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The Floreana mockingbird (''Mimus trifasciatus'') or the Charles Island mockingbird, is a
species A species () is often defined as the largest group of organisms in which any two individuals of the appropriate sexes or mating types can produce fertile offspring, typically by sexual reproduction. It is the basic unit of Taxonomy (biology), ...
of
bird Birds are a group of warm-blooded vertebrates constituting the class (biology), class Aves (), characterised by feathers, toothless beaked jaws, the Oviparity, laying of Eggshell, hard-shelled eggs, a high Metabolism, metabolic rate, a fou ...
in the
family Family (from ) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). It forms the basis for social order. Ideally, families offer predictabili ...
Mimidae. It was
endemic Endemism is the state of a species being found only in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also foun ...
to Floreana, one of the
Galápagos Islands The Galápagos Islands () are an archipelago of volcanic islands in the Eastern Pacific, located around the equator, west of the mainland of South America. They form the Galápagos Province of the Republic of Ecuador, with a population of sli ...
, but now is found only on two nearby islets, Campeón and Gardner-near-Floreana.Cody, M. L. and C. J. Sharpe (2020). Floreana Mockingbird (''Mimus trifasciatus''), version 1.0. In Birds of the World (J. del Hoyo, A. Elliott, J. Sargatal, D. A. Christie, and E. de Juana, Editors). Cornell Lab of Ornithology, Ithaca, NY, USA. https://doi.org/10.2173/bow.chamoc1.01 retrieved July 24, 2021Remsen, J. V., Jr., J. I. Areta, E. Bonaccorso, S. Claramunt, A. Jaramillo, D. F. Lane, J. F. Pacheco, M. B. Robbins, F. G. Stiles, and K. J. Zimmer. Version 23 May 2021. A classification of the bird species of South America. American Ornithological Society. https://www.museum.lsu.edu/~Remsen/SACCBaseline.htm retrieved May 24, 2021 The Floreana mockingbird is also known as Darwin's mockingbird, as it was the arguable inspiration for
Charles Darwin Charles Robert Darwin ( ; 12 February 1809 – 19 April 1882) was an English Natural history#Before 1900, naturalist, geologist, and biologist, widely known for his contributions to evolutionary biology. His proposition that all speci ...
's work on the origins of species; he noticed distinct differences between them and previous species he had encountered and consequently established the existence of other variants on neighboring islands.


Taxonomy and systematics

The Floreana mockingbird, Galapagos mockingbird (''Mimus parvulus''), Espanola mockingbird (''M. macdonaldi''), and San Cristobal mockingbird (''M. melanotis'') were previously placed in genus ''Nesomimus'' and were considered
conspecific Biological specificity is the tendency of a characteristic such as a behavior or a biochemical variation to occur in a particular species. Biochemist Linus Pauling stated that "Biological specificity is the set of characteristics of living organism ...
. They now form a
superspecies In biology, a species complex is a group of closely related organisms that are so similar in appearance and other features that the boundaries between them are often unclear. The taxa in the complex may be able to hybridize readily with each oth ...
. The Floreana mockingbird is
monotypic In biology, a monotypic taxon is a taxonomic group (taxon) that contains only one immediately subordinate taxon. A monotypic species is one that does not include subspecies or smaller, infraspecific taxa. In the case of genera, the term "unisp ...
.


Description

The Floreana mockingbird is long. Males weigh an average of and females . Adults have a pied appearance. They have a pale
supercilium The supercilium is a plumage feature found on the heads of some bird species. It is a stripe which runs from the base of the bird's beak above its eye, finishing somewhere towards the rear of the bird's head.Dunn and Alderfer (2006), p. 10 Also k ...
, a black patch in front of the eye, a darkish one below it, and white cheeks. Their crown, upperparts, and tail are grayish brown with a few faint darker streaks. Their whitish underparts have a dark patch on the side of the breast and faint spots on the breast and flanks. The folded wing shows two white bars. The juvenile is similar to the adult but more heavily streaked.


