Glowworm or glow-worm is the common name for various groups of insect
larvae and adult
larviform females that glow through
bioluminescence. They include the European
common glow-worm and other members of the
Lampyridae, but bioluminescence also occurs in the families
Elateridae,
Phengodidae and
Rhagophthalmidae among
beetles; as well as members of the genera ''
Arachnocampa
''Arachnocampa'' is a genus of nine fungus gnat species which have a bioluminescent larval stage, akin to the larval stage of glowworm beetles. The species of ''Arachnocampa'' are endemic to Australia and New Zealand, dwelling in caves and grotto ...
'', ''
Keroplatus'' and ''
Orfelia'' among
keroplatid fungus gnats.
Beetles
Four families of beetles are
bioluminescent. The wingless
larviform females and larvae of these bioluminescent species are usually known as "glowworms". Winged males may or may not also exhibit bioluminescence. Their light may be emitted as flashes or as a constant glow, and usually range in colour from green, yellow, to orange.
The families are closely related, and are all members of the beetle superfamily,
Elateroidea. Phylogenetic analyses have indicated that bioluminescence may have a single evolutionary origin among the families Lampyridae, Phengodidae, and Rhagophthalmidae; but is likely to have arisen independently among Elateridae.
* Family
Elateridae – The click beetles. Of the estimated 10,000 species classified under this family, around 200 species from tropical regions of the Americas and some
Melanesian islands are bioluminescent. All of them are members of the subfamily Pyrophorinae, except for one species, ''
Campyloxenus pyrothorax'', which belongs to subfamily Campyloxeninae, and ''
Balgus schnusei'', in
Thylacosterninae.
* Family
Lampyridae – True fireflies. Contains around 2,000 species found throughout the world. Some "glow worms" are in this family.
* Family
Phengodidae – Usually known as glowworm beetles. Contains around 230 species
endemic to the
New World. This family also includes
railroad worms, which are unique among all terrestrial bioluminescent organisms in producing red light.
* Family
Rhagophthalmidae – Contains around 30 species found in
Asia. The validity of this family has not been fully resolved. Rhagophthalmidae was formerly considered to be a subfamily under Phengodidae before being treated as a distinct family. Some authors now believe that it should be classified under Lampyridae.
Fungus gnats
Three genera of
fungus gnats are bioluminescent, and known as "glowworms" in their larval stage. They produce a blue-green light.
The larvae spin sticky webs to catch food. They are found in caves, overhangs, rock cavities, and other sheltered, wet areas. They are usually classified under the family
Keroplatidae, but this is not universally accepted and some authors place them under
Mycetophilidae instead. Despite the similarities in function and appearance, the bioluminescent systems of the three genera are not
homologous and are believed to have evolved separately.
* Genus ''
Arachnocampa
''Arachnocampa'' is a genus of nine fungus gnat species which have a bioluminescent larval stage, akin to the larval stage of glowworm beetles. The species of ''Arachnocampa'' are endemic to Australia and New Zealand, dwelling in caves and grotto ...
'' – around five species found only in
New Zealand
New Zealand () is an island country in the southwestern Pacific Ocean. It consists of two main landmasses—the North Island () and the South Island ()—and List of islands of New Zealand, over 600 smaller islands. It is the List of isla ...
and
Australia
Australia, officially the Commonwealth of Australia, is a country comprising mainland Australia, the mainland of the Australia (continent), Australian continent, the island of Tasmania and list of islands of Australia, numerous smaller isl ...
. The best-known member of the genus is the New Zealand glowworm, ''
Arachnocampa luminosa
''Arachnocampa luminosa'' (Skuse, 1891), commonly known as New Zealand glowworm or simply glowworm, is a species of fungus gnat solely endemic to New Zealand. The larval stage and the imago produce a blue-green bioluminescence. The species is kno ...
.'' The larvae are predatory and use their lights to lure prey into their webs.
* Genus ''
Orfelia'' – sometimes known as "dismalites". Contains a single species, ''
Orfelia fultoni'', found only in
North America
North America is a continent in the Northern Hemisphere, Northern and Western Hemisphere, Western hemispheres. North America is bordered to the north by the Arctic Ocean, to the east by the Atlantic Ocean, to the southeast by South Ameri ...
. Like ''Arachnocampa'' spp., their larvae may use their lights to attract prey like springtails and other small insects, but their main food is fungal spores.
* Genus ''
Keroplatus'' – found in
Eurasia. Unlike ''Arachnocampa'' and ''Orfelia'', the larvae of ''Keroplatus'' feed only on fungal spores.
Their bioluminescence is believed to have no function and is
vestigial.
See also
*
Larviform
References
External links
{{wikiquote, Glowworms
Springbrook Glow Worms Research Centre.All about glow worms
rachnocampa flavain the Gold Coast hinterland, Queensland Australia.
All About Glow Worm in Tamborine, Gold Coast Descriptive information on Glow Worm Life Cycle and Where are found in Australia.
The UK Glow Worm Surveyhas county-by-county lists of glowworm sites in the UK.
Fly biology
Beetles by adaptation
Bioluminescent insects
Insect common names