
A flyway is a flight path used by large numbers of birds while
migrating between their
breeding grounds and their
overwintering
Overwintering is the process by which some organisms pass through or wait out the winter season, or pass through that period of the year when "winter" conditions (cold or sub-zero temperatures, ice, snow, limited food supplies) make normal activ ...
quarters. Flyways generally span
continent
A continent is any of several large geographical regions. Continents are generally identified by convention (norm), convention rather than any strict criteria. A continent could be a single large landmass, a part of a very large landmass, as ...
s and often pass over
ocean
The ocean is the body of salt water that covers approximately 70.8% of Earth. The ocean is conventionally divided into large bodies of water, which are also referred to as ''oceans'' (the Pacific, Atlantic, Indian Ocean, Indian, Southern Ocean ...
s. Although applying to any species of migrating bird, the concept was first conceived and applied to
waterfowl
Anseriformes is an order of birds also known as waterfowl that comprises about 180 living species of birds in three families: Anhimidae (three species of screamers), Anseranatidae (the magpie goose), and Anatidae, the largest family, which i ...
and
shore birds. The flyways can be thought of as wide
arterial highways to which the migratory routes of different species are
tributaries
A tributary, or an ''affluent'', is a stream or river that flows into a larger stream ('' main stem'' or ''"parent"''), river, or a lake. A tributary does not flow directly into a sea or ocean. Tributaries, and the main stem river into which the ...
.
An alternative definition is that a flyway is the entire range of a migratory bird, encompassing both its breeding and non-breeding grounds, and the resting and feeding locations it uses while migrating. There are four major north–south flyways in North America and six covering Eurasia, Africa, and Australasia.
History
The passing of the
Migratory Bird Treaty Act of 1918
The Migratory Bird Treaty Act of 1918 (MBTA), codified at (although §709 is omitted), is a United States federal law, first enacted in 1918 to implement the convention for the protection of migratory birds between the United States and Canada. ...
in the United States resulted in a need for more information on bird migration.
Frederick Charles Lincoln was put in charge and improved methods for trapping and
banding, developed record-keeping procedures, recruited banders, fostered international cooperation, and promoted banding as a tool for research and wildlife management. He found it was possible to establish the routes used by waterfowl during their annual migrations and he developed the flyway concept, a key idea in the management and regulation of hunting of migratory birds; by establishing the routes used, estimates of population sizes could be made and suitable protection could be put in place.

The special vulnerability of waterfowl and shorebirds on their international migrations, with their specific needs for suitable wetland stopovers, resulted in the signing of the
Ramsar Convention
The Ramsar Convention on Wetlands of International Importance Especially as Waterfowl Habitat is an international treaty for the conservation and sustainable use of Ramsar site, Ramsar sites (wetlands). It is also known as the Convention on We ...
in 1971. As a result, over 2300
Ramsar sites have been established around the world, many being situated on flyways where they provide the vital habitat needed by the birds on their journeys.
Flyways
Many bird populations migrate long distances twice a year. The most common pattern involves flying north in the spring to breed in the temperate parts of the northern hemisphere or the
Arctic
The Arctic (; . ) is the polar regions of Earth, polar region of Earth that surrounds the North Pole, lying within the Arctic Circle. The Arctic region, from the IERS Reference Meridian travelling east, consists of parts of northern Norway ( ...
during summer and returning southward in the autumn to wintering grounds in warmer regions, often on the other side of the equator. A similar pattern occurs in the southern hemisphere with birds flying south to breed and north to overwinter, but on a much smaller scale. The flyway, or route, taken by different bird species varies, but each population has its traditional staging points along the route where birds feed to build up their energy reserves to prepare for the next migratory stage; the route used on the spring migration may be different from that used in the autumn and will depend on such factors as wind direction and the availability of food at staging points.
Flyways may not be the shortest route available but may have curves or doglegs. Birds of different species may follow similar routes, and populations from one area may merge with other groups and diverge to reach different destinations. Flyways tend to avoid obstacles such as mountain ranges and oceans, running parallel to the barriers and following routes along the coast or along major river valleys. Passerines often fly on a broader front across the terrain, either flying over or circumventing obstacles on the route, according to their evolutionary adaptations. In selecting routes, birds may overcompensate for predicted winds. Terrestrial birds tend to travel over land, raptors need routes where
thermal
A thermal column (or thermal) is a rising mass of buoyant air, a convective current in the atmosphere, that transfers heat energy vertically. Thermals are created by the uneven heating of Earth's surface from solar radiation, and are an example ...
s can give them the lift they require, sea birds prefer ocean routes and wetland birds need routes with suitable staging sites; deltas and coastal wetlands provide reliable food sources for this purpose whereas inland wetlands are less predictable.
