Fordilloidea is an
extinct
Extinction is the termination of an organism by the death of its Endling, last member. A taxon may become Functional extinction, functionally extinct before the death of its last member if it loses the capacity to Reproduction, reproduce and ...
superfamily of early
bivalve
Bivalvia () or bivalves, in previous centuries referred to as the Lamellibranchiata and Pelecypoda, is a class (biology), class of aquatic animal, aquatic molluscs (marine and freshwater) that have laterally compressed soft bodies enclosed b ...
s
containing two described families,
Fordillidae and
Camyidae and the only superfamily in the order Fordillida.
The superfamily is known from fossils of
early to middle Cambrian age found in North America, Greenland, Europe, the Middle East, Asia, and Australia.
[The Paleobiology Database Fordillidae entry](_blank)
accessed 4 January 2012[The Paleobiology Database ''Fordilla'' entry](_blank)
accessed 4 January 2012 Fordillidae currently contains two genera, ''
Fordilla'' and ''
Pojetaia'' each with up to three described species while Camyidae only contains a single genus ''
Camya'' with one described species, ''Camya asy''.
Due to the size and age of the fossil specimens, Fordillidae species are included as part of the Turkish
Small shelly fauna.
Description
''Fordilla'' are small bivalves with
valves
A valve is a device or natural object that regulates, directs or controls the flow of a fluid (gases, liquids, fluidized solids, or slurries) by opening, closing, or partially obstructing various passageways. Valves are technically fittings, ...
that are equal in size and suboval in shape. In size, ''Fordilla'' specimens reach a total shell length of up to and a height of .
The shells are compressed laterally and the back edge is slightly broadened. The rear adductor is less developed and smaller than the front adductor, while the small pedal retractor muscle scar is positioned near the front adductor scar.
The valve hinge is usually straight to slightly convexly curved and each valve will have at most one tooth present. The external surface of the shell occasionally shows faint ribbing.
Similar to ''Fordilla'', species of ''Pojetaia'' are small, with valves to less than in length. ''Pojetaia'' species have an overall shape which is suboval, with the subequal valves slightly elongated. The ligament is straight with an umbo which is central to subcentral. In contrast to ''Fordilla'', the rear adductor muscle was larger and more developed than the front adductor, with pallial muscles arranged along the valve margins. Also in contrast to ''Fordilla'', valves of ''Pojetaia'' possess between one and three teeth, with up to two teeth per valve. The exteriors of the shells show faint ribbing and fine comarginal growth lines.
''Camya'' is based on the fossils of two juvenile specimens which are both incomplete due to only the left
valve
A valve is a device or natural object that regulates, directs or controls the flow of a fluid (gases, liquids, fluidized solids, or Slurry, slurries) by opening, closing, or partially obstructing various passageways. Valves are technically Pip ...
of each being recovered. The valves have a distinct subtriangular shape and possess a long straight hinge. The umbo is positioned notably anterior on the shell and the beak is bracketed by two teeth of indistinctly pyramidal shape.
The presence of the teeth was later questioned in a 1998 study by G. Geyer and M. Streng and cited the lack of preserved muscle scars as reason to suspect the placement of ''Camya asy'' in Bivalvia.
The inner shell layers of ''Fordilla'' and ''Pojetaia'' species both consist of layers of carbonate which is akin to the laminar aragonite layer found in extant monoplacophora.
The structuring is similar to shell layering found in the extinct genera ''
Anabarella'' and ''
Watsonella'' which is thought to suggest members of the phylum
Mollusca
Mollusca is a phylum of protostome, protostomic invertebrate animals, whose members are known as molluscs or mollusks (). Around 76,000 extant taxon, extant species of molluscs are recognized, making it the second-largest animal phylum ...
developed
nacre
Nacre ( , ), also known as mother-of-pearl, is an organicinorganic composite material produced by some molluscs as an inner shell layer. It is also the material of which pearls are composed. It is strong, resilient, and iridescent.
Nacre is ...
independently several times.
Of the four accepted bivalve genera to have been described from the Cambrian, Fordilloidea contains three, ''Camya'', ''Fordilla'', and ''
Pojetaia'',
with the remaining genus ''
Tuarangia'' in the possibly related order
Tuarangiida.
References
{{Taxonbar, from=Q5468083
Bivalve taxonomy
Prehistoric animal superfamilies
Prehistoric bivalves
Taxa described in 1975