The Spanish flag (''Gonioplectrus hispanus'') is a species of marine

ray-finned fish Actinopterygii (; ), members of which are known as ray-finned fish or actinopterygians, is a class of bony fish that comprise over 50% of living vertebrate species. They are so called because of their lightly built fins made of webbings of sk ...

, a

grouper Groupers are a diverse group of marine ray-finned fish in the family Epinephelidae, in the order Perciformes. Groupers were long considered a subfamily of the seabasses in Serranidae, but are now treated as distinct. Not all members of this f ...

from the

subfamily In biological classification, a subfamily (Latin: ', plural ') is an auxiliary (intermediate) taxonomic rank, next below family but more inclusive than genus. Standard nomenclature rules end botanical subfamily names with "-oideae", and zo ...

Epinephelinae Groupers are a diverse group of marine Actinopterygii, ray-finned fish in the family Epinephelidae, in the order Perciformes. Groupers were long considered a subfamily of the seabasses in Serranidae, but are now treated as distinct. Not all m ...

which is part of the

family Family (from ) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). It forms the basis for social order. Ideally, families offer predictabili ...

Serranidae Serranidae is a large family (biology), family of fishes belonging to the order Perciformes. The family contains about 450 species in 65 genera, including the sea basses and the groupers (subfamily Epinephelinae). Although many species are small, ...

, which also includes the anthias and sea basses. It is found in the western Atlantic Ocean. It is the only species in the genus ''Gonioplectrus''.

Description

The Spanish flag has a body which is oblong with a large head, the length of the head is the same as the body's depth. The

standard length Fish measurement is the measuring of individual fish and various parts of fish anatomy, their anatomies, for data used in many areas of ichthyology, including Taxonomy (biology), taxonomy and fishery biology. Overall length Standard length (SL) is ...

is 2.3 to 2.5 times the depth of the body. The dorsal profile of the head and the intraorbital area are convex. They have a rounded preopercle which has a large, curved spine at its angle and fleshy lower edge. The central spine on the gill cover is and reaches past the rear of the opercular membrane. The

dorsal fin A dorsal fin is a fin on the back of most marine and freshwater vertebrates. Dorsal fins have evolved independently several times through convergent evolution adapting to marine environments, so the fins are not all homologous. They are found ...

contains 8 spines and 13 soft rays while the

anal fin Fins are moving appendages protruding from the body of fish that interact with water to generate thrust and help the fish swim. Apart from the tail or caudal fin, fish fins have no direct connection with the back bone and are supported o ...

has 3 spines and 7 soft rays. The membranes between the dorsal fin spines are deeplyncised and the spines have fleshy, scale covered bases. The

caudal fin Fins are moving appendages protruding from the body of fish that interact with water to generate thrust and help the fish swim. Apart from the tail or caudal fin, fish fins have no direct connection with the back bone and are supported only ...

is truncate. There are 47-49 scales in the

lateral line The lateral line, also called the lateral line organ (LLO), is a system of sensory organs found in fish, used to detect movement, vibration, and pressure gradients in the surrounding water. The sensory ability is achieved via modified epithelia ...

, which curves upwards underneath the soiny part of the dorsal in and then downwards below the soft-rayed part. The background colour of the head, body, dorsal and caudal fins is yellow and there are 6-7 pink stripes running horizontally from the head to the soft-rayed part of the dorsal fin and the caudal fin. There is a bright red blotch on the anterior part of the anal fin and a white blotch on the side of the belly. The

pelvic fin Pelvic fins or ventral fins are paired fins located on the ventral (belly) surface of fish, and are the lower of the only two sets of paired fins (the other being the laterally positioned pectoral fins). The pelvic fins are homologous to the hi ...

s are pinkish purple in colour. This species attains a maximum

total length Fish measurement is the measuring of individual fish and various parts of their anatomies, for data used in many areas of ichthyology, including taxonomy and fishery biology. Overall length Standard length (SL) is the length of a fish measured f ...

of .

Distribution

The Spanish flag occurs in the Western Atlantic Ocean in the

Gulf of Mexico The Gulf of Mexico () is an oceanic basin and a marginal sea of the Atlantic Ocean, mostly surrounded by the North American continent. It is bounded on the northeast, north, and northwest by the Gulf Coast of the United States; on the southw ...

where its range extends as far north as

Louisiana Louisiana ( ; ; ) is a state in the Deep South and South Central regions of the United States. It borders Texas to the west, Arkansas to the north, and Mississippi to the east. Of the 50 U.S. states, it ranks 31st in area and 25 ...

and the Flower Garden Banks National Marine Sanctuary, the

Florida Keys The Florida Keys are a coral island, coral cay archipelago off the southern coast of Florida, forming the southernmost part of the continental United States. They begin at the southeastern coast of the Florida peninsula, about south of Miami a ...

