Gordon Arthur Riley (1 June 1911 – 7 October 1985) was an American biological oceanographer most associated with his studies of the dynamics of

plankton Plankton are the diverse collection of organisms that drift in Hydrosphere, water (or atmosphere, air) but are unable to actively propel themselves against ocean current, currents (or wind). The individual organisms constituting plankton are ca ...

ecosystem An ecosystem (or ecological system) is a system formed by Organism, organisms in interaction with their Biophysical environment, environment. The Biotic material, biotic and abiotic components are linked together through nutrient cycles and en ...

Early life and education

Born in

Webb City, Missouri Webb City is a city in Jasper County, Missouri, United States. The population was 13,031 at the 2020 census. It is part of the Joplin, Missouri Metropolitan Statistical Area. Webb City also has a police department, a fire department, and animal ...

on June 1, 1911, Riley was educated within the state at

Drury College Drury University, formerly Drury College and originally Springfield College, is a private university in Springfield, Missouri, United States. The university's mission statement describes itself as "church-related". It enrolls about 1,590 undergr ...

and

Washington University in St. Louis Washington University in St. Louis (WashU) is a private research university in St. Louis, Missouri, United States. Founded in 1853 by a group of civic leaders and named for George Washington, the university spans 355 acres across its Danforth ...

, graduating with a MS in

embryology Embryology (from Ancient Greek, Greek ἔμβρυον, ''embryon'', "the unborn, embryo"; and -λογία, ''-logy, -logia'') is the branch of animal biology that studies the Prenatal development (biology), prenatal development of gametes (sex ...

. He moved to

Yale University Yale University is a Private university, private Ivy League research university in New Haven, Connecticut, United States. Founded in 1701, Yale is the List of Colonial Colleges, third-oldest institution of higher education in the United Stat ...

in 1934, intending to work with the

anatomist Anatomy () is the branch of morphology concerned with the study of the internal structure of organisms and their parts. Anatomy is a branch of natural science that deals with the structural organization of living things. It is an old scien ...

Ross Harrison, but instead became interested in

limnology Limnology ( ; ) is the study of inland aquatic ecosystems. It includes aspects of the biological, chemical, physical, and geological characteristics of fresh and saline, natural and man-made bodies of water. This includes the study of lakes, ...

. Working with the

ecologist Ecology () is the natural science of the relationships among living organisms and their environment. Ecology considers organisms at the individual, population, community, ecosystem, and biosphere levels. Ecology overlaps with the closely re ...

G. Evelyn Hutchinson, he completed his

doctoral thesis A thesis (: theses), or dissertation (abbreviated diss.), is a document submitted in support of candidature for an academic degree or professional qualification presenting the author's research and findings.International Standard ISO 7144: D ...

on the

copper Copper is a chemical element; it has symbol Cu (from Latin ) and atomic number 29. It is a soft, malleable, and ductile metal with very high thermal and electrical conductivity. A freshly exposed surface of pure copper has a pinkish-orang ...

cycle Cycle, cycles, or cyclic may refer to: Anthropology and social sciences * Cyclic history, a theory of history * Cyclical theory, a theory of American political history associated with Arthur Schlesinger, Sr. * Social cycle, various cycles in ...

of lakes in

Connecticut Connecticut ( ) is a U.S. state, state in the New England region of the Northeastern United States. It borders Rhode Island to the east, Massachusetts to the north, New York (state), New York to the west, and Long Island Sound to the south. ...

. Subsequently, he continued to be interested in the

productivity Productivity is the efficiency of production of goods or services expressed by some measure. Measurements of productivity are often expressed as a ratio of an aggregate output to a single input or an aggregate input used in a production proce ...

of lakes, but gradually expanded his studies to encompass

salt water Saline water (more commonly known as salt water) is water that contains a high concentration of dissolved salts (mainly sodium chloride). On the United States Geological Survey (USGS) salinity scale, saline water is saltier than brackish wate ...

, ultimately moving into biological oceanography.