[
]
Flyways of the Americas
In North America
North America is a continent in the Northern Hemisphere, Northern and Western Hemisphere, Western hemispheres. North America is bordered to the north by the Arctic Ocean, to the east by the Atlantic Ocean, to the southeast by South Ameri ...
, the flyways used by migratory waterfowl
Anseriformes is an order of birds also known as waterfowl that comprises about 180 living species of birds in three families: Anhimidae (three species of screamers), Anseranatidae (the magpie goose), and Anatidae, the largest family, which i ...
are divided into four geographical paths. Each flyway has a different composition of species
A species () is often defined as the largest group of organisms in which any two individuals of the appropriate sexes or mating types can produce fertile offspring, typically by sexual reproduction. It is the basic unit of Taxonomy (biology), ...
and habitat
In ecology, habitat refers to the array of resources, biotic factors that are present in an area, such as to support the survival and reproduction of a particular species. A species' habitat can be seen as the physical manifestation of its ...
. The United States Fish and Wildlife Service
The United States Fish and Wildlife Service (USFWS or FWS) is a List of federal agencies in the United States, U.S. federal government agency within the United States Department of the Interior which oversees the management of fish, wildlife, ...
established the flyways to help with the management of migratory birds
Bird migration is a seasonal movement of birds between breeding and wintering grounds that occurs twice a year. It is typically from north to south or from south to north. Migration is inherently risky, due to predation and mortality.
The ...
. They studied all migratory birds and established the Mississippi Flyway
The Mississippi Flyway is a bird migration route that generally follows the Mississippi, Missouri, and Lower Ohio Rivers in the United States across the western Great Lakes to the Mackenzie River and Hudson Bay in Canada. The main endpoints of t ...
, Atlantic Flyway, Mountain Flyway, and Pacific Flyway
The Pacific Flyway is a major north-south flyway for migratory birds in the Americas, extending from Alaska to Patagonia. Every year, migratory birds travel some or all of this distance both in spring and in fall, following food sources, heading ...
.
The Atlantic Flyway starts in northern Canada and Greenland
Greenland is an autonomous territory in the Danish Realm, Kingdom of Denmark. It is by far the largest geographically of three constituent parts of the kingdom; the other two are metropolitan Denmark and the Faroe Islands. Citizens of Greenlan ...
and follows the East Coast of the United States
The East Coast of the United States, also known as the Eastern Seaboard, the Atlantic Coast, and the Atlantic Seaboard, is the region encompassing the coast, coastline where the Eastern United States meets the Atlantic Ocean; it has always pla ...
to the Caribbean Sea
The Caribbean Sea is a sea of the Atlantic Ocean, North Atlantic Ocean in the tropics of the Western Hemisphere, located south of the Gulf of Mexico and southwest of the Sargasso Sea. It is bounded by the Greater Antilles to the north from Cuba ...
, and on to the tropical
The tropics are the regions of Earth surrounding the equator, where the sun may shine directly overhead. This contrasts with the temperate or polar regions of Earth, where the Sun can never be directly overhead. This is because of Earth's ax ...
Central America
Central America is a subregion of North America. Its political boundaries are defined as bordering Mexico to the north, Colombia to the southeast, the Caribbean to the east, and the Pacific Ocean to the southwest. Central America is usually ...
. Examples of birds in this flyway are the American Goldfinch, Blue Jay, and Northern Cardinal.
The Mississippi Flyway
The Mississippi Flyway is a bird migration route that generally follows the Mississippi, Missouri, and Lower Ohio Rivers in the United States across the western Great Lakes to the Mackenzie River and Hudson Bay in Canada. The main endpoints of t ...
starts from northeastern Canada and passes over the Great Lakes, following the lower Ohio River
The Ohio River () is a river in the United States. It is located at the boundary of the Midwestern and Southern United States, flowing in a southwesterly direction from Pittsburgh, Pennsylvania, to its river mouth, mouth on the Mississippi Riv ...
, the Missouri
Missouri (''see #Etymology and pronunciation, pronunciation'') is a U.S. state, state in the Midwestern United States, Midwestern region of the United States. Ranking List of U.S. states and territories by area, 21st in land area, it border ...
and the Mississippi
Mississippi ( ) is a U.S. state, state in the Southeastern United States, Southeastern and Deep South regions of the United States. It borders Tennessee to the north, Alabama to the east, the Gulf of Mexico to the south, Louisiana to the s ...
to the Gulf of Mexico
The Gulf of Mexico () is an oceanic basin and a marginal sea of the Atlantic Ocean, mostly surrounded by the North American continent. It is bounded on the northeast, north, and northwest by the Gulf Coast of the United States; on the southw ...
, and on to Central and South America. There are no mountains on this route.[ Examples of birds in this flyway are the Black-capped Chickadee, Baltimore Oriole, and House Finch.