, the Bahamas and northwestern Cuba south to

Quintana Roo Quintana Roo, officially the Free and Sovereign State of Quintana Roo, is one of the 31 states which, along with Mexico City, constitute the 32 administrative divisions of Mexico, federal entities of Mexico. It is divided into municipalities of ...

ion Mexico and the Bay Islands of Honduras. They are also found off

Santa Marta Santa Marta (), officially the Distrito Turístico, Cultural e Histórico de Santa Marta (), is a port List of cities in Colombia, city on the coast of the Caribbean Sea in northern Colombia. It is the capital of Magdalena Department and the fou ...

in Colombia, east along the South American coast to Venezuela and south to

Vitória, Espírito Santo Vitória (; ) is the capital of the state of Espírito Santo, Brazil. It is located on a small island within a bay where a number of rivers meet the sea. It was founded in 1551. The city proper is and has a population of 322,869 (2022), whilst ...

and Trindade island.

Vagrants Vagrancy is the condition of wandering homelessness without regular employment or income. Vagrants usually live in poverty and support themselves by travelling while engaging in begging, scavenging, or petty theft. In Western countries, ...

have been recorded as far north as

North Carolina North Carolina ( ) is a U.S. state, state in the Southeastern United States, Southeastern region of the United States. It is bordered by Virginia to the north, the Atlantic Ocean to the east, South Carolina to the south, Georgia (U.S. stat ...

Habitat and biology

The Spanish flag is a fish of deeper waters and is found at depths of . It is a rather solitary species which occurs over hard bottoms and reefs, especially on the edge of the continental shelfwhere there are steep slopes and walls. It is largely

piscivorous A piscivore () is a carnivorous animal that primarily eats fish. Fish were the diet of early tetrapod evolution (via water-bound amphibians during the Devonian period); insectivory came next; then in time, the more terrestrially adapted rept ...

, feeding on other fishes, but will also prey on

benthic The benthic zone is the ecological region at the lowest level of a body of water such as an ocean, lake, or stream, including the sediment surface and some sub-surface layers. The name comes from the Ancient Greek word (), meaning "the depths". ...

crustacean Crustaceans (from Latin meaning: "those with shells" or "crusted ones") are invertebrate animals that constitute one group of arthropods that are traditionally a part of the subphylum Crustacea (), a large, diverse group of mainly aquatic arthrop ...

s such as crabs or shrimps.

Taxonomy

The Spanish flag was first formally described as ''Plectropoma hispanum'' in 1828 by the French

naturalist Natural history is a domain of inquiry involving organisms, including animals, fungi, and plants, in their natural environment, leaning more towards observational than experimental methods of study. A person who studies natural history is cal ...

Georges Cuvier Jean Léopold Nicolas Frédéric, baron Cuvier (23 August 1769 – 13 May 1832), known as Georges Cuvier (; ), was a French natural history, naturalist and zoology, zoologist, sometimes referred to as the "founding father of paleontology". Cuv ...

(1769-1832) with the type locality given as

Martinique Martinique ( ; or ; Kalinago language, Kalinago: or ) is an island in the Lesser Antilles of the West Indies, in the eastern Caribbean Sea. It was previously known as Iguanacaera which translates to iguana island in Carib language, Kariʼn ...

. In 1862 the American

ichthyologist Ichthyology is the branch of zoology devoted to the study of fish, including bony fish (Osteichthyes), cartilaginous fish (Chondrichthyes), and jawless fish (Agnatha). According to FishBase, 35,800 species of fish had been described as of March 2 ...

Theodore Nicholas Gill Theodore Nicholas Gill (March 21, 1837 – September 25, 1914) was an American ichthyologist, mammalogist, malacologist, and librarian. Career Born and educated in New York City under private tutors, Gill early showed interest in natural histor ...

(1837-1914) placed it in the

monotypic In biology, a monotypic taxon is a taxonomic group (taxon) that contains only one immediately subordinate taxon. A monotypic species is one that does not include subspecies or smaller, infraspecific taxa. In the case of genera, the term "unisp ...

genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...

''Gonioplectrus''. The relationships of the genus ''Gonioplectrus'' to the other groupers are regarded as unclear but it does sit within the subfamily Epinephelinae.

Utilisation

The Spanish flag is a smallish rare species of deeper waters and as such is not targeted by commercial fisheries. However, as a relatively small, colourful species it is found in the aquarium trade.

References

{{Taxonbar, From=Q2075469 Epinephelini Fish described in 1828 Taxa named by Georges Cuvier Fish of the Western Atlantic