The gopher rockfish (''Sebastes carnatus''), also known as the gopher sea perch, is a

species A species () is often defined as the largest group of organisms in which any two individuals of the appropriate sexes or mating types can produce fertile offspring, typically by sexual reproduction. It is the basic unit of Taxonomy (biology), ...

of marine

ray-finned fish Actinopterygii (; ), members of which are known as ray-finned fish or actinopterygians, is a class of bony fish that comprise over 50% of living vertebrate species. They are so called because of their lightly built fins made of webbings of sk ...

belonging to the subfamily

Sebastinae Sebastinae is a subfamily of marine fish belonging to the Family (biology), family Scorpaenidae in the Order (biology), order Scorpaeniformes. Their common names include rockfishes, rock perches, ocean perches, sea perches, thornyheads, scorpion ...

, the rockfishes, part of the

family Family (from ) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). It forms the basis for social order. Ideally, families offer predictabili ...

Scorpaenidae The Scorpaenidae (also known as scorpionfish) are a family (biology), family of mostly ocean, marine fish that includes many of the world's most venomous species. As their name suggests, scorpionfish have a type of "sting" in the form of sharp ...

. It is found in the eastern

Pacific The Pacific Ocean is the largest and deepest of Earth's five oceanic divisions. It extends from the Arctic Ocean in the north to the Southern Ocean, or, depending on the definition, to Antarctica in the south, and is bounded by the cont ...

, primarily off

California California () is a U.S. state, state in the Western United States that lies on the West Coast of the United States, Pacific Coast. It borders Oregon to the north, Nevada and Arizona to the east, and shares Mexico–United States border, an ...

Taxonomy

The gopher rockfish was originally described in 1880 as ''Sebastichthys carnatus'' by the American

ichthyologists Ichthyology is the branch of zoology devoted to the study of fish, including bony fish (Osteichthyes), cartilaginous fish (Chondrichthyes), and jawless fish (Agnatha). According to FishBase, 35,800 species of fish had been described as of March 2 ...

David Starr Jordan David Starr Jordan (January 19, 1851 – September 19, 1931) was the founding president of Stanford University, serving from 1891 to 1913. He was an ichthyologist during his research career. Prior to serving as president of Stanford Universi ...

and

Charles Henry Gilbert Charles Henry Gilbert (December 5, 1859 in Rockford, Illinois – April 20, 1928 in Palo Alto, California) was a pioneer ichthyologist and Fisheries science, fishery biologist of particular significance to natural history of the western Unit ...

with the type locality given as the

Monterey Bay, California Monterey Bay is a bay of the Pacific Ocean located on the coast of the U.S. state of California, south of the San Francisco Bay Area. San Francisco itself is further north along the coast, by about 75 miles (120 km), accessible via CA 1 and U ...

. Some authorities place this species in the

subgenus In biology, a subgenus ( subgenera) is a taxonomic rank directly below genus. In the International Code of Zoological Nomenclature, a subgeneric name can be used independently or included in a species name, in parentheses, placed between the ge ...

''Pteropodus''. The

specific name Specific name may refer to: * in Database management systems, a system-assigned name that is unique within a particular database In taxonomy, either of these two meanings, each with its own set of rules: * Specific name (botany), the two-part (bino ...

''carnatus'' means "fleshy" or "flesh-colored", alluding to the background color of this fish.

Description

The gopher rockfish is a deep, stout bodied fish with a steep dorsal profile. The body is as deep as 34% to 38% of its

standard length Fish measurement is the measuring of individual fish and various parts of fish anatomy, their anatomies, for data used in many areas of ichthyology, including Taxonomy (biology), taxonomy and fishery biology. Overall length Standard length (SL) is ...

