, deposit an egg, and seal the hole with a gelatinous secretion. This may be how other females know the grain has an egg in it already. This ensures the young will survive and produce another generation. One pair of weevils may produce up to 6,000 offspring per year. The species is not found "in the wild", but only in human food-storage situations.


Human impact

Wheat weevils are pests of stores of grains of the cereal crops
wheat Wheat is a group of wild and crop domestication, domesticated Poaceae, grasses of the genus ''Triticum'' (). They are Agriculture, cultivated for their cereal grains, which are staple foods around the world. Well-known Taxonomy of wheat, whe ...
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oat The oat (''Avena sativa''), sometimes called the common oat, is a species of cereal grain grown for its seed, which is known by the same name (usually in the plural). Oats appear to have been domesticated as a secondary crop, as their seeds ...
s,
rye Rye (''Secale cereale'') is a grass grown extensively as a grain, a cover crop and a forage crop. It is grown principally in an area from Eastern and Northern Europe into Russia. It is much more tolerant of cold weather and poor soil than o ...
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barley Barley (), a member of the grass family, is a major cereal grain grown in temperate climates globally. It was one of the first cultivated grains; it was domesticated in the Fertile Crescent around 9000 BC, giving it nonshattering spikele ...
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rice Rice is a cereal grain and in its Domestication, domesticated form is the staple food of over half of the world's population, particularly in Asia and Africa. Rice is the seed of the grass species ''Oryza sativa'' (Asian rice)—or, much l ...
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maize Maize (; ''Zea mays''), also known as corn in North American English, is a tall stout grass that produces cereal grain. It was domesticated by indigenous peoples in southern Mexico about 9,000 years ago from wild teosinte. Native American ...
. Unfortunately the impact of wheat weevils worldwide is unknown, because information is not well shared. It is believed to be especially bad in places where the grain harvests are not accurately measured. The weevils are hard to detect and usually all of the grain in an infested storage facility must be destroyed. Many methods have been attempted to get rid of the wheat weevil, such as pesticides, different methods of masking the odor of the grain with unpleasant scents, and some have even gone as far as introducing predator organisms.Giacinto, G. S., Antonio, D. C., & Giuseppe, R. 2008. Behavioral responses of adult ''Sitophilus granarius'' to individual cereal volatiles. Journal of Chemical Ecology, 34: 523-529.


Prevention and control

Sanitation and inspection are key to preventing infestation. Grains should be stored in preferably metallic (cardboard, even fortified, is easily drilled through by the weevil) containers with tight lids in a refrigerator or a freezer, and should be purchased in small quantities. If any suspicion has arisen, carefully examine the grains for adult insects or holes in the grain kernels. Another method is to immerse them in water. If they float to the surface, it is a good indication of infestation. Even if identified early, disposal may be the only effective solution.


See also

* '' Lixus concavus'', the rhubarb curculio weevil *
Maize weevil The maize weevil (''Sitophilus zeamais''), known in the United States as the greater rice weevil, is a species of beetle in the family Curculionidae. It can be found in numerous tropical areas around the world, and in the United States, and is a m ...
(''Sitophilus zeamais'') *
Rice weevil The rice weevil (''Sitophilus oryzae'') is a stored product Pest (organism), pest which attacks seeds of several crops, including wheat, rice, and maize. Description The adults are usually between long, with a long snout. The body color appears ...
(''Sitophilus oryzae'')


References

{{Authority control Dryophthorinae Agricultural pest insects Beetles described in 1758 Taxa named by Carl Linnaeus Beetles of Africa Storage pests Insect pests of wheat