Francis Gotch (13 July 1853 – 15 July 1913) was a British neurophysiologist who was professor of

physiology Physiology (; ) is the science, scientific study of function (biology), functions and mechanism (biology), mechanisms in a life, living system. As a branches of science, subdiscipline of biology, physiology focuses on how organisms, organ syst ...

at University College Liverpool and

Oxford University The University of Oxford is a collegiate research university in Oxford, England. There is evidence of teaching as early as 1096, making it the oldest university in the English-speaking world and the second-oldest continuously operating u ...

. He was educated at Amersham Hall School and then at

London University The University of London (UoL; abbreviated as Lond or more rarely Londin in post-nominals) is a federal public research university located in London, England, United Kingdom. The university was established by royal charter in 1836 as a degr ...

graduating B.A. in 1873 and then B.Sc. After studying medicine he qualified M.R.C.S. in 1881. In 1887 he married Rosamund Brunel Horsley (1864-1949), the younger daughter of the artist

John Callcott Horsley John Callcott Horsley (29 January 1817 – 18 October 1903) was a British academic Painting, painter of genre painting, genre and historical scenes, illustrator, and designer of the first Christmas card. He was a member of the artist's colony ...

. He made several pioneer contributions to British neurophysiology. With his brother-in-law,

Victor Horsley Sir Victor Alexander Haden Horsley (14 April 1857 – 16 July 1916) was a British scientist and professor. He was born in Kensington, London. Educated at Cranbrook School, Kent, he studied medicine at University College London and in Berlin, G ...

(1857–1916), he performed research involving

localization of brain function In neuroscience, functional specialization is a theory which suggests that List of regions in the human brain, different areas in the brain are specialized for different functions.Flourens, M. J. P. (1824) Recherces experimentales sur les propr ...

via

electrical stimulation Functional electrical stimulation (FES) is a technique that uses low-energy electrical pulses to artificially generate body movements in individuals who have been paralyzed due to injury to the central nervous system. More specifically, FES ca ...

of the

cortex Cortex or cortical may refer to: Biology * Cortex (anatomy), the outermost layer of an organ ** Cerebral cortex, the outer layer of the vertebrate cerebrum, part of which is the ''forebrain'' *** Motor cortex, the regions of the cerebral cortex i ...

, and also demonstrated that the mammalian brain was capable of producing

electric current An electric current is a flow of charged particles, such as electrons or ions, moving through an electrical conductor or space. It is defined as the net rate of flow of electric charge through a surface. The moving particles are called charge c ...

. In 1899 he described the "inexcitable" or "refractory phase" that takes place between

nerve impulse An action potential (also known as a nerve impulse or "spike" when in a neuron) is a series of quick changes in voltage across a cell membrane. An action potential occurs when the membrane potential of a specific Cell (biology), cell rapidly ri ...

s. He also performed significant research in the field of

electroretinography Electroretinography measures the electrical responses of various cell types in the retina, including the Photoreceptor cell, photoreceptors (rod cell, rods and cone cell, cones), inner retinal cells (Retinal bipolar cell, bipolar and amacrine ce ...

. In 1891, with Horsley, he delivered the

Croonian Lecture The Croonian Medal and Lecture is a prestigious award, a medal, and lecture given at the invitation of the Royal Society and the Royal College of Physicians. Among the papers of William Croone at his death in 1684, was a plan to endow a singl ...

before the

Royal Society of London The Royal Society, formally The Royal Society of London for Improving Natural Knowledge, is a learned society and the United Kingdom's national academy of sciences. The society fulfils a number of roles: promoting science and its benefits, r ...

, entitled "On The Mammalian Nervous System: Its Functions, And Their Localization Determined By An Electrical Method". In June 1892 he was elected a

Fellow of the Royal Society Fellowship of the Royal Society (FRS, ForMemRS and HonFRS) is an award granted by the Fellows of the Royal Society of London to individuals who have made a "substantial contribution to the improvement of natural science, natural knowledge, incl ...

His body was interred at

Wolvercote Cemetery Wolvercote Cemetery is a cemetery in the parish of Wolvercote and district of Cutteslowe in Oxford, England. Its main entrance is on Banbury Road and it has a side entrance in Five Mile Drive. It has a funeral chapel, public toilets and a small a ...

References

Further reading