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''Pituophis'' is a
genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
of non venomous colubrid
snake Snakes are elongated limbless reptiles of the suborder Serpentes (). Cladistically squamates, snakes are ectothermic, amniote vertebrates covered in overlapping scales much like other members of the group. Many species of snakes have s ...
s, commonly referred to as gopher snakes, pine snakes, and bullsnakes, which are
endemic Endemism is the state of a species being found only in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also foun ...
to
North America North America is a continent in the Northern Hemisphere, Northern and Western Hemisphere, Western hemispheres. North America is bordered to the north by the Arctic Ocean, to the east by the Atlantic Ocean, to the southeast by South Ameri ...
. They are often yellow or cream in color with dark spots and a dark line across their face. Some species can exceed seven feet in length. Gopher snakes can live for 15 years. The gopher snake is commonly misidentified as a
rattlesnake Rattlesnakes are venomous snakes that form the genus, genera ''Crotalus'' and ''Sistrurus'' of the subfamily Crotalinae (the pit vipers). All rattlesnakes are vipers. Rattlesnakes are predators that live in a wide array of habitats, hunting sm ...
because of its similar coloration and its defensive behavior when feeling threatened. A scared gopher snake will flatten its head, hiss loudly, and shake its tail rapidly, doing a very convincing rattlesnake imitation.


Nomenclature

The genus name ''Pituophis'' is a Latinized modern scientific Greek compound Πιτυόφις : "pine snake"; from (pítus, "pine"), and (óphis, "snake").


Geographic range

Species A species () is often defined as the largest group of organisms in which any two individuals of the appropriate sexes or mating types can produce fertile offspring, typically by sexual reproduction. It is the basic unit of Taxonomy (biology), ...
and
subspecies In Taxonomy (biology), biological classification, subspecies (: subspecies) is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (Morphology (biology), morpholog ...
within the
genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
''Pituophis'' are found throughout
Mexico Mexico, officially the United Mexican States, is a country in North America. It is the northernmost country in Latin America, and borders the United States to the north, and Guatemala and Belize to the southeast; while having maritime boundar ...
, the Southern and
Western United States The Western United States (also called the American West, the Western States, the Far West, the Western territories, and the West) is List of regions of the United States, census regions United States Census Bureau. As American settlement i ...
and
Western Canada Western Canada, also referred to as the Western provinces, Canadian West, or Western provinces of Canada, and commonly known within Canada as the West, is a list of regions of Canada, Canadian region that includes the four western provinces and t ...
. Conant R (1975). ''A Field Guide to Reptiles and Amphibians of Eastern and Central North America, Second Edition''. Boston: Houghton Mifflin. (paperback). (Genus ''Pituophis'', pp. 198–201 + Plate 27 + figure 57 + Map 147).


Description

All species of ''Pituophis'' are large and powerfully built. The head is relatively small in proportion to the body and it is only slightly distinct from the neck. The
rostral Rostral may refer to: Anatomy * Rostral (anatomical term), situated toward the oral or nasal region * Rostral bone, in ceratopsian dinosaurs * Rostral organ, of certain fish * Rostral scale The rostral scale, or rostral, in snakes and other sca ...
is enlarged and elongated, imparting a characteristic somewhat pointed shape to the head. All the species occurring in the United States have four prefrontals instead of the usual two.


Modified epiglottis

In all snakes of the genus ''Pituophis'', the epiglottis is peculiarly modified so that it is thin, erect and flexible. When a stream of air is forced from the
trachea The trachea (: tracheae or tracheas), also known as the windpipe, is a cartilaginous tube that connects the larynx to the bronchi of the lungs, allowing the passage of air, and so is present in almost all animals' lungs. The trachea extends from ...
, the epiglottis vibrates, thereby producing the peculiarly loud, hoarse hissing for which bullsnakes, gopher snakes, and pine snakes are well known. Schmidt KP, Davis DD (1941). ''Field Book of Snakes of the United States and Canada''. New York: G.P. Putnam's Sons. 365 pp. (Genus ''Pituophis'', pp. 158–160).


Species and subspecies


References


Further reading

*


External links

* *Accounts of four subspecies, ''P. deppei deppei, P. deppei jani, P. lineaticollis lineaticollis'' and ''P. lineaticollis gibsoni'', are given in Duellman WE (1960)
"A Taxonomic Study of the Middle American Snake, ''Pituophis deppei'' ".
''University of Kansas Publications, Museum of Natural History'' 10: 599–610. {{Authority control Snake genera Snakes of North America Taxa named by John Edwards Holbrook