''Fockea multiflora'', or python vine, is a plant of the dogbane family,

Apocynaceae Apocynaceae (, from '' Apocynum'', Greek for "dog-away") is a family of flowering plants that includes trees, shrubs, herbs, stem succulents, and vines, commonly known as the dogbane family, because some taxa were used as dog poison. Notable mem ...

, native to

Tanzania Tanzania, officially the United Republic of Tanzania, is a country in East Africa within the African Great Lakes region. It is bordered by Uganda to the northwest; Kenya to the northeast; the Indian Ocean to the east; Mozambique and Malawi to t ...

Mozambique Mozambique, officially the Republic of Mozambique, is a country located in Southeast Africa bordered by the Indian Ocean to the east, Tanzania to the north, Malawi and Zambia to the northwest, Zimbabwe to the west, and Eswatini and South Afr ...

Zimbabwe file:Zimbabwe, relief map.jpg, upright=1.22, Zimbabwe, relief map Zimbabwe, officially the Republic of Zimbabwe, is a landlocked country in Southeast Africa, between the Zambezi and Limpopo Rivers, bordered by South Africa to the south, Bots ...

Zambia Zambia, officially the Republic of Zambia, is a landlocked country at the crossroads of Central Africa, Central, Southern Africa, Southern and East Africa. It is typically referred to being in South-Central Africa or Southern Africa. It is bor ...

Angola Angola, officially the Republic of Angola, is a country on the west-Central Africa, central coast of Southern Africa. It is the second-largest Portuguese-speaking world, Portuguese-speaking (Lusophone) country in both total area and List of c ...

Botswana Botswana, officially the Republic of Botswana, is a landlocked country in Southern Africa. Botswana is topographically flat, with approximately 70 percent of its territory part of the Kalahari Desert. It is bordered by South Africa to the sou ...

Namibia Namibia, officially the Republic of Namibia, is a country on the west coast of Southern Africa. Its borders include the Atlantic Ocean to the west, Angola and Zambia to the north, Botswana to the east and South Africa to the south; in the no ...

, including the

Caprivi Strip The Caprivi Strip, also known simply as Caprivi, is a geographic salient protruding from the northeastern corner of Namibia. It is bordered by Botswana to the south and Angola and Zambia to the north. Namibia, Botswana and Zambia meet at a sing ...

, and

Malawi Malawi, officially the Republic of Malawi, is a landlocked country in Southeastern Africa. It is bordered by Zambia to the west, Tanzania to the north and northeast, and Mozambique to the east, south, and southwest. Malawi spans over and ...

. It is a large semisucculent

liana A liana is a long-Plant stem, stemmed Woody plant, woody vine that is rooted in the soil at ground level and uses trees, as well as other means of vertical support, to climb up to the Canopy (biology), canopy in search of direct sunlight. T ...

, growing to some 15m in length and up to 60 cm in diameter, found primarily in the seasonally dry tropical biome.

Taxonomy

The genus ''

Fockea ''Fockea'' is a genus of succulent scrubs native to Africa south of the equator. They are members of the Asclepiadoideae (milkweeds), a subfamily of the dogbane family Apocynaceae. Of the six accepted species, only the two most widely distributed ...

'' comprises six species belonging to the

Asclepiadoideae The Asclepiadoideae are a subfamily of plants in the family Apocynaceae. Formerly, it was treated as a separate family under the name Asclepiadaceae, e.g. by APG II, and known as the milkweed family. They form a group of perennial herbs, twinin ...

(milkweeds)

subfamily In biological classification, a subfamily (Latin: ', plural ') is an auxiliary (intermediate) taxonomic rank, next below family but more inclusive than genus. Standard nomenclature rules end botanical subfamily names with "-oideae", and zo ...

of the

family. Massive specimens of ''F. multiflora'' are probably the largest known members of this subfamily. All six ''Fockea'' species occur south of the equator in Africa, with ''F. multiflora'' the second most widely distributed, after ''F. angustifolia''. Unlike the other five ''Fockea'' species, all of which are relatively small climbers with swollen, mostly subterranean

tuber Tubers are a type of enlarged structure that plants use as storage organs for nutrients, derived from stems or roots. Tubers help plants perennate (survive winter or dry months), provide energy and nutrients, and are a means of asexual reproduc ...

s, ''F. multiflora'' is a massive, tropical

without a tuber, It is considered a

sister A sister is a woman or a girl who shares parents or a parent with another individual; a female sibling. The male counterpart is a brother. Although the term typically refers to a familial relationship, it is sometimes used endearingly to ref ...

to the other five species.

