Foot Arch
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The arches of the foot, formed by the tarsal and
metatarsal The metatarsal bones or metatarsus (: metatarsi) are a group of five long bones in the midfoot, located between the tarsal bones (which form the heel and the ankle) and the phalanges ( toes). Lacking individual names, the metatarsal bones are ...
bones, strengthened by
ligaments A ligament is a type of fibrous connective tissue in the body that connects bones to other bones. It also connects flight feathers to bones, in dinosaurs and birds. All 30,000 species of amniotes (land animals with internal bones) have ligam ...
and
tendons A tendon or sinew is a tough band of dense fibrous connective tissue that connects muscle to bone. It sends the mechanical forces of muscle contraction to the skeletal system, while withstanding tension. Tendons, like ligaments, are made of ...
, allow the
foot The foot (: feet) is an anatomical structure found in many vertebrates. It is the terminal portion of a limb which bears weight and allows locomotion. In many animals with feet, the foot is an organ at the terminal part of the leg made up o ...
to support the weight of the body in the erect posture with the least weight. They are categorized as longitudinal and transverse
arch An arch is a curved vertical structure spanning an open space underneath it. Arches may support the load above them, or they may perform a purely decorative role. As a decorative element, the arch dates back to the 4th millennium BC, but stru ...
es.


Structure


Longitudinal arches

The longitudinal arches of the foot can be divided into medial and lateral arches.


Medial arch

The medial arch is higher than the lateral longitudinal arch. It is made up by the
calcaneus In humans and many other primates, the calcaneus (; from the Latin ''calcaneus'' or ''calcaneum'', meaning heel; : calcanei or calcanea) or heel bone is a bone of the Tarsus (skeleton), tarsus of the foot which constitutes the heel. In some other ...
, the talus, the navicular, the three cuneiforms (medial, intermediate, and lateral), and the first, second, and third metatarsals. Its summit is at the superior articular surface of the talus, and its two extremities or piers, on which it rests in standing, are the tuberosity on the plantar surface of the calcaneus posteriorly and the heads of the first, second, and third metatarsal bones anteriorly. The chief characteristic of this arch is its elasticity, due to its height and to the number of small joints between its component parts. Its weakest part (i.e., the part most liable to yield from overpressure) is the joint between the talus and navicular, but this portion is braced by the plantar calcaneonavicular ligament a.k.a. spring ligament, which is elastic and is thus able to quickly restore the arch to its original condition when the disturbing force is removed. The ligament is strengthened medially by blending with the
deltoid ligament The deltoid ligament (or medial ligament of talocrural joint) is a strong, flat, triangular band, attached, above, to the apex and anterior and posterior borders of the medial malleolus. The deltoid ligament supports the ankle joint and also resi ...
of the ankle joint, and is supported inferiorly by the tendon of the tibialis posterior, which is spread out in a fanshaped insertion and prevents undue tension of the ligament or such an amount of stretching as would permanently elongate it. The arch is further supported by the plantar aponeurosis, by the small muscles in the sole of the foot (short muscles of the big toe), by the tendons of the
tibialis anterior The tibialis anterior muscle is a muscle of the anterior compartment of the lower leg. It originates from the upper portion of the tibia; it inserts into the medial cuneiform and first metatarsal bones of the foot. It acts to dorsiflex and invert ...
and posterior and fibularis longus, flexor digitorum longus, flexor hallucis longus and by the ligaments of all the articulations involved.


Lateral arch

The lateral arch is composed of the calcaneus, the cuboid, and the fourth and fifth metatarsals. Two notable features of this arch are its solidity and its slight elevation. Two strong ligaments, the long plantar and the plantar calcaneocuboid, together with the extensor tendons and the short muscles of the little toe, preserve its integrity.


Fundamental longitudinal arch

While these medial and lateral arches may be readily demonstrated as the component antero-posterior arches of the foot, the fundamental longitudinal arch is contributed to by both, and consists of the calcaneus, cuboid, third cuneiform, and third metatarsal: all the other bones of the foot may be removed without destroying this arch.


Transversal arch

In addition to the longitudinal arches the foot presents a series of transverse arches. At the posterior part of the metatarsus and the anterior part of the tarsus the arches are complete, but in the middle of the tarsus they present more the characters of half-domes, the concavities of which are directed downward and medialward, so that when the medial borders of the feet are placed in apposition a complete tarsal dome is formed. The transverse arch is composed of the three cuneiforms, the cuboid, and the five metatarsal bases. The transverse arch is strengthened by the interosseous, plantar, and dorsal ligaments, by the short muscles of the first and fifth toes (especially the transverse head of the adductor hallucis), and by the fibularis longus, whose tendon stretches across between the piers of the arches.


