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Glowworm or glow-worm is the common name for various groups of insect larvae and adult larviform females that glow through bioluminescence. They include the European common glow-worm and other members of the Lampyridae, but bioluminescence also occurs in the families Elateridae, Phengodidae and Rhagophthalmidae among beetles; as well as members of the genera ''
Arachnocampa ''Arachnocampa'' is a genus of nine fungus gnat species which have a bioluminescent larval stage, akin to the larval stage of glowworm beetles. The species of ''Arachnocampa'' are endemic to Australia and New Zealand, dwelling in caves and grotto ...
'', '' Keroplatus'' and '' Orfelia'' among keroplatid fungus gnats.


Beetles

Four families of beetles are bioluminescent. The wingless larviform females and larvae of these bioluminescent species are usually known as "glowworms". Winged males may or may not also exhibit bioluminescence. Their light may be emitted as flashes or as a constant glow, and usually range in colour from green, yellow, to orange. The families are closely related, and are all members of the beetle superfamily, Elateroidea. Phylogenetic analyses have indicated that bioluminescence may have a single evolutionary origin among the families Lampyridae, Phengodidae, and Rhagophthalmidae; but is likely to have arisen independently among Elateridae. * Family Elateridae – The click beetles. Of the estimated 10,000 species classified under this family, around 200 species from tropical regions of the Americas and some Melanesian islands are bioluminescent. All of them are members of the subfamily Pyrophorinae, except for one species, '' Campyloxenus pyrothorax'', which belongs to subfamily Campyloxeninae, and '' Balgus schnusei'', in Thylacosterninae. * Family Lampyridae – True fireflies. Contains around 2,000 species found throughout the world. Some "glow worms" are in this family. * Family Phengodidae – Usually known as glowworm beetles. Contains around 230 species endemic to the New World. This family also includes railroad worms, which are unique among all terrestrial bioluminescent organisms in producing red light. * Family Rhagophthalmidae – Contains around 30 species found in Asia. The validity of this family has not been fully resolved. Rhagophthalmidae was formerly considered to be a subfamily under Phengodidae before being treated as a distinct family. Some authors now believe that it should be classified under Lampyridae.


Fungus gnats

Three genera of fungus gnats are bioluminescent, and known as "glowworms" in their larval stage. They produce a blue-green light. The larvae spin sticky webs to catch food. They are found in caves, overhangs, rock cavities, and other sheltered, wet areas. They are usually classified under the family Keroplatidae, but this is not universally accepted and some authors place them under Mycetophilidae instead. Despite the similarities in function and appearance, the bioluminescent systems of the three genera are not homologous and are believed to have evolved separately. * Genus ''
Arachnocampa ''Arachnocampa'' is a genus of nine fungus gnat species which have a bioluminescent larval stage, akin to the larval stage of glowworm beetles. The species of ''Arachnocampa'' are endemic to Australia and New Zealand, dwelling in caves and grotto ...
'' – around five species found only in
New Zealand New Zealand () is an island country in the southwestern Pacific Ocean. It consists of two main landmasses—the North Island () and the South Island ()—and List of islands of New Zealand, over 600 smaller islands. It is the List of isla ...
and
Australia Australia, officially the Commonwealth of Australia, is a country comprising mainland Australia, the mainland of the Australia (continent), Australian continent, the island of Tasmania and list of islands of Australia, numerous smaller isl ...
. The best-known member of the genus is the New Zealand glowworm, ''
Arachnocampa luminosa ''Arachnocampa luminosa'' (Skuse, 1891), commonly known as New Zealand glowworm or simply glowworm, is a species of fungus gnat solely endemic to New Zealand. The larval stage and the imago produce a blue-green bioluminescence. The species is kno ...
.'' The larvae are predatory and use their lights to lure prey into their webs. * Genus '' Orfelia'' – sometimes known as "dismalites". Contains a single species, '' Orfelia fultoni'', found only in
North America North America is a continent in the Northern Hemisphere, Northern and Western Hemisphere, Western hemispheres. North America is bordered to the north by the Arctic Ocean, to the east by the Atlantic Ocean, to the southeast by South Ameri ...
. Like ''Arachnocampa'' spp., their larvae may use their lights to attract prey like springtails and other small insects, but their main food is fungal spores. * Genus '' Keroplatus'' – found in Eurasia. Unlike ''Arachnocampa'' and ''Orfelia'', the larvae of ''Keroplatus'' feed only on fungal spores. Their bioluminescence is believed to have no function and is vestigial.


See also

* Larviform


References


External links

{{wikiquote, Glowworms
Springbrook Glow Worms Research Centre.
All about glow worms rachnocampa flavain the Gold Coast hinterland, Queensland Australia.
All About Glow Worm in Tamborine, Gold Coast
Descriptive information on Glow Worm Life Cycle and Where are found in Australia.
The UK Glow Worm Survey
has county-by-county lists of glowworm sites in the UK. Fly biology Beetles by adaptation Bioluminescent insects Insect common names