''Gegepterus'' is a

genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial n ...

ctenochasmatid Ctenochasmatidae is a group of pterosaurs within the suborder Pterodactyloidea. They are characterized by their distinctive teeth, which are thought to have been used for filter-feeding. Ctenochasmatids lived from the Late Jurassic to the Early ...

pterosaur Pterosaurs (; from Greek ''pteron'' and ''sauros'', meaning "wing lizard") is an extinct clade of flying reptiles in the Order (biology), order, Pterosauria. They existed during most of the Mesozoic: from the Late Triassic to the end of the Cre ...

from the

Early Cretaceous The Early Cretaceous (geochronological name) or the Lower Cretaceous ( chronostratigraphic name), is the earlier or lower of the two major divisions of the Cretaceous. It is usually considered to stretch from 145 Ma to 100.5 Ma. Geology Pr ...

period of what is now the

Yixian Formation The Yixian Formation (; formerly transcribed as Yihsien Formation) is a geological formation in Jinzhou, Liaoning, People's Republic of China, that spans the late Barremian and early Aptian stages of the Early Cretaceous. It is known for its ...

of Liaoning, China. Only one species is known, ''G. changi''.

History and etymology

The genus was named in 2007 by

Wang Xiaolin Wang may refer to: Names * Wang (surname) (王), a common Chinese surname * Wāng (汪), a less common Chinese surname * Titles in Chinese nobility * A title in Korean nobility * A title in Mongolian nobility Places * Wang River in Thailand ...

Alexander Kellner Alexander Wilhelm Armin Kellner (born September 26, 1961) is a Brazilian geologist and paleontologist who is a leading expert in the field of studying pterosaurs. His research has focused mainly on fossil reptiles from the Cretaceous Period, i ...

Zhou Zhonge Zhou Zhongge (; born 15 February 1967) is a Chinese track and field coach and former high jumper. His personal best was , set in 1990. He was the gold medallist at the 1990 Asian Games and the 1998 Asian Games silver medallist. He was the winner a ...

and

Diogenes de Almeida Campos Diogenes ( ; grc, Διογένης, Diogénēs ), also known as Diogenes the Cynic (, ) or Diogenes of Sinope, was a Greek philosopher and one of the founders of Cynicism (philosophy). He was born in Sinope, an Ionian colony on the Black Sea ...

. The

type species In zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the species that contains the biological type specimen( ...

is ''Gegepterus changi''. The generic name is derived from

Manchu The Manchus (; ) are a Tungusic East Asian ethnic group native to Manchuria in Northeast Asia. They are an officially recognized ethnic minority in China and the people from whom Manchuria derives its name. The Later Jin (1616–1636) and Q ...

''ge ge'', the title of a princess, in reference to the dainty gracility of the specimen, and a Latinized Greek ''pteron'', "wing". The specific name honors female paleontologist Chang Meemann, who over the years established a cordial relationship between the Chinese and Brazilian authors. In 2008 Wang emended the epithet to ''changae'', but such changes are no longer allowed by the

ICZN The International Code of Zoological Nomenclature (ICZN) is a widely accepted convention in zoology that rules the formal scientific naming of organisms treated as animals. It is also informally known as the ICZN Code, for its publisher, the Int ...

. It is known from two specimens. The first is

holotype A holotype is a single physical example (or illustration) of an organism, known to have been used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of seve ...

IVPP V 11981, which was in 2001 found in grey

shale Shale is a fine-grained, clastic sedimentary rock formed from mud that is a mix of flakes of clay minerals (hydrous aluminium phyllosilicates, e.g. kaolin, Al2 Si2 O5( OH)4) and tiny fragments (silt-sized particles) of other minerals, especia ...

s from the lower part of the

formation Formation may refer to: Linguistics * Back-formation, the process of creating a new lexeme by removing or affixes * Word formation, the creation of a new word by adding affixes Mathematics and science * Cave formation or speleothem, a secondary ...

(estimated at 125 million years old), near the city of

Beipiao Beipiao () is a city in Chaoyang prefecture, Liaoning province, in Northeast China. It has a population of 202,807. The main industry in the area is coal mining. With vertical shafts of almost 1000m, these are some of the deepest coal mines in ...

