Gigantornis Eaglesomei
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''Gigantornis eaglesomei'' is a very large prehistoric bird described from a fragmentary specimen from the
Eocene The Eocene ( ) is a geological epoch (geology), epoch that lasted from about 56 to 33.9 million years ago (Ma). It is the second epoch of the Paleogene Period (geology), Period in the modern Cenozoic Era (geology), Era. The name ''Eocene'' comes ...
of
Nigeria Nigeria, officially the Federal Republic of Nigeria, is a country in West Africa. It is situated between the Sahel to the north and the Gulf of Guinea in the Atlantic Ocean to the south. It covers an area of . With Demographics of Nigeria, ...
. It was originally described as a representative of the
albatross Albatrosses, of the biological family Diomedeidae, are large seabirds related to the procellariids, storm petrels, and diving petrels in the order Procellariiformes (the tubenoses). They range widely in the Southern Ocean and the North Paci ...
family, Diomedeidae, but was later referred to the pseudotooth or bony-toothed bird family, Pelagornithidae. ''Gigantornis'' is one of the largest pseudotooth birds, with an
inferred Inferences are steps in logical reasoning, moving from premises to logical consequences; etymologically, the word ''infer'' means to "carry forward". Inference is theoretically traditionally divided into deduction and induction, a distinction ...
wingspan of about 6 m (20 ft) it is among the largest birds ever. Its identified remains consist of a broken
sternum The sternum (: sternums or sterna) or breastbone is a long flat bone located in the central part of the chest. It connects to the ribs via cartilage and forms the front of the rib cage, thus helping to protect the heart, lungs, and major bl ...
found in
Middle Eocene The Eocene ( ) is a geological epoch that lasted from about 56 to 33.9 million years ago (Ma). It is the second epoch of the Paleogene Period in the modern Cenozoic Era. The name ''Eocene'' comes from the Ancient Greek (''Ēṓs'', ' Dawn') a ...
Ameki Formation The Ameki Formation is a Lutetian to Bartonian geological formation located in Nigeria. It belongs to the Bende-Ameki Group, which also consists of two formations namely Nanka Formation, Nsugbe Formation. Fossil content Among others, the foll ...
deposits at Ameki (
Nigeria Nigeria, officially the Federal Republic of Nigeria, is a country in West Africa. It is situated between the Sahel to the north and the Gulf of Guinea in the Atlantic Ocean to the south. It covers an area of . With Demographics of Nigeria, ...
). The
fossil A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserve ...
bird was considered to belong the
albatross Albatrosses, of the biological family Diomedeidae, are large seabirds related to the procellariids, storm petrels, and diving petrels in the order Procellariiformes (the tubenoses). They range widely in the Southern Ocean and the North Paci ...
family Family (from ) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). It forms the basis for social order. Ideally, families offer predictabili ...
(Diomedeidae), as no sterna of pseudotooth birds were known until its discovery, and it remained the only such specimen for decades. Only in the 1970s its true affinities were realized, after it had become clear that although it must have been from a dynamic soarer with wings like an albatross, it resembled pelicans (
order Order, ORDER or Orders may refer to: * A socio-political or established or existing order, e.g. World order, Ancien Regime, Pax Britannica * Categorization, the process in which ideas and objects are recognized, differentiated, and understood ...
Ciconiiformes) rather than
tubenoses Procellariiformes is an order of seabirds that comprises four families: the albatrosses, the petrels and shearwaters, and two families of storm petrels. Formerly called Tubinares and still called tubenoses in English, procellariiforms are of ...
(order Procellariiformes, to which albatrosses belong) in its details. It is not known whether this bird belongs to a distinct genus; it might even be the very same species as the similar-sized ''
Dasornis emuinus ''Dasornis'' is a genus of prehistoric pseudotooth birds. These were probably close relatives of either pelicans and storks or waterfowl; they are placed in the order Odontopterygiformes to account for this uncertainty. Almost all known mater ...
'', whose fossils are not uncommon in the
Ypresian In the geologic timescale the Ypresian is the oldest age (geology), age or lowest stage (stratigraphy), stratigraphic stage of the Eocene. It spans the time between , is preceded by the Thanetian Age (part of the Paleocene) and is followed by th ...
(Early Eocene)
London Clay The London Clay Formation is a Sediment#Shores and shallow seas, marine formation (geology), geological formation of Ypresian (early Eocene Epoch, c. 54-50 million years ago) age which outcrop, crops out in the southeast of England. The London C ...
of the
Isle of Sheppey The Isle of Sheppey is an island off the northern coast of Kent, England, neighbouring the Thames Estuary, centred from central London. It has an area of . The island forms part of the districts of England, local government district of Borough ...
(
England England is a Countries of the United Kingdom, country that is part of the United Kingdom. It is located on the island of Great Britain, of which it covers about 62%, and List of islands of England, more than 100 smaller adjacent islands. It ...
). Remains of a large pseudotooth bird were also found in the Middle Eocene of Kpogamé-Hahotoé (
Togo Togo, officially the Togolese Republic, is a country in West Africa. It is bordered by Ghana to Ghana–Togo border, the west, Benin to Benin–Togo border, the east and Burkina Faso to Burkina Faso–Togo border, the north. It is one of the le ...
) and have been provisionally termed " Aequornis traversei"; their analysis is likely to provide at least some insight on the
taxonomic 280px, Generalized scheme of taxonomy Taxonomy is a practice and science concerned with classification or categorization. Typically, there are two parts to it: the development of an underlying scheme of classes (a taxonomy) and the allocation ...
status of ''G. eaglesomei''. The ''Gigantornis'' sternum is of the typical short and deep-crested shape found in dynamic soarers. Compared to LHNB (CCCP)-1, a
Middle Middle or The Middle may refer to: * Centre (geometry), the point equally distant from the outer limits. Places * Middle (sheading), a subdivision of the Isle of Man * Middle Bay (disambiguation) * Middle Brook (disambiguation) * Middle Creek ...
to
Late Miocene The Late Miocene (also known as Upper Miocene) is a sub-epoch of the Miocene epoch (geology), Epoch made up of two faunal stage, stages. The Tortonian and Messinian stages comprise the Late Miocene sub-epoch, which lasted from 11.63 Ma (million ye ...
pseudotooth bird sternum found in
Portugal Portugal, officially the Portuguese Republic, is a country on the Iberian Peninsula in Southwestern Europe. Featuring Cabo da Roca, the westernmost point in continental Europe, Portugal borders Spain to its north and east, with which it share ...
and tentatively assigned to ''
Pelagornis ''Pelagornis'' is an extinct genus of prehistoric pseudotooth birds, a group of extinct seabirds. Species span from the Oligocene to the Early Pleistocene. Members of ''Pelagornis'' represent among the largest pseudotooth birds, with one species ...
'', its
articular facet A joint or articulation (or articular surface) is the connection made between bones, ossicles, or other hard structures in the body which link an animal's skeletal system into a functional whole.Saladin, Ken. Anatomy & Physiology. 7th ed. McGraw- ...
for the
furcula The (Latin for "little fork"; : furculae) or wishbone is a forked bone found in most birds and some species of non-avian dinosaurs, and is either an interclavicle or formed by the fusion of the two clavicles. In birds, its primary function is ...
consists of a flat section at the very tip of the
sternal keel The sternum (: sternums or sterna) or breastbone is a long flat bone located in the central part of the chest. It connects to the ribs via cartilage and forms the front of the rib cage, thus helping to protect the heart, lungs, and major blood ve ...
and a similar one set immediately above it at an outward angle, and the
spina externa Spina was an Etruscan civilization, Etruscan port city, established by the end of the 6th century BCE, on the Adriatic Sea, Adriatic at the ancient mouth of the Po River, Po. Discovery The site of Spina was lost until modern times, when drainag ...
is shaped like an Old French shield in cross-section. The slightly smaller LHNB (CCCP)-1 has a less sharply protruding sternal keel, the articular facet for the furcula consists of a large knob at the forward margin, and the spina externa is narrow in cross-section.Mayr ''et al.'' (2008)


