Fockea
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''Fockea'' is a genus of succulent scrubs native to
Africa Africa is the world's second-largest and second-most populous continent after Asia. At about 30.3 million km2 (11.7 million square miles) including adjacent islands, it covers 20% of Earth's land area and 6% of its total surfac ...
south of the equator. They are members of the
Asclepiadoideae The Asclepiadoideae are a subfamily of plants in the family Apocynaceae. Formerly, it was treated as a separate family under the name Asclepiadaceae, e.g. by APG II, and known as the milkweed family. They form a group of perennial herbs, twinin ...
(milkweeds), a
subfamily In biological classification, a subfamily (Latin: ', plural ') is an auxiliary (intermediate) taxonomic rank, next below family but more inclusive than genus. Standard nomenclature rules end botanical subfamily names with "-oideae", and zo ...
of the dogbane family
Apocynaceae Apocynaceae (, from '' Apocynum'', Greek for "dog-away") is a family of flowering plants that includes trees, shrubs, herbs, stem succulents, and vines, commonly known as the dogbane family, because some taxa were used as dog poison. Notable mem ...
. Of the six accepted species, only the two most widely distributed extend north of
southern Africa Southern Africa is the southernmost region of Africa. No definition is agreed upon, but some groupings include the United Nations geoscheme for Africa, United Nations geoscheme, the intergovernmental Southern African Development Community, and ...
, with '' F. multiflora'' reaching as far north as Tanzania and ''F. angustifolia'' reaching to southern
Kenya Kenya, officially the Republic of Kenya, is a country located in East Africa. With an estimated population of more than 52.4 million as of mid-2024, Kenya is the 27th-most-populous country in the world and the 7th most populous in Africa. ...
. ''Fockea'' are known as water roots, a reference to the bulbous
caudex A caudex (: caudices) of a plant is a stem, but the term is also used to mean a rootstock and particularly a basal stem structure from which new growth arises.pages 456 and 695 In the strict sense of the term, meaning a stem, "caudex" is most ...
characteristic of most species, which is also edible in at least some species.