The Central Flyway starts from central Canada and crosses the ]Great Plains
The Great Plains is a broad expanse of plain, flatland in North America. The region stretches east of the Rocky Mountains, much of it covered in prairie, steppe, and grassland. They are the western part of the Interior Plains, which include th ...
before continuing southwards to the Gulf of Mexico, merging with the Mississippi Flyway. There are no mountain barriers on this route. Examples of birds in this flyway are the American Crow, American Robin, Common Grackle.
The Pacific Flyway
The Pacific Flyway is a major north-south flyway for migratory birds in the Americas, extending from Alaska to Patagonia. Every year, migratory birds travel some or all of this distance both in spring and in fall, following food sources, heading ...
is a north–south flyway for birds migrating from breeding grounds in Alaska
Alaska ( ) is a non-contiguous U.S. state on the northwest extremity of North America. Part of the Western United States region, it is one of the two non-contiguous U.S. states, alongside Hawaii. Alaska is also considered to be the north ...
and Canada to their overwintering areas in South America, some species travelling as far south as Patagonia
Patagonia () is a geographical region that includes parts of Argentina and Chile at the southern end of South America. The region includes the southern section of the Andes mountain chain with lakes, fjords, temperate rainforests, and glaciers ...
.[ Examples of birds in this flyway are the Northern Flicker, Purple Finch, and Rufous Hummingbird.
The Allegheny Front flyway in the central ]Appalachian Mountains
The Appalachian Mountains, often called the Appalachians, are a mountain range in eastern to northeastern North America. The term "Appalachian" refers to several different regions associated with the mountain range, and its surrounding terrain ...
is an important flyway for migratory birds traveling from their northern breeding grounds to their southern wintering sites.
Flyways of Eurasia, Africa, and Australasia
The East Atlantic Flyway starts from northern North America, Greenland
Greenland is an autonomous territory in the Danish Realm, Kingdom of Denmark. It is by far the largest geographically of three constituent parts of the kingdom; the other two are metropolitan Denmark and the Faroe Islands. Citizens of Greenlan ...
, Iceland
Iceland is a Nordic countries, Nordic island country between the Atlantic Ocean, North Atlantic and Arctic Oceans, on the Mid-Atlantic Ridge between North America and Europe. It is culturally and politically linked with Europe and is the regi ...
, northern Europe and western Siberia and leads to wintering areas in western Europe and North Africa, with some birds continuing down the west coast of the continent to South Africa.[
The Black Sea-Mediterranean Flyway starts from northern and western Siberia and leads across Asia, the ]Black Sea
The Black Sea is a marginal sea, marginal Mediterranean sea (oceanography), mediterranean sea lying between Europe and Asia, east of the Balkans, south of the East European Plain, west of the Caucasus, and north of Anatolia. It is bound ...
and the Mediterranean Sea
The Mediterranean Sea ( ) is a sea connected to the Atlantic Ocean, surrounded by the Mediterranean basin and almost completely enclosed by land: on the east by the Levant in West Asia, on the north by Anatolia in West Asia and Southern Eur ...
to northern Africa. Little has been published about birds using this flyway.[
The Asian–East African Flyway starts from the northern breeding grounds of water birds in Siberia and leads across Asia to East Africa. Little has been published about birds using this flyway.][
The Central Asian-Indian Flyway starts from the northern breeding grounds of water birds in Siberia and leads across Asia to the Indian subcontinent. Little has been published about birds using this flyway.][
The East Asian–Australasian Flyway starts at the ]Taymyr Peninsula
The Taymyr Peninsula ( ) is a peninsula in the Far North of Russia, in the Siberian Federal District, that forms the northernmost part of the mainland of Eurasia. Administratively it is part of the Krasnoyarsk Krai Federal subject of Russia.
Ge ...
in Russia and Alaska
Alaska ( ) is a non-contiguous U.S. state on the northwest extremity of North America. Part of the Western United States region, it is one of the two non-contiguous U.S. states, alongside Hawaii. Alaska is also considered to be the north ...
and extends southwards to southeastern Asia, Australia and New Zealand. This flyway overlaps with the West Pacific Flyway. About 60 species of shorebird use this route.[
The West Pacific Flyway links New Zealand and the east coast of Australia, through the central Pacific Ocean and the east coast of northern Asia, including Japan and the Korean Peninsula, ending up in eastern Siberia, including the Chukchi and ]Kamchatka
The Kamchatka Peninsula (, ) is a peninsula in the Russian Far East, with an area of about . The Pacific Ocean and the Sea of Okhotsk make up the peninsula's eastern and western coastlines, respectively.
Immediately offshore along the Pacific ...
peninsulas, and Alaska. This flyway overlaps with the East Asian–Australasian Flyway.[
]
References
{{reflist