Career

Riley's oceanographic work focused on the influences affecting the population ecology of plankton systems in

coastal A coast (coastline, shoreline, seashore) is the land next to the sea or the line that forms the boundary between the land and the ocean or a lake. Coasts are influenced by the topography of the surrounding landscape and by aquatic erosion, su ...

and

open ocean The pelagic zone consists of the water column of the open ocean and can be further divided into regions by depth. The word ''pelagic'' is derived . The pelagic zone can be thought of as an imaginary cylinder or water column between the surfa ...

waters. His early work

correlated In statistics, correlation or dependence is any statistical relationship, whether causal or not, between two random variables or bivariate data. Although in the broadest sense, "correlation" may indicate any type of association, in statistic ...

phytoplankton Phytoplankton () are the autotrophic (self-feeding) components of the plankton community and a key part of ocean and freshwater Aquatic ecosystem, ecosystems. The name comes from the Greek language, Greek words (), meaning 'plant', and (), mea ...

production Production may refer to: Economics and business * Production (economics) * Production, the act of manufacturing goods * Production, in the outline of industrial organization, the act of making products (goods and services) * Production as a stat ...

with regulating factors such as

nutrient A nutrient is a substance used by an organism to survive, grow and reproduce. The requirement for dietary nutrient intake applies to animals, plants, fungi and protists. Nutrients can be incorporated into cells for metabolic purposes or excret ...

light Light, visible light, or visible radiation is electromagnetic radiation that can be visual perception, perceived by the human eye. Visible light spans the visible spectrum and is usually defined as having wavelengths in the range of 400– ...

and zooplankton abundance. From this

empirical Empirical evidence is evidence obtained through sense experience or experimental procedure. It is of central importance to the sciences and plays a role in various other fields, like epistemology and law. There is no general agreement on how t ...

base he went on to develop

ecosystem model An ecosystem model is an abstract, usually mathematical, representation of an ecological system (ranging in scale from an individual population, to an ecological community, or even an entire biome), which is studied to better understand the r ...

s that explained the

annual cycle Annual may refer to: *Annual publication, periodical publications appearing regularly once per year **Yearbook A yearbook, also known as an annual, is a type of Annual publication, a book published annually. One use is to record, highlight, an ...

of plankton ecosystems, most notably in his analysis of the

Georges Bank Georges Bank (formerly known as St. Georges Bank) is a large elevated area of the sea floor between Cape Cod, Massachusetts (United States), and Cape Sable Island, Nova Scotia (Canada). It separates the Gulf of Maine from the Atlantic Ocean. ...

region. His final publication concerned patchiness in plankton, the potential role of

diel vertical migration Diel vertical migration (DVM), also known as diurnal vertical migration, is a pattern of movement used by some organisms, such as copepods, living in the ocean and in lakes. The adjective "diel" ( IPA: , ) comes from , and refers to a 24-hour ...

in this, and reflected on what this implied for plankton modelling studies, including his own 1946 study. After an extended period at Yale, in 1965 Riley moved to become a professor, and the director, at the Institute of Oceanography at

Dalhousie University Dalhousie University (commonly known as Dal) is a large public research university in Nova Scotia, Canada, with three campuses in Halifax, Nova Scotia, Halifax, a fourth in Bible Hill, Nova Scotia, Bible Hill, and a second medical school campus ...

. Much of his work continued to be in

collaboration Collaboration (from Latin ''com-'' "with" + ''laborare'' "to labor", "to work") is the process of two or more people, entities or organizations working together to complete a task or achieve a goal. Collaboration is similar to cooperation. The ...

with researchers at the

Woods Hole Oceanographic Institution The Woods Hole Oceanographic Institution (WHOI, acronym pronounced ) is a private, nonprofit research and higher education facility dedicated to the study of marine science and engineering. Established in 1930 in Woods Hole, Massachusetts, it i ...

. Towards the end of his life, Riley wrote a candid

autobiography An autobiography, sometimes informally called an autobio, is a self-written account of one's own life, providing a personal narrative that reflects on the author's experiences, memories, and insights. This genre allows individuals to share thei ...

of his scientific life, in part to document the early days of

oceanography Oceanography (), also known as oceanology, sea science, ocean science, and marine science, is the scientific study of the ocean, including its physics, chemistry, biology, and geology. It is an Earth science, which covers a wide range of to ...

as a discipline.

References

External links

Autobiography

University of California San Diego The University of California, San Diego (UC San Diego in communications material, formerly and colloquially UCSD) is a public university, public Land-grant university, land-grant research university in San Diego, California, United States. Es ...

Biography

Obituary
''

Limnology and Oceanography Limnology ( ; ) is the study of inland aquatic ecosystems. It includes aspects of the biological, chemical, physical, and geological characteristics of fresh and saline, natural and man-made bodies of water. This includes the study of lakes ...

'' 1911 births 1985 deaths American marine biologists American oceanographers Systems ecologists Washington University in St. Louis alumni 20th-century American zoologists Presidents of the Association for the Sciences of Limnology and Oceanography Yale University alumni {{US-scientist-stub