Footnotes


References

* * Bourdon, Estelle (2005): Osteological evidence for sister group relationship between pseudo-toothed birds (Aves: Odontopterygiformes) and waterfowls (Anseriformes). ''
Naturwissenschaften ''The Science of Nature'', formerly ''Naturwissenschaften'', is a monthly peer-reviewed scientific journal published by Springer Science+Business Media covering all aspects of the natural sciences relating to questions of biological significance. I ...
'' 92(12): 586–591. (HTML abstract
Electronic supplement
(requires subscription) * Bourdon, Estelle (2006): L'avifaune du Paléogène des phosphates du Maroc et du Togo: diversité, systématique et apports à la connaissance de la diversification des oiseaux modernes (Neornithes) Paleogene avifauna of phosphates of Morocco and Togo: diversity, systematics and contributions to the knowledge of the diversification of the Neornithes" Doctoral thesis,
Muséum national d'histoire naturelle The French National Museum of Natural History ( ; abbr. MNHN) is the national natural history museum of France and a of higher education part of Sorbonne University. The main museum, with four galleries, is located in Paris, France, within the Ja ...
n French N, or n, is the fourteenth letter of the Latin alphabet, used in the modern English alphabet, the alphabets of other western European languages, and others worldwide. Its name in English is ''en'' (pronounced ), plural ''ens''. History ...
br>HTML abstract
* Brodkorb, Pierce (1963): Catalogue of fossil birds. Part 1 (Archaeopterygiformes through Ardeiformes). ''Bulletin of the Florida State Museum, Biological Sciences'' 7(4): 179–293
PDF or JPEG fulltext
*
International Commission on Zoological Nomenclature The International Commission on Zoological Nomenclature (ICZN) is an organization dedicated to "achieving stability and sense in the scientific naming of animals". Founded in 1895, it currently comprises 26 commissioners from 20 countries. Orga ...
(ICZN) (1999): ''International Code of Zoological Nomenclature'' (4th ed.). International Trust for Zoological Nomenclature, London. HTML fulltext
* Mayr, Gerald (2009): ''Paleogene Fossil Birds''. Springer-Verlag, Heidelberg & New York. * Mayr, Gerald; Hazevoet, Cornelis J.; Dantas, Pedro; & Cachāo, Mário (2008): A sternum of a very large bony-toothed bird (Pelagornithidae) from the Miocene of Portugal. ''
J. Vertebr. Paleontol. The ''Journal of Vertebrate Paleontology'' is a bimonthly peer-reviewed scientific journal that was established in 1980 by Jiri Zidek (University of Oklahoma). It covers all aspects of vertebrate paleontology, including vertebrate origins, evolu ...
'' 28(3): 762–769. DOI:10.1671/0272-4634(2008)28 62:ASOAVL.0.CO;2HTML abstract
* Olson, Storrs L. (1985): The Fossil Record of Birds. ''In:'' Farner, D.S.; King, J.R. & Parkes, Kenneth C. (eds.): ''Avian Biology'' 8: 79-252
PDF fulltext
{{Taxonbar, from=Q5560236 Pelagornithidae Eocene birds Eocene animals of Africa Prehistoric bird genera