Goniasteridae (the biscuit stars) constitute the largest
family
Family (from ) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). It forms the basis for social order. Ideally, families offer predictabili ...
of
sea star
Starfish or sea stars are Star polygon, star-shaped echinoderms belonging to the class (biology), class Asteroidea (). Common usage frequently finds these names being also applied to brittle star, ophiuroids, which are correctly referred to ...
s, included in the order
Valvatida
The Valvatida are an order of starfish in the class Asteroidea, which contains 695 species in 172 genera in 17 families.
Description
The order encompasses both tiny species, which are only a few millimetres in diameter, like those in the gen ...
. They are mostly deep-dwelling species, but the family also include several colorful shallow tropical species.
Description
Goniasteridae are usually middle-sized sea stars with a characteristic double range of marginal plates bordering the disk and arms. Most of them have five arms, often short and triangular, around a broad central disc; many species are pentagonal or subpentagonal, covered densely with granular, seed-like protuberances, hence the name of the family "seed-star" (''gonium+aster''). The aboral face is often covered with tiny spines looking like
paxillae.
Pedicellaria
A pedicellaria (: pedicellariae) is a small wrench- or claw-shaped appendage with movable jaws, called valves, commonly found on echinoderms (phylum Echinodermata), particularly in sea stars (class Asteroidea) and sea urchins (class Echinoidea).
...
e are often valvate, and the gonads are located at the interradius.
Main identification keys for this group include the presence of paxillae, granules, teeth, spines, or the shape and dimensions of marginal plate.
Location and habitat
They occur predominantly on deep-water
continental shelf
A continental shelf is a portion of a continent that is submerged under an area of relatively shallow water, known as a shelf sea. Much of these shelves were exposed by drops in sea level during glacial periods. The shelf surrounding an islan ...
habitats (but a part of them inhabit shallow waters) in all the world's oceans, being the most diverse in the
Indo-Pacific
The Indo-Pacific is a vast biogeographic region of Earth. In a narrow sense, sometimes known as the Indo-West Pacific or Indo-Pacific Asia, it comprises the tropical waters of the Indian Ocean, the western and central Pacific Ocean, and the ...
region.
List of genera
About 260 extant species within 70 genera are currently known, which make this family the most diverse of all the sea stars,
even if half of the genera are monospecific. Species belonging to the ''
Ferdininae'' subfamily have been imported from
Ophidiasteridae thanks to a large revision of these two families in 2017
According to
World Register of Marine Species
The World Register of Marine Species (WoRMS) is a taxonomic database that aims to provide an authoritative and comprehensive catalogue and list of names of marine organisms.
Content
The content of the registry is edited and maintained by scien ...
, this family includes the following genera:
Image:Akelbaster caledoniae.JPG, '' Akelbaster novaecaledoniae''
Image:Anthenoides peircei - NOAA Photo Library.jpg, '' Anthenoides piercei''
Image:Apollonaster kelleyi.jpg, '' Apollonaster kelleyi''
Image:Astroceramus eldredgei.jpg, '' Astroceramus eldredgei''
Image:Calliaster baccatus P3164673.JPG, '' Calliaster baccatus''
Image:Ceramaster granularis-25mm.jpg, '' Ceramaster granularis''
Image:Chitonaster johannae.jpg, '' Chitonaster johannae''
Image:Circeaster sandrae.JPG, '' Circeaster sandrae''
Image:Cladaster analogus.JPG, '' Cladaster analogus''
Image:Eknomiaster beccae.JPG, '' Eknomiaster beccae''
Image:Eratosaster jenae.JPG, '' Eratosaster jennae''
Image:Evoplosoma.jpg, '' Evoplosoma sp.''
Image:Eosaster nadiae.jpg, '' Eosaster nadiae''
Image:Ferdina flavescens.jpg, '' Ferdina flavescens''
Image:Kanakaster solidus.jpg, '' Kanakaster solidus''
Image:Neoferdina cumingi.jpg, '' Neoferdina cumingi''
Image:Paraferdina plakos.jpg, '' Paraferdina plakos''
Image:Fromia monilis (Seastar).jpg, '' Fromia monilis''
Image:Gilbertaster caribaea.jpg, '' Gilbertaster caribaea''
Image:Glyphodiscus magnificus.jpg, '' Glyphodiscus magnificus''
Image:Goniaster tessellatus São Tomé.jpg, '' Goniaster tessellatus''
Image:Olympic Coast National Marine Sanctuary 2010 Hippasteria spinosa.jpg, '' Hippasteria spinosa''
Image:Iconaster longimanus.jpg, '' Iconaster longimanus''
Image:Mariaster giganteus 01.jpg, `''Mariaster giganteus''
Image:Mediaster aequalis 25March2010 14.jpg, '' Mediaster aequalis''
Image:Milteliphaster woodmasoni.jpg, '' Milteliphaster woodmasoni''
Image:Nectria macrobrachia Large plated seastar P1021082.JPG, '' Nectria macrobrachia''
Image:Nymphaster arenatus.png, '' Nymphaster arenatus''
Image:Peltaster placenta.jpg, '' Peltaster placenta''
Image:Pawsonaster parvus.jpg, '' Pawsonaster parvus''
Image:Pentagonaster duebeni 1.jpg, '' Pentagonaster duebeni''
Image:Pillsburiaster calvus.JPG, '' Pillsburiaster calvus''
Image:Plinthaster dentatus.jpg, '' Plinthaster dentatus''
Image:Pseudoceramaster regularis.jpg, '' Pseudoceramaster regularis''
Image:Rosaster.jpg, '' Rosaster sp.''
Image:Ryukuaster onnae.JPG, '' Ryukuaster onnae''
Image:Sphaeriodiscus mirabilis.JPG, '' Sphaeriodiscus mirabilis''
Image:Galloping sea star (Stellaster equestris).jpg, '' Stellaster childreni''
Image:Stellasteropsis fouadi (MNHN-IE-2014-49) 01.jpg, '' Stellasteropsis fouadi''
Image:Sthenaster emmae.JPG, '' Sthenaster emmae''
Image:Tosia australis P1222858.JPG, '' Tosia australis''
Extinct genera

Lists of genera containing extinct species according to fossilworks.
References
{{Authority control
Echinoderm families
Extant Jurassic first appearances