Taxonomy

The genus ''Fockea'' Endl. was established in 1838 by the Austrian botanist
Stephan Ladislaus Endlicher Stephan Friedrich Ladislaus Endlicher, also known as Endlicher István László (24 June 1804 – 28 March 1849), was an Austrian botanist, numismatist and Sinologist. He was a director of the Botanical Garden of Vienna. Biography Endlicher ...
through illustration and description of a specimen of ''Fockea capensis'' collected in
Cape Colony The Cape Colony (), also known as the Cape of Good Hope, was a British Empire, British colony in present-day South Africa named after the Cape of Good Hope. It existed from 1795 to 1802, and again from 1806 to 1910, when it united with three ...
circa 1786 by
Franz Boos Franz Boos (23 December 1753 in Frauenalb – 9 February 1832 in Vienna) was an Austrian List of gardener-botanist explorers of the Enlightenment, gardener-botanist in the Age of Enlightenment, a voyager and collector of natural history specimens ...
and Georg Scholl and cultivated at
Schönbrunn Garden Schönbrunn may refer to: * Schönbrunn Palace in Vienna, Austria * Schönbrunn Station of the Vienna U-Bahn * Schönbrunn Zoo in Schönbrunn Palace gardens * Schönbrunn (Baden), a municipality in Rhein-Neckar, Baden-Württemberg, Germany * Schö ...
in Vienna. The genus was named after the German physician and naturalist
Gustav Woldemar Focke Gustav Woldemar Focke (24 January 1810, Bremen – 1 June 1877, Bremen) was a German physician and naturalist. He was a nephew of naturalist Gottfried Reinhold Treviranus. He studied medicine at the University of Heidelberg, receiving his do ...
, author of the commentary ''De respiratione vegetabilium'' (''Of the respiration of vegetables''). The "Old Lady of Schönbrunn" — the oldest potted succulent in cultivation — continued to be cultivated at least through 1988, over 200 years after it was first collected by Boos and Scholl. It was believed to be the last surviving member of its species until the South African botanist
Rudolf Marloth Hermann Wilhelm Rudolf Marloth (28 December 1855 Lübben, Germany – 15 May 1931 Caledon, Cape Province) was a German-born South African botanist, pharmacist and analytical chemist, best known for his ''Flora of South Africa'' which appeared i ...
collected another specimen of ''F. capensis'' near
Prince Albert Prince Albert most commonly refers to: *Prince Albert of Saxe-Coburg and Gotha (1819–1861), consort of Queen Victoria *Albert II, Prince of Monaco (born 1958), present head of state of Monaco Prince Albert may also refer to: Royalty * Alb ...
in 1906. Most ''Fockea'' species are relatively small climbers with swollen, mostly subterranean
tuber Tubers are a type of enlarged structure that plants use as storage organs for nutrients, derived from stems or roots. Tubers help plants perennate (survive winter or dry months), provide energy and nutrients, and are a means of asexual reproduc ...
s, whereas ''
Fockea multiflora ''Fockea multiflora'', or python vine, is a plant of the dogbane family, Apocynaceae, native to Tanzania, Mozambique, Zimbabwe, Zambia, Angola, Botswana, Namibia, including the Caprivi Strip, and Malawi. It is a large semisucculent liana, growi ...
'', a widely distributed but exclusively tropical species, is a massive, tropical
liana A liana is a long-Plant stem, stemmed Woody plant, woody vine that is rooted in the soil at ground level and uses trees, as well as other means of vertical support, to climb up to the Canopy (biology), canopy in search of direct sunlight. T ...
without a tuber. It is considered a
sister A sister is a woman or a girl who shares parents or a parent with another individual; a female sibling. The male counterpart is a brother. Although the term typically refers to a familial relationship, it is sometimes used endearingly to ref ...
to the other five species. ''
Fockea angustifolia ''Fockea'' is a genus of succulent scrubs native to Africa south of the equator. They are members of the Asclepiadoideae (milkweeds), a subfamily of the dogbane family Apocynaceae. Of the six accepted species, only the two most widely distribute ...
'', also widely distributed, is mainly tropical, and sister to the remaining four species, which are
endemic Endemism is the state of a species being found only in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also foun ...
to southern Africa. ;Species # ''
Fockea angustifolia ''Fockea'' is a genus of succulent scrubs native to Africa south of the equator. They are members of the Asclepiadoideae (milkweeds), a subfamily of the dogbane family Apocynaceae. Of the six accepted species, only the two most widely distribute ...
'' K.Schum. — A climbing tuberous
geophyte A storage organ is a part of a plant specifically modified for storage of energy (generally in the form of carbohydrates) or water. Storage organs often grow underground, where they are better protected from attack by herbivores. Plants that have ...
found primarily in the seasonally dry tropical biome, ranging from Southeast
Kenya Kenya, officially the Republic of Kenya, is a country located in East Africa. With an estimated population of more than 52.4 million as of mid-2024, Kenya is the 27th-most-populous country in the world and the 7th most populous in Africa. ...
to
South Africa South Africa, officially the Republic of South Africa (RSA), is the Southern Africa, southernmost country in Africa. Its Provinces of South Africa, nine provinces are bounded to the south by of coastline that stretches along the Atlantic O ...
. # ''
Fockea capensis ''Fockea'' is a genus of succulent scrubs native to Africa south of the equator. They are members of the Asclepiadoideae (milkweeds), a subfamily of the dogbane family Apocynaceae. Of the six accepted species, only the two most widely distributed ...
'' Endl. — A climbing tuberous geophyte found primarily in the subtropical biome; its native range is the southern
Cape Provinces The Cape Provinces of South Africa is a biogeographical area used in the World Geographical Scheme for Recording Plant Distributions (WGSRPD). It is part of the WGSRPD region 27 Southern Africa. The area has the code "CPP". It includes the Sou ...
of South Africa. According to Jacobsen, the turnip-shaped tuber can be up to diameter. # ''
Fockea comaru ''Fockea'' is a genus of succulent scrubs native to Africa south of the equator. They are members of the Asclepiadoideae (milkweeds), a subfamily of the dogbane family Apocynaceae. Of the six accepted species, only the two most widely distribute ...
'' (E.Mey.) N.E.Br. - A tuberous geophyte found primarily in the desert or dry shrubland biome, ranging from southern
Namibia Namibia, officially the Republic of Namibia, is a country on the west coast of Southern Africa. Its borders include the Atlantic Ocean to the west, Angola and Zambia to the north, Botswana to the east and South Africa to the south; in the no ...
to South Africa's Cape Provinces. # ''
Fockea edulis ''Fockea edulis'' is a species of caudiciform plant in the family Apocynaceae that is native to the Cape Provinces and KwaZulu-Natal in South Africa South Africa, officially the Republic of South Africa (RSA), is the Southern Africa, s ...
'' (Thunb.) K.Schum. - A climbing tuberous geophyte found primarily in the subtropical biome, ranging from the southern Cape Provinces to
KwaZulu-Natal KwaZulu-Natal (, also referred to as KZN) is a Provinces of South Africa, province of South Africa that was created in 1994 when the government merged the Zulu people, Zulu bantustan of KwaZulu ("Place of the Zulu" in Zulu language, Zulu) and ...
in South Africa. # ''
Fockea multiflora ''Fockea multiflora'', or python vine, is a plant of the dogbane family, Apocynaceae, native to Tanzania, Mozambique, Zimbabwe, Zambia, Angola, Botswana, Namibia, including the Caprivi Strip, and Malawi. It is a large semisucculent liana, growi ...
'' K.Schum. - A semisucculent
liana A liana is a long-Plant stem, stemmed Woody plant, woody vine that is rooted in the soil at ground level and uses trees, as well as other means of vertical support, to climb up to the Canopy (biology), canopy in search of direct sunlight. T ...
growing primarily in the seasonally dry tropical biome, ranging from Tanzania to northern Namibia. # ''
Fockea sinuata ''Fockea'' is a genus of succulent scrubs native to Africa south of the equator. They are members of the Asclepiadoideae (milkweeds), a subfamily of the dogbane family Apocynaceae. Of the six accepted species, only the two most widely distribute ...
'' (E.Mey.) Druce - A climbing tuberous geophyte growing primarily in the desert or dry shrubland biome, ranging from south-central and southern Namibia to the Cape Provinces and Free State province of South Africa.


References

{{Taxonbar, from=Q5863412 Apocynaceae genera Caudiciform plants Taxa named by Stephan Endlicher