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Francis "Frank" Milburn Howlett (5 January 1877 – 20 August 1920) was a British entomologist who served in India in the position of a Second Imperial Entomologist, a position which was later changed to the Imperial Pathological Entomologist in India. He specialized in insects (mainly Diptera - sandflies) and parasitic ticks of medical and veterinary importance. A major discovery by him was the attractant methyl eugenol and its effect on flies of the genus ''
Bactrocera ''Bactrocera'' is a large genus of tephritid fruit flies, with close to 500 species currently described and accepted. Name The genus name is derived from Ancient Greek Ancient Greek (, ; ) includes the forms of the Greek language used ...
''.


Life and work

Howlett was born in
Wymondham Wymondham ( ) is a market town and civil parishes in England, civil parish in the South Norfolk district of Norfolk, England. It lies on the River Tiffey, south-west of Norwich and just off the A11 road (England), A11 road to London. The pari ...
,
Norfolk Norfolk ( ) is a Ceremonial counties of England, ceremonial county in England, located in East Anglia and officially part of the East of England region. It borders Lincolnshire and The Wash to the north-west, the North Sea to the north and eas ...
, the son of Francis John Howlett, a
solicitor A solicitor is a lawyer who traditionally deals with most of the legal matters in some jurisdictions. A person must have legally defined qualifications, which vary from one jurisdiction to another, to be described as a solicitor and enabled to p ...
, and Mary Jane née Milburn. He was educated at Wymondham Grammar School and Bury St Edmunds Grammar School, and then at
Christ's College, Cambridge Christ's College is a Colleges of the University of Cambridge, constituent college of the University of Cambridge, England. The college includes the Master, the Fellows of the College, and about 450 undergraduate and 250 graduate students. The c ...
. He was an assistant master at
Edinburgh Academy The Edinburgh Academy is a Private schools in the United Kingdom, private day school in Edinburgh, Scotland, which was opened in 1824. The original building, on Henderson Row in Stockbridge, Edinburgh, Stockbridge, is now part of the Senior Scho ...
from 1900 to 1903 and at Holt Grammar School before being posted as a professor of natural science (which included the teaching of chemistry) at Muir Central College, Allahabad, from 1905 to 1908, initially in a temporary position (to replace E.G. Hill who was on furlough) which was then extended. He joined the Imperial Agricultural Research Institute at Pusa in December 1907 as Second (i.e. deputy) Imperial Entomologist under
Harold Maxwell-Lefroy Harold Maxwell-Lefroy (20 January 1877 – 14 October 1925) was an England, English entomologist. He served as a Professor of Entomology at Imperial College London before moving to India where he took over the position of entomologist to the gover ...
and from 1912 as Imperial Pathological Entomologist for the Government of India. In 1910 he was in England and Harold Maxwell-Lefroy deputed him to attend the first International Entomological Congress in Brussels, where he presented on the state of economic entomology in India and also on issues in preserving specimens in India. He left India during the
First World War World War I or the First World War (28 July 1914 – 11 November 1918), also known as the Great War, was a World war, global conflict between two coalitions: the Allies of World War I, Allies (or Entente) and the Central Powers. Fighting to ...
and worked with the Royal Army Medical College, while also attending various meetings of learned societies, and returned to India only in 1917. In 1919 he presided over the zoological section of the 6th Indian Science Congress at Bombay, giving a talk on "tactics against insects." One of his most important findings was in noting the attraction of tephritid flies to
methyl eugenol Methyl eugenol (allylveratrol) is a natural chemical compound classified as a phenylpropene, a type of phenylpropanoid. It is the methyl ether of eugenol and is important to insect behavior and pollination. It is found in various essential oils. ...
, a component that he identified from several others present in
citronella oil Citronella oilCitronella oil is an essential oil obtained from the leaves and stems of different species of '' Cymbopogon'' (lemongrass). The oil is used extensively as a source of perfumery chemicals such as citronellal, citronellol, and geran ...
. He also noted how fleas disliked wet grass and noted a decline in plague during the onset of the rains and suggested that this might be a reason for the plague being rare in Bengal. He published a technique to trap
thrips Thrips (Order (biology) , order Thysanoptera) are minute (mostly long or less), slender insects with fringed wings and unique asymmetrical mouthparts. Entomologists have species description , described approximately 7,700 species. They fly on ...
with attractant mixtures of
Benzaldehyde Benzaldehyde (C6H5CHO) is an organic compound consisting of a benzene ring with a formyl substituent. It is among the simplest aromatic aldehydes and one of the most industrially useful. It is a colorless liquid with a characteristic almond-li ...
,
Cinnamaldehyde Cinnamaldehyde is an organic compound with the formula or . Occurring naturally as predominantly the ''trans'' (''E'') isomer, it gives cinnamon its flavor and odor. It is a phenylpropanoid that is naturally synthesized by the shikimate pathway ...
and Anisaldehyde. Howlett also discovered that he could induce ''Stomoxys calcitrans'' to oviposit on cotton impregnated with valerianic acid, which is a component of fermenting vegetable matter. He also noted the life history of mosquito larvae (''Stegomyia'', now ''
Anopheles ''Anopheles'' () is a genus of mosquito first described by the German entomologist Johann Wilhelm Meigen, J. W. Meigen in 1818, and are known as nail mosquitoes and marsh mosquitoes. Many such mosquitoes are Disease vector, vectors of the paras ...
'') that could survive in dry soil. He also studied the biology of sandflies. Howlett was also known for his humorous sketches at Pusa. He later moved to the Agricultural Research Institute at Pune. Howlett was an athlete and an artist who illustrated his own works. His health was poor during his service in India and he died a premature death from complications following a surgical procedure at
Mussoorie Mussoorie () is a hill station and a municipal board, in Dehradun city in the Dehradun district of the Indian state of Uttarakhand. It is about from the state capital of Dehradun and north of the national capital of New Delhi. The hil ...
. At the time of his death he was working on a book titled ''The Control of Harmful Insects''. An obituarist in the ''Agricultural Journal of India'' noted that he was:
... a man of almost childlike simplicity and originality of outlook, and with many interests. ... He had the faults as well as the merits of the artistic temperament. ... He was a born schoolmaster, delighted in teaching, and could make all subjects interesting, and had the gift of implanting in his disciples some of his own enthusiasm. He was a combative apostle of pure research, and his disappearance from the ranks of scientists of this order will be a serious blow to the cause.
Howlett assisted Harold Maxwell-Lefroy in writing and illustrating the book ''Indian Insect Life''. He wrote the sections on the flies and trained staff at Pusa in technical illustration. A species of tick, ''Haemaphysalis howletti'', was described by Warburton in 1913 from a pony in Pakistan and in 1962 was found on rodents and birds in Pune, Maharashtra. Howlett developed techniques for collecting and preserving insects and for marking insects (houseflies) to study dispersal. Brunetti, named a fly after Howlett as ''Howlettia'' (now considered a synonym of '' Platypalpus'' of family Hybotidae).


Publications

Apart from the publications cited, Howlett's works include: * * * * * * * * *


References

{{DEFAULTSORT:Howlett, Frank Milburn British entomologists 1877 births 1920 deaths Alumni of the University of Cambridge