Foxtail Pines
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''Pinus balfouriana'', the foxtail pine, is a rare high-elevation
pine A pine is any conifer tree or shrub in the genus ''Pinus'' () of the family Pinaceae. ''Pinus'' is the sole genus in the subfamily Pinoideae. ''World Flora Online'' accepts 134 species-rank taxa (119 species and 15 nothospecies) of pines as cu ...
that is
endemic Endemism is the state of a species being found only in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also foun ...
to
California California () is a U.S. state, state in the Western United States that lies on the West Coast of the United States, Pacific Coast. It borders Oregon to the north, Nevada and Arizona to the east, and shares Mexico–United States border, an ...
,
United States The United States of America (USA), also known as the United States (U.S.) or America, is a country primarily located in North America. It is a federal republic of 50 U.S. state, states and a federal capital district, Washington, D.C. The 48 ...
. It is closely related to the
Great Basin The Great Basin () is the largest area of contiguous endorheic watersheds, those with no outlets to the ocean, in North America. It spans nearly all of Nevada, much of Utah, and portions of California, Idaho, Oregon, Wyoming, and Baja Californi ...
and Rocky Mountain bristlecone pines, in the subsection ''
Balfourianae The term bristlecone pine covers three species of pine tree (family Pinaceae, genus ''Pinus'', subsection ''Balfourianae''). All three species are long-lived and highly resilient to harsh weather and bad soils. One of the three species, ''Pinus ...
''.


Description

''P. balfouriana'' is a
tree In botany, a tree is a perennial plant with an elongated stem, or trunk, usually supporting branches and leaves. In some usages, the definition of a tree may be narrower, e.g., including only woody plants with secondary growth, only ...
to tall, exceptionally , with a
trunk Trunk may refer to: Biology * Trunk (anatomy), synonym for torso * Trunk (botany), a tree's central superstructure, and the stem of woody plants * Trunk of corpus callosum, in neuroanatomy * Elephant trunk, the proboscis of an elephant Comput ...
up to across. Its
leaves A leaf (: leaves) is a principal appendage of the stem of a vascular plant, usually borne laterally above ground and specialized for photosynthesis. Leaves are collectively called foliage, as in "autumn foliage", while the leaves, stem, ...
are needle-like, in bundles of five (or sometimes four, in the southern Sierra) with a semi-persistent basal sheath, and long, deep glossy green on the outer face, and white on the inner faces; they persist for 10–15 years. The
cones In geometry, a cone is a three-dimensional figure that tapers smoothly from a flat base (typically a circle) to a point not contained in the base, called the ''apex'' or '' vertex''. A cone is formed by a set of line segments, half-lines, ...
are long, dark purple ripening red-brown, with soft, flexible scales each with a central prickle.


Distribution

''P. balfouriana'' occurs in the
subalpine Montane ecosystems are found on the slopes of mountains. The alpine climate in these regions strongly affects the ecosystem because temperatures fall as elevation increases, causing the ecosystem to stratify. This stratification is a crucial f ...
forest at an elevation of in the
Klamath Mountains The Klamath Mountains are a rugged and lightly populated mountain range in northwestern California and southwestern Oregon in the western United States. As a mountain system within both the greater Pacific Coast Ranges and the California Coast R ...
, and at in the
Sierra Nevada The Sierra Nevada ( ) is a mountain range in the Western United States, between the Central Valley of California and the Great Basin. The vast majority of the range lies in the state of California, although the Carson Range spur lies primari ...
. In the Sierra Nevada, Foxtail pines are limited to the area around Sequoia and
Kings Canyon National Park Kings Canyon National Park is a national park of the United States in the southern Sierra Nevada, in Fresno and Tulare Counties, California. Originally established in 1890 as General Grant National Park, the park was greatly expanded and ren ...
s. In both areas, it is often a
tree line The tree line is the edge of a habitat at which trees are capable of growing and beyond which they are not. It is found at high elevations and high latitudes. Beyond the tree line, trees cannot tolerate the environmental conditions (usually low ...
species. There are two
disjunct population In biology, a taxon with a disjunct distribution is one that has two or more groups that are related but considerably separated from each other geographically. The causes are varied and might demonstrate either the expansion or contraction of a s ...
s: A small outlying population was reported in southern Oregon, but was proven to have been misidentified.Kauffmann, Michael E. (2012). Conifer Country. Kneeland, CA: Backcountry Press. . .


Age

It is thought that ''P. balfouriana'' can live up to 3000 years in the Sierra Nevada, although the highest currently proven age is 2110 years. In the Klamath Mountains, ages are only known to about 1000 years.


Related species

''P. balfouriana'' is closely related to the
bristlecone pine The term bristlecone pine covers three species of pine tree (family Pinaceae, genus ''Pinus'', subsection ''Balfourianae''). All three species are long-lived and highly resilient to harsh weather and bad soils. One of the three species, ''Pinus ...
s, being classified in the same subsection ''
Balfourianae The term bristlecone pine covers three species of pine tree (family Pinaceae, genus ''Pinus'', subsection ''Balfourianae''). All three species are long-lived and highly resilient to harsh weather and bad soils. One of the three species, ''Pinus ...
''; it has been hybridised with the
Great Basin Bristlecone Pine ''Pinus longaeva'' (commonly referred to as the Great Basin bristlecone pine, intermountain bristlecone pine, or western bristlecone pine) is a long-living species of bristlecone pine tree found in the higher mountains of California, Nevada, and ...
in cultivation, though no hybrids have ever been found in the wild.


References


Further reading

* * Bailey, D.K. 1970. Phytogeography and taxonomy of Pinus subsection Balfourianae. ''Ann. Missouri Bot. Gard.'' 57: 210–249. * Mastrogiuseppe, R.J. & Mastrogiuseppe, J.D. 1980. A study of Pinus balfouriana Grev. & Balf. (Pinaceae). ''Systematic Botany'' 5: 86–104. * Richardson, D.M. (ed.). 1998. ''Ecology and Biogeography of Pinus''. Cambridge University Press, Cambridge. 530 p. . *


External links


Calflora: Pinus balfouriana CalFlora Database: ''Pinus balfouriana'' (foxtail pine)

Gymnosperm Database – ''Pinus balfouriana''

USDA Plants Profile for ''Pinus balfouriana'' (foxtail pine)




* * {{Authority control balfouriana Endemic flora of California Trees of Northern America Taxa named by John Hutton Balfour