Function

The medial longitudinal arch in particular creates a space for soft tissues with elastic properties, which act as springs, particularly the thick plantar aponeurosis, passing from the heel to the toes. Because of their elastic properties, these
soft tissue Soft tissue connective tissue, connects and surrounds or supports internal organs and bones, and includes muscle, tendons, ligaments, Adipose tissue, fat, fibrous tissue, Lymphatic vessel, lymph and blood vessels, fasciae, and synovial membranes.Β ...
s can spread ground contact reaction forces over a longer time period, and thus reduce the risk of musculoskeletal wear or damage, and they can also store the energy of these forces, returning it at the next step and thus reducing the cost of walking and, particularly, running, where vertical forces are higher.


Clinical significance

Arched feet are generally more common in men, and studies have reported that women are more likely to have flat feet than men. Women who do have arched feet who experience
pregnancy Pregnancy is the time during which one or more offspring gestation, gestates inside a woman's uterus. A multiple birth, multiple pregnancy involves more than one offspring, such as with twins. Conception (biology), Conception usually occurs ...
and may experience a flattening of their arched feet, as the surge of female sex hormones such as
estrogen Estrogen (also spelled oestrogen in British English; see spelling differences) is a category of sex hormone responsible for the development and regulation of the female reproductive system and secondary sex characteristics. There are three ...
can cause the tendons in their feet to soften and relax, to the extent that their arched feet become flat. The anatomy and shape of a person's longitudinal and transverse arch can dictate the types of injuries to which that person is susceptible. The height of a person's arch is determined by the height of the navicular bone. Collapse of the longitudinal arches results in what is known as
flat feet Flat feet, also called pes planus or fallen arches, is a Posture (psychology), postural deformity in which the arches of the foot collapse, with the entire sole (foot), sole of the foot coming into complete or near-complete contact with the gro ...
. A person with a low longitudinal arch, or flat feet will likely stand and walk with their feet in a pronated position, where the foot everts or rolls inward. This makes the person susceptible to heel pain, arch pain and
plantar fasciitis Plantar fasciitis or plantar heel pain is a disorder of the plantar fascia, which is the connective tissue that supports the Arches of the foot, arch of the foot. It results in pain in the heel and Sole (foot), bottom of the foot that is usuall ...
. Flat footed people may also have more difficulty performing exercises that require supporting their weight on their toes. People who have high longitudinal arches or a cavus foot tend to walk and stand with their feet in a supinated position where the foot inverts or rolls outward. High arches can also cause plantar fasciitis as they cause the plantar fascia to be stretched away from the calcaneus or heel bone. Additionally, high or low arches can increase the risk of shin splints as the anterior tibialis must work harder to keep the foot from slapping the ground.


