''Fraxinus ornus'', the manna ash or South European flowering ash, is a species of ''

Fraxinus ''Fraxinus'' (), commonly called ash, is a genus of plants in the olive and lilac family, Oleaceae, and comprises 45–65 species of usually medium-to-large trees, most of which are deciduous trees, although some Subtropics, subtropical specie ...

'' native to

Southern Europe Southern Europe is also known as Mediterranean Europe, as its geography is marked by the Mediterranean Sea. Definitions of southern Europe include some or all of these countries and regions: Albania, Andorra, Bosnia and Herzegovina, Bulgaria, C ...

and Southwestern Asia, from

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and

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north to

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and the

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, and east through the

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, and western

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and

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.Rushforth, K. (1999). ''Trees of Britain and Europe''. Collins .Flora Europaea
''Fraxinus ornus''
/ref>Med-Checklist
''Fraxinus ornus''
/ref>

Description

''Fraxinus ornus'' is a medium-sized

deciduous In the fields of horticulture and botany, the term deciduous () means "falling off at maturity" and "tending to fall off", in reference to trees and shrubs that seasonally shed Leaf, leaves, usually in the autumn; to the shedding of petals, aft ...

tree In botany, a tree is a perennial plant with an elongated stem, or trunk, usually supporting branches and leaves. In some usages, the definition of a tree may be narrower, e.g., including only woody plants with secondary growth, only ...

growing to tall with a trunk up to 1 m diameter. The

bark Bark may refer to: Common meanings * Bark (botany), an outer layer of a woody plant such as a tree or stick * Bark (sound), a vocalization of some animals (which is commonly the dog) Arts and entertainment * ''Bark'' (Jefferson Airplane album), ...

is dark grey, remaining smooth even on old trees. The

bud In botany, a bud is an undeveloped or Plant embryogenesis, embryonic Shoot (botany), shoot and normally occurs in the axil of a leaf or at the tip of a Plant stem, stem. Once formed, a bud may remain for some time in a dormancy, dormant conditi ...

s are pale pinkish-brown to grey-brown, with a dense covering of short grey hairs. The

leaves A leaf (: leaves) is a principal appendage of the stem of a vascular plant, usually borne laterally above ground and specialized for photosynthesis. Leaves are collectively called foliage, as in "autumn foliage", while the leaves, stem, ...

are in opposite pairs, pinnate, long, with 5 to 9 leaflets; the leaflets are broad ovoid, long and broad, with a finely serrated and wavy margin, and short but distinct petiolules long; the autumn colour is variable, yellow to purplish. The

flower Flowers, also known as blooms and blossoms, are the reproductive structures of flowering plants ( angiosperms). Typically, they are structured in four circular levels, called whorls, around the end of a stalk. These whorls include: calyx, m ...

s are produced in dense

panicle In botany, a panicle is a much-branched inflorescence. (softcover ). Some authors distinguish it from a compound spike inflorescence, by requiring that the flowers (and fruit) be pedicellate (having a single stem per flower). The branches of a p ...

s long after the new leaves appear in late spring, each flower with four slender creamy white petals long; they are pollinated by

insect Insects (from Latin ') are Hexapoda, hexapod invertebrates of the class (biology), class Insecta. They are the largest group within the arthropod phylum. Insects have a chitinous exoskeleton, a three-part body (Insect morphology#Head, head, ...

s. The

fruit In botany, a fruit is the seed-bearing structure in flowering plants (angiosperms) that is formed from the ovary after flowering. Fruits are the means by which angiosperms disseminate their seeds. Edible fruits in particular have long propaga ...

is a slender

samara Samara, formerly known as Kuybyshev (1935–1991), is the largest city and administrative centre of Samara Oblast in Russia. The city is located at the confluence of the Volga and the Samara (Volga), Samara rivers, with a population of over 1.14 ...

long, the seed broad and the wing broad, green ripening brown.Rushforth, K. (1999). ''Trees of Britain and Europe''. Collins .Mitchell, A. F. (1974). ''A Field Guide to the Trees of Britain and Northern Europe''. Collins Mitchell, A. F. (1982). ''The Trees of Britain and Northern Europe''. Collins

Cultivation and uses

''Fraxinus ornus'' is frequently grown as an

ornamental tree Ornamental plants or ''garden plants'' are plants that are primarily grown for their beauty but also for qualities such as scent or how they shape physical space. Many flowering plants and garden varieties tend to be specially bred cultivars th ...

in Europe north of its native range for its decorative flowers—the species is also sometimes called "flowering ash". Some cultivated specimens are grafted on rootstocks of ''

Fraxinus excelsior ''Fraxinus excelsior'', known as the ash, or European ash or common ash to distinguish it from other types of ash, is a flowering plant species in the olive family Oleaceae. It is native throughout mainland Europe east to the Caucasus and Alb ...

'', with an often very conspicuous change in the bark at the graft line to the fissured bark of the rootstock species. A

sugar Sugar is the generic name for sweet-tasting, soluble carbohydrates, many of which are used in food. Simple sugars, also called monosaccharides, include glucose Glucose is a sugar with the Chemical formula#Molecular formula, molecul ...

y extract from the sap may be obtained from the bark of the tree;Traverso V. The return of Sicily's ancient 'white gold'. ''BBC News'' 30 March 2025
/ref> which was compared to biblical

manna Manna (, ; ), sometimes or archaically spelled Mahna or Mana, is described in the Bible and the Quran as an edible substance that God in Abrahamic religions, God bestowed upon the Israelites while they were wandering the desert during the 40-year ...

in late medieval times (attested circa 1400 AD''

Oxford English Dictionary The ''Oxford English Dictionary'' (''OED'') is the principal historical dictionary of the English language, published by Oxford University Press (OUP), a University of Oxford publishing house. The dictionary, which published its first editio ...

''), giving rise to the name of the tree in English, Spanish (''fresno del maná''), and Italian (''frassino da manna''). Likewise, the sugar

mannose Mannose is a sugar with the formula , which sometimes is abbreviated Man. It is one of the monomers of the aldohexose series of carbohydrates. It is a C-2 epimer of glucose. Mannose is important in human metabolism, especially in the glycosylatio ...

and the sugar alcohol

mannitol Mannitol is a type of sugar alcohol used as a sweetener and medication. It is used as a low calorie sweetener as it is poorly absorbed by the intestines. As a medication, it is used to decrease pressure in the eyes, as in glaucoma, and to l ...

both derive their names from the extract.

References

External links

Production of manna in Sicily

''Fraxinus ornus''
- information, genetic conservation units and related resources.

European Forest Genetic Resources Programme European Forest Genetic Resources Programme (EUFORGEN) is an international network that promotes the conservation and sustainable use of forest genetic resources in Europe as an integral part of sustainable forest management. It was established ...

(EUFORGEN) {{Taxonbar, from=Q161421 ornus Flora of Southeastern Europe Flora of Western Asia Flora of Ukraine Flora of Spain Flora of Sardinia Flora of Portugal Flora of France Flora of Corsica Flora of Switzerland Flora of the Czech Republic Flora of Hungary Flora of Armenia Flora of Azerbaijan Flora of Georgia (country) Plants described in 1753 Taxa named by Carl Linnaeus Garden plants of Asia Garden plants of Europe Ornamental trees Trees of mild maritime climate Trees of Mediterranean climate