. They have many spines on the head and body. The overall color is dark brown, black, and greenish fading to reddish brown on the belly. There is a row of flesh-colored or whitish spotting and blotches on their back reaching up to and onto the

dorsal fin A dorsal fin is a fin on the back of most marine and freshwater vertebrates. Dorsal fins have evolved independently several times through convergent evolution adapting to marine environments, so the fins are not all homologous. They are found ...

and irregular pale patches on their flanks. There is a dark stripe running rearwards from the eye and another on the upper jaw. The head is of average length for this genus and has a short snout with a small terminal mouth and large eyes which bulge over the dorsal profile of the head. The

caudal fin Fins are moving appendages protruding from the body of fish that interact with water to generate thrust and help the fish swim. Apart from the tail or caudal fin, fish fins have no direct connection with the back bone and are supported only ...

is truncate. The dorsal fin has 13 spines and 12 to 14 rays while the

anal fin Fins are moving appendages protruding from the body of fish that interact with water to generate thrust and help the fish swim. Apart from the tail or caudal fin, fish fins have no direct connection with the back bone and are supported o ...

has 3 spines and 5 to 7 soft rays. This species grows to a maximum

total length Fish measurement is the measuring of individual fish and various parts of their anatomies, for data used in many areas of ichthyology, including taxonomy and fishery biology. Overall length Standard length (SL) is the length of a fish measured f ...

of and .

Distribution and habitat

Gopher rockfish are found in the northeastern Pacific Ocean off the Western coast of North America. They are known from as far north as Cape Blanco in

Oregon Oregon ( , ) is a U.S. state, state in the Pacific Northwest region of the United States. It is a part of the Western U.S., with the Columbia River delineating much of Oregon's northern boundary with Washington (state), Washington, while t ...

, down to

Ensenada Ensenada ("inlet") is a city in Ensenada Municipality, Baja California, situated on the Pacific Coast of Mexico. Located on Bahía de Todos Santos, the city had a population of 279,765 in 2018, making it the third-largest city in Baja Californ ...

in southern

Baja California Baja California, officially the Free and Sovereign State of Baja California, is a state in Mexico. It is the northwesternmost of the 32 federal entities of Mexico. Before becoming a state in 1952, the area was known as the North Territory of B ...

. They are commonest between Northern Baja California and

Northern California Northern California (commonly shortened to NorCal) is a geocultural region that comprises the northern portion of the U.S. state of California, spanning the northernmost 48 of the state's List of counties in California, 58 counties. Northern Ca ...

. It is a

demersal The demersal zone is the part of the sea or ocean (or deep lake) consisting of the part of the water column near to (and significantly affected by) the seabed and the benthos. The demersal zone is just above the benthic zone and forms a layer o ...

species that is encountered as solitary and highly territorial individuals with nearby shelters on rock structures or within

kelp forest Kelp forests are underwater areas with a high density of kelp, which covers a large part of the world's coastlines. Smaller areas of anchored kelp are called kelp beds. They are recognized as one of the most productive and dynamic ecosystems on E ...

s at depths from .

Biology and diet

The gopher rockfish is a

territorial A territory is an area of land, sea, or space, belonging or connected to a particular country, person, or animal. In international politics, a territory is usually a geographic area which has not been granted the powers of self-government, ...

species which defends a home territory on the seabed excluding other rockfish. It is a

nocturnal Nocturnality is a ethology, behavior in some non-human animals characterized by being active during the night and sleeping during the day. The common adjective is "nocturnal", versus diurnality, diurnal meaning the opposite. Nocturnal creatur ...

fish, spending the day sheltering in cavities and crevices. They leave their shelter at dusk to forage. The juveniles feed on planktonic crustaceans while the adults prey on

cephalopod A cephalopod is any member of the molluscan Taxonomic rank, class Cephalopoda (Greek language, Greek plural , ; "head-feet") such as a squid, octopus, cuttlefish, or nautilus. These exclusively marine animals are characterized by bilateral symm ...

gastropod Gastropods (), commonly known as slugs and snails, belong to a large Taxonomy (biology), taxonomic class of invertebrates within the phylum Mollusca called Gastropoda (). This class comprises snails and slugs from saltwater, freshwater, and fro ...

brittle star Brittle stars, serpent stars, or ophiuroids (; ; referring to the serpent-like arms of the brittle star) are echinoderms in the class Ophiuroidea, closely related to starfish. They crawl across the sea floor using their flexible arms for locomot ...

crabs Crabs are decapod crustaceans of the infraorder Brachyura (meaning "short tailed" in Greek), which typically have a very short projecting tail-like abdomen, usually hidden entirely under the thorax. Their exoskeleton is often thickened and ha ...