. It consists of a crushed and damaged partial skeleton of a

subadult A juvenile is an individual organism that has not yet reached its adult form, sexual maturity or size. Juveniles can look very different from the adult form, particularly in colour, and may not fill the same Ecological niche, niche as the adult ...

including skull, lower jaws, cervical and sacral

vertebra The spinal column, a defining synapomorphy shared by nearly all vertebrates, Hagfish are believed to have secondarily lost their spinal column is a moderately flexible series of vertebrae (singular vertebra), each constituting a characterist ...

e, ribs,

gastralia Gastralia (singular gastralium) are dermal bones found in the ventral body wall of modern crocodilians and tuatara, and many prehistoric tetrapods. They are found between the sternum and pelvis, and do not articulate with the vertebrae. In thes ...

("belly ribs"),

shoulder girdle The shoulder girdle or pectoral girdle is the set of bones in the appendicular skeleton which connects to the arm on each side. In humans it consists of the clavicle and scapula; in those species with three bones in the shoulder, it consists ...

and hindlimb remains, along with dark soft tissue remains near the skull and gastralia and in the orbit; unfortunately, the soft tissue remains show no structure except for some small, unbranched fibers at the back of the head. The jaws are very elongated; the snout is flat and concave on top, with a low and thin crest. The forehead slightly projects to the front. The cervicals are elongated.Wang, X., Kellner, A.W.A., Zhou, Z., and Campos, D.A. (2007). "A new pterosaur (Ctenochasmatidae, Archaeopterodactyloidea) from the Lower Cretaceous Yixian Formation of China." ''Cretaceous Research'', 28(2): 2245-260. In 2011, a second, smaller specimen was described, IVPP V 11972, which increased the known skeletal elements and showed a more extensive covering of hair-like structures.Jiang Shun-Xing & Wang Xiao-Lin, 2011, "Important features of ''Gegepterus changae'' (Pterosauria: Archaeopterodactyloidea, Ctenochasmatidae) from a new specimen", ''Vertebrata PalAsiatica'' 49(2): 172-184 http://www.ivpp.cas.cn/cbw/gjzdwxb/xbwzxz/201105/P020110530510419201105.pdf

Classification

The authors assigned ''Gegepterus'' to the

Ctenochasmatidae Ctenochasmatidae is a group of pterosaurs within the suborder Pterodactyloidea. They are characterized by their distinctive teeth, which are thought to have been used for filter-feeding. Ctenochasmatids lived from the Late Jurassic to the Early C ...

, on the basis of its long

rostrum Rostrum may refer to: * Any kind of a platform for a speaker: **dais **pulpit * Rostrum (anatomy), a beak, or anatomical structure resembling a beak, as in the mouthparts of many sucking insects * Rostrum (ship), a form of bow on naval ships * Ros ...

and numerous needle-like

teeth A tooth ( : teeth) is a hard, calcified structure found in the jaws (or mouths) of many vertebrates and used to break down food. Some animals, particularly carnivores and omnivores, also use teeth to help with capturing or wounding prey, te ...

, about 150 in total. This is the first uncontroversial report of the Ctenochasmatidae from the Yixian Formation, as the fossils of other assumed ctenochasmatids have not preserved the dentition. It was at first suspected to be the juvenile of some known species. Below is

cladogram A cladogram (from Greek ''clados'' "branch" and ''gramma'' "character") is a diagram used in cladistics to show relations among organisms. A cladogram is not, however, an evolutionary tree because it does not show how ancestors are related to ...

following a topology by Andres, Clark and Xu (2014). In the analysis, they recovered ''Gegepterus'' within the family Ctenochasmatidae, more precisely within the tribe Pterodaustrini, sister taxon to '' Beipiaopterus''.

References

{{Portal bar, Paleontology, Cretaceous, China Early Cretaceous pterosaurs of Asia Ctenochasmatoids Fossil taxa described in 2007 Taxa named by Alexander Kellner Yixian fauna

History and etymology

Classification

See also

References