Evolution and other animals

The non-
human Humans (''Homo sapiens'') or modern humans are the most common and widespread species of primate, and the last surviving species of the genus ''Homo''. They are Hominidae, great apes characterized by their Prehistory of nakedness and clothing ...
ape Apes (collectively Hominoidea ) are a superfamily of Old World simians native to sub-Saharan Africa and Southeast Asia (though they were more widespread in Africa, most of Asia, and Europe in prehistory, and counting humans are found global ...
s (the
gibbon Gibbons () are apes in the family Hylobatidae (). The family historically contained one genus, but now is split into four extant genera and 20 species. Gibbons live in subtropical and tropical forests from eastern Bangladesh and Northeast Indi ...
s,
gorilla Gorillas are primarily herbivorous, terrestrial great apes that inhabit the tropical forests of equatorial Africa. The genus ''Gorilla'' is divided into two species: the eastern gorilla and the western gorilla, and either four or five su ...
s,
orangutan Orangutans are great apes native to the rainforests of Indonesia and Malaysia. They are now found only in parts of Borneo and Sumatra, but during the Pleistocene they ranged throughout Southeast Asia and South China. Classified in the genus ...
s,
chimpanzee The chimpanzee (; ''Pan troglodytes''), also simply known as the chimp, is a species of Hominidae, great ape native to the forests and savannahs of tropical Africa. It has four confirmed subspecies and a fifth proposed one. When its close rel ...
s and
bonobo The bonobo (; ''Pan paniscus''), also historically called the pygmy chimpanzee (less often the dwarf chimpanzee or gracile chimpanzee), is an endangered great ape and one of the two species making up the genus ''Pan (genus), Pan'' (the other bei ...
s) tend to walk on the lateral side of the foot, that is with an 'inverted' foot, which may reflect a basic adaptation to walking on branches. It is often held that their feet lack longitudinal arches, but footprints made by bipedally walking apes, which must directly or indirectly reflect the pressure they exert to support and propel themselves do suggest that they exert lower foot pressure under the medial part of their midfoot. However, human feet, and the human medial longitudinal arch, differ in that the anterior part of the foot is medially twisted on the posterior part of the foot, so that all the toes may contact the ground at the same time, and the twisting is so marked that the most medial toe, the big toe or hallux, (in some individuals the second toe) tends to exert the greatest propulsive force in walking and running. This gives the human foot an 'everted' or relatively outward-facing appearance compared to that of other apes. The strong twisting of the anterior part of the human foot on the posterior part tends to increase the height of the medial longitudinal arch. However, there is now considerable evidence that shoe-wearing also accentuates the height of the medial longitudinal arch and that the height of the medial longitudinal arch also differs very considerably between individuals and at different speeds. The presence of high-arched feet in modern humans is a result of natural selection for long-distance running. On the other hand, the primitive trait of arch-less feet in our
great ape The Hominidae (), whose members are known as the great apes or hominids (), are a taxonomic family of primates that includes eight extant species in four genera: '' Pongo'' (the Bornean, Sumatran and Tapanuli orangutan); '' Gorilla'' (the ...
relatives has been maintained because of selection for grasping tree branches as a part of their arboreal lifestyle. Divergence between ape feet and human feet began with the early human ancestor '' Ardipithecus ramidus'', when strengthened plantar tissue evolved, which supported early terrestrial propulsion before evolving a true arch. However, the skeletal longitudinal arch structure itself did not begin to evolve until ''
Australopithecus afarensis ''Australopithecus afarensis'' is an extinct species of australopithecine which lived from about 3.9–2.9 million years ago (mya) in the Pliocene of East Africa. The first fossils were discovered in the 1930s, but major fossil finds would not ta ...
'' had evolved a relatively low longitudinal arch (compared to modern humans) and the first signs of a transverse arch accompanying it. It is not yet agreed to what extent the early human ancestor ''
Australopithecus afarensis ''Australopithecus afarensis'' is an extinct species of australopithecine which lived from about 3.9–2.9 million years ago (mya) in the Pliocene of East Africa. The first fossils were discovered in the 1930s, but major fossil finds would not ta ...
'', (3.75 million years ago onwards) had acquired a functionally human-like foot, but the medial twist of the forefoot evident in fossil footbones of this species, and in the
Laetoli Laetoli is a pre-historic site located in Enduleni ward of Ngorongoro District in Arusha Region, Tanzania. The site is dated to the Plio-Pleistocene and famous for its Hominina footprints, preserved in volcanic ash. The site of the Laetoli footp ...
footprint trail in Tanzania generally attributed to this species, certainly appears less marked than is evident in fossil footbones of ''
Homo erectus ''Homo erectus'' ( ) is an extinction, extinct species of Homo, archaic human from the Pleistocene, spanning nearly 2 million years. It is the first human species to evolve a humanlike body plan and human gait, gait, to early expansions of h ...
'' (sometimes called '' Homo georgicus'') from
Dmanisi Dmanisi ( ka, αƒ“αƒ›αƒαƒœαƒ˜αƒ‘αƒ˜, tr, , ) is a town and archaeological site in the Kvemo Kartli region of Georgia (country), Georgia approximately 93 km southwest of the nation’s capital Tbilisi in the river valley of Mashavera. Abandoned i ...
, Georgia (c. 1. 8 million years ago) and the roughly contemporaneous fossil footprint trail at Ileret, Kenya attributed to '' Homo erectus ergaster''.


See also

* Pes cavus *
Flat feet Flat feet, also called pes planus or fallen arches, is a Posture (psychology), postural deformity in which the arches of the foot collapse, with the entire sole (foot), sole of the foot coming into complete or near-complete contact with the gro ...
* Arch support * Comparison of orthotics * Foot gymnastics


References


Bibliography

* * * * * * * * *


External links


Overview at gla.ac.uk



Arches of the foot
{{DEFAULTSORT:Arches Of The Foot Foot