Grammistops
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The ocellate soapfish (''Grammistops ocellatus'') is a species of marine
ray-finned fish Actinopterygii (; ), members of which are known as ray-finned fish or actinopterygians, is a class of bony fish that comprise over 50% of living vertebrate species. They are so called because of their lightly built fins made of webbings of sk ...
with a wide
Indo-Pacific The Indo-Pacific is a vast biogeographic region of Earth. In a narrow sense, sometimes known as the Indo-West Pacific or Indo-Pacific Asia, it comprises the tropical waters of the Indian Ocean, the western and central Pacific Ocean, and the ...
distribution. It is the only species in the genus ''Grammistops''. It is also known as the ocellated soapfish, ocellated podge, or false-eyed soapfish. The
specific name Specific name may refer to: * in Database management systems, a system-assigned name that is unique within a particular database In taxonomy, either of these two meanings, each with its own set of rules: * Specific name (botany), the two-part (bino ...
''ocellatus'' refers to the ocellate (eye-like) spot on the operculum (gill covering).


Taxonomy

''Grammistops ocellatus'' was first formally described in 1953 by the
American American(s) may refer to: * American, something of, from, or related to the United States of America, commonly known as the "United States" or "America" ** Americans, citizens and nationals of the United States of America ** American ancestry, p ...
ichthyologist Ichthyology is the branch of zoology devoted to the study of fish, including bony fish (Osteichthyes), cartilaginous fish (Chondrichthyes), and jawless fish (Agnatha). According to FishBase, 35,800 species of fish had been described as of March 2 ...
Leonard Peter Schultz Leonard Peter Schultz (1901–1986) was an American ichthyologist. Biography Schultz was born in 1901, at Albion, Michigan. He received education on ichthyology at Albion College, in which he got his bachelor's degree, in 1924. In 1926, he got ...
(1901–1986) with the type locality given as the Lagoon at Arji Island,
Bikini Atoll Bikini Atoll ( or ; Marshallese language, Marshallese: , , ), known as Eschscholtz Atoll between the 19th century and 1946, is a coral reef in the Marshall Islands consisting of 23 islands surrounding a central lagoon. The atoll is at the no ...
in the
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. It is related to the
grouper Groupers are a diverse group of marine ray-finned fish in the family Epinephelidae, in the order Perciformes. Groupers were long considered a subfamily of the seabasses in Serranidae, but are now treated as distinct. Not all members of this f ...
s, and like them this species is classified within the
subfamily In biological classification, a subfamily (Latin: ', plural ') is an auxiliary (intermediate) taxonomic rank, next below family but more inclusive than genus. Standard nomenclature rules end botanical subfamily names with "-oideae", and zo ...
Epinephelinae Groupers are a diverse group of marine Actinopterygii, ray-finned fish in the family Epinephelidae, in the order Perciformes. Groupers were long considered a subfamily of the seabasses in Serranidae, but are now treated as distinct. Not all m ...
of the
family Family (from ) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). It forms the basis for social order. Ideally, families offer predictabili ...
Serranidae Serranidae is a large family (biology), family of fishes belonging to the order Perciformes. The family contains about 450 species in 65 genera, including the sea basses and the groupers (subfamily Epinephelinae). Although many species are small, ...
. It is often considered part of a separate subfamily, Grammistinae (the soapfishes). , it is the only species in the genus ''Grammistops''.


Description

The ocellate soapfish has a body which is moderately elongate with a depth which is less than the length of the head. The body is dull yellowish-brown body and it has similarly coloured fins with a single dark, ocellate (eye-like) spot on the gill cover and a dark brown spot on either side of the chin. The
dorsal fin A dorsal fin is a fin on the back of most marine and freshwater vertebrates. Dorsal fins have evolved independently several times through convergent evolution adapting to marine environments, so the fins are not all homologous. They are found ...
contains 7 spines and 12–13 soft rays while the
anal fin Fins are moving appendages protruding from the body of fish that interact with water to generate thrust and help the fish swim. Apart from the tail or caudal fin, fish fins have no direct connection with the back bone and are supported o ...
has 3 spines and 9 soft rays. It attains a maximum total length of . The preopercle has a smooth margin with 1 or 2 short spines and the
caudal fin Fins are moving appendages protruding from the body of fish that interact with water to generate thrust and help the fish swim. Apart from the tail or caudal fin, fish fins have no direct connection with the back bone and are supported only ...
is rounded.


Distribution

''Grammistops ocellatus'' has a disjunct Indo-Pacific distribution. It is found in the Indian Ocean off the coast of East Africa from Kenya south to Mozambique and
Phuket Phuket (; , , or ''Tongkah'') is one of the Southern Thailand, southern Provinces of Thailand, provinces (''changwat'') of Thailand. It consists of the island of Phuket, List of islands of Thailand, the country's largest island, and another 3 ...
in Thailand and
Christmas Island Christmas Island, officially the Territory of Christmas Island, is an States and territories of Australia#External territories, Australian external territory in the Indian Ocean comprising the island of the same name. It is about south o ...
. In the Pacific Ocean it is found from eastern Indonesia east to the
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, north to the
Ryukyu Islands The , also known as the or the , are a chain of Japanese islands that stretch southwest from Kyushu to Geography of Taiwan, Taiwan: the Ryukyu Islands are divided into the Satsunan Islands (Ōsumi Islands, Ōsumi, Tokara Islands, Tokara and A ...
of southern Japan and south as far as Australia,
New Caledonia New Caledonia ( ; ) is a group of islands in the southwest Pacific Ocean, southwest of Vanuatu and east of Australia. Located from Metropolitan France, it forms a Overseas France#Sui generis collectivity, ''sui generis'' collectivity of t ...
and Tonga.


Habitat and biology

The ocellate soapfish is found on coral heads in lagoons and seaward reefs at depths of . It is a cryptic species which is encountered either solitarily or as pairs. Like other species named "soapfish" this species secretes a skin toxin from epidermal glandular cells when stressed.


References

{{Taxonbar, from=Q3541245 Grammistini Fish described in 1953