Distribution and habitat

The Floreana mockingbird was formerly abundant on Floreana Island but had been
extirpate Local extinction, also extirpation, is the termination of a species (or other taxon) in a chosen geographic area of study, though it still exists elsewhere. Local extinctions are contrasted with global extinctions. Local extinctions mark a chan ...
d from there by 1888. It now occurs only on two islets off Floreana's shore, Campeón and Gardner-near-Floreana. The islets provide a habitat of low vegetation along their shores and arid scrub with cacti and a few trees inland.


Behavior


Feeding

The Floreana mockingbird mostly forages on the ground, but also in vegetation. It is primarily insectivorous but also eats cactus fruit, carrion, and probably seabird eggs.


Breeding

The Floreana mockingbird's breeding season spans from October to April and two broods are usual. It is a cooperative breeder with up to five adults in each territory of about . The nest is a cup made of twigs lined with finer material; it is almost always placed in a cactus. The clutch size is two to four.


Vocalization

The Floreana mockingbird has a " ud, melodious song nddoes not mimic other species."


Status

The
IUCN The International Union for Conservation of Nature (IUCN) is an international organization working in the field of nature conservation and sustainable use of natural resources. Founded in 1948, IUCN has become the global authority on the status ...
in 1994 initially assessed the Floreana mockingbird as Endangered. In 2008 it was reclassified as Critically Endangered but then in 2017 returned to Endangered status. It was extirpated from Floreana by 1888; introduced
rat Rats are various medium-sized, long-tailed rodents. Species of rats are found throughout the order Rodentia, but stereotypical rats are found in the genus ''Rattus''. Other rat genera include '' Neotoma'' (pack rats), '' Bandicota'' (bandicoo ...
s,
mice A mouse (: mice) is a small rodent. Characteristically, mice are known to have a pointed snout, small rounded ears, a body-length scaly tail, and a high breeding rate. The best known mouse species is the common house mouse (''Mus musculus' ...
,
cat The cat (''Felis catus''), also referred to as the domestic cat or house cat, is a small domesticated carnivorous mammal. It is the only domesticated species of the family Felidae. Advances in archaeology and genetics have shown that the ...
s,
dog The dog (''Canis familiaris'' or ''Canis lupus familiaris'') is a domesticated descendant of the gray wolf. Also called the domestic dog, it was selectively bred from a population of wolves during the Late Pleistocene by hunter-gatherers. ...
s, and
goat The goat or domestic goat (''Capra hircus'') is a species of Caprinae, goat-antelope that is mostly kept as livestock. It was domesticated from the wild goat (''C. aegagrus'') of Southwest Asia and Eastern Europe. The goat is a member of the ...
s have all been implicated as causes. The population on the two islets has fluctuated with the presence or absence of
El Niño EL, El or el may refer to: Arts and entertainment Fictional entities * El, a character from the manga series ''Shugo Chara!'' by Peach-Pit * Eleven (''Stranger Things'') (El), a fictional character in the TV series ''Stranger Things'' * El, fami ...
events but is believed to exceed 250 individuals and be stable.


Repopulation efforts

Studies are under way to attempt to reintroduce the Floreana mockingbird to the main island. These include assessments of Floreana's habitat and the status of the introduced
predator Predation is a biological interaction in which one organism, the predator, kills and eats another organism, its prey. It is one of a family of common List of feeding behaviours, feeding behaviours that includes parasitism and micropredation ...
s and
competitors Competition is a rivalry where two or more parties strive for a common goal which cannot be shared: where one's gain is the other's loss (an example of which is a zero-sum game). Competition can arise between entities such as organisms, indi ...
.


References

{{Taxonbar, from=Q3285195 Floreana mockingbird Endemic birds of the Galápagos Islands Floreana mockingbird Taxonomy articles created by Polbot Taxa named by John Gould