(notably the Cancrinae family),

shrimp A shrimp (: shrimp (American English, US) or shrimps (British English, UK)) is a crustacean with an elongated body and a primarily Aquatic locomotion, swimming mode of locomotion – typically Decapods belonging to the Caridea or Dendrobranchi ...

and

polychaete worm Polychaeta () is a paraphyletic class of generally marine annelid worms, commonly called bristle worms or polychaetes (). Each body segment has a pair of fleshy protrusions called parapodia that bear many bristles, called chaetae, which are ma ...

s. They also eat smaller fish such as juvenile rockfish, particularly blue rockfish (''Sebastes mystinus''),

sculpin A sculpin is a type of fish that belongs to the superfamily Cottoidea in the order Perciformes.Kane, E. A. and T. E. Higham. (2012)Life in the flow lane: differences in pectoral fin morphology suggest transitions in station-holding demand acros ...

s, juvenile

surfperch The surfperches are a family of percomorph fishes, the Embiotocidae. They are mainly found in northeast Pacific Ocean (as far south as Baja California), but a few species (genera '' Ditrema'' and '' Neoditrema'') are found in the northwest Pacifi ...

, kelpfishes, and

plainfin midshipman The plainfin midshipman (''Porichthys notatus'') is a species of bottom-dwelling Batrachoididae, batrachoid toadfish. It is a member of the midshipman fish, midshipman genus, ''Porichthys'', the only batrachoid fishes that have photophores. It is ...

(''Porichthys notatus''). Like other rockfish, it is a

viviparous In animals, viviparity is development of the embryo inside the body of the mother, with the maternal circulation providing for the metabolic needs of the embryo's development, until the mother gives birth to a fully or partially developed juve ...

species. spawning happens from January through July, and they give birth to live young after

gestation Gestation is the period of development during the carrying of an embryo, and later fetus, inside viviparous animals (the embryo develops within the parent). It is typical for mammals, but also occurs for some non-mammals. Mammals during pregn ...

. Compared to other

rockfish Rockfish is a common term for several species of fish, referring to their tendency to hide among rocks. The name rockfish is used for many kinds of fish used for food. This common name belongs to several groups that are not closely related, and ca ...

, gophers mature quicker, at 3-4 years of age, but the rate of maturity depends on the region. Males mature and grow faster than females. They are known to live for up to 30 years, possibly to 35, but it is rare for the fish to be older than 20.

Fisheries and relation to humans

Gopher rockfish are considered of moderate importance to

commercial fisheries Commercial fishing is the activity of catching fish and other seafood for commercial profit, mostly from wild fisheries. It provides a large quantity of food to many countries around the world, but those who practice it as an industry must often p ...

. They are mainly caught by recreational anglers fishing from shore for othe
near-shore rockfishes
and can be a relatively common catch, including from boats and kayaks. Larger fish are found in deeper waters. They are considered good eating fish, and may be sold live or frozen.

Genetics

A PCR-RFLP genetic sex marker has been identified for gopher rockfish, which can successfully distinguish males and females. The marker potentially also works in the closely related black-and-yellow rockfish, but it does not seem to successfully distinguish males and females in various other rockfish species. Chromosome 19 has been identified as a putative sex chromosome for the species.

References

* * Milton S. Love, Mary Yoklavich, Lyman K. Thorsteinson, (2002), ''The Rockfishes of the Northeast Pacific'', University of California Press, pp. 140–143 * Shawn Narum, Vincent Buonaccorsi, Carol Kimbrell, and Russell Vetter. (2004). Genetic Divergence between Gopher Rockfish (''Sebastes carnatus'') and Black and Yellow Rockfish (''Sebastes chrysomelas''). ''Copeia'', 4, pp. 926–931. {{Taxonbar, from=Q2717740 gopher rockfish Taxa named by David Starr Jordan Taxa named by Charles Henry Gilbert Fauna of California Western North American coastal fauna gopher rockfish