Description

''Fockea multiflora'' is a large climber up to 15 m, with a thick, fleshy trunk up to 30 cm thick, swollen toward the base but lacking a distinct basal tuber. In mature plants the rootstock consists of a network of fleshy roots radiating from the base of the stem. By comparison, in the other five ''Fockea'' species, stems arise from a tuber that is much broader than the main stem. ''F. multiflora'' is rarely shrub-like; rather, its stems sprawl on the ground or twist around surrounding trees as massive lianas, appearing to strangle them, although there’s no evidence that the supporting trees are harmed. The stems produces a white, milky latex. Young stems are tomentose (covered with densely matted wooly hairs), later becoming covered with grey, shiny bark. Leaves are opposite, broadly elliptic to ovate, large (100mm x 80mm), with grey-felted undersides, felty to smooth above. These leaves, as well as the fruits and seeds, are much larger than for other ''Fockea'' species.

Inflorescence In botany, an inflorescence is a group or cluster of flowers arranged on a plant's Plant stem, stem that is composed of a main branch or a system of branches. An inflorescence is categorized on the basis of the arrangement of flowers on a mai ...

is a many-flowered axillary cluster of as many as 30 flowers approximately 15mm in diameter, grayish-green on the outside and yellow-green to brown on the inside, with flowers opening simultaneously or in rapid succession. The fruits are smooth, paired and horn-shaped, 10–22 cm × 1.5–3 cm, dehiscing to release multiple winged seeds; seeds are ovate and flattened, 10 mm × 7–8 mm, shortly winged.http://database.prota.org/PROTAhtml/Fockea%20multiflora_En.htm

Ecology

Growing in the altitude range of 600–1000 m, ''Fockea multiflora'' occurs on low hills or among rocks around the base of hills in dry, open, often deciduous woodland or scrub, especially mixed ''

Acacia ''Acacia'', commonly known as wattles or acacias, is a genus of about of shrubs and trees in the subfamily Mimosoideae of the pea family Fabaceae. Initially, it comprised a group of plant species native to Africa, South America, and Austral ...

''-''

Commiphora ''Commiphora'' is the most species-rich genus of flowering plants in the frankincense and myrrh family, Burseraceae. The genus contains approximately 190 species of shrubs and trees, which are distributed throughout the (sub-) tropical regions of A ...

''-''

Balanites ''Balanites'' is an Afrotropical, Palearctic and Indomalayan genus of flowering plants in the caltrop family, Zygophyllaceae. The name ''Balanites'' derives from the Greek word for an acorn and refers to the fruit. It was coined by Alire Del ...

'' or

mopane ''Colophospermum mopane'', commonly called mopane, mopani, butterfly tree, turpentine tree, or balsam tree, is a tree in the legume family (Fabaceae), that grows in hot, dry, low-lying areas, in elevation, in parts of Southern Africa. The tree ...

woodland. It does not tend to occur in more mesic

miombo Miombo woodland is a tropical and subtropical grasslands, savannas, and shrublands biome (in the World Wide Fund for Nature scheme) located in central and southern tropical Africa. It includes three woodland savanna ecoregions (listed below) chara ...

woodland dominated by ''

Brachystegia ''Brachystegia'' is a genus of tree of the subfamily Detarioideae that is native to tropical Africa. Trees of the genus are commonly known as miombo, and are dominant in the miombo woodlands of central and southern tropical Africa. The Zambezia ...

''.

References

{{Taxonbar, from=Q15398617 multiflora Flora of Botswana Flora of the Caprivi Strip Flora of Namibia Flora of South Tropical Africa Flora of Tanzania