Floral Diagram
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A floral diagram is a graphic representation of the structure of a
flower Flowers, also known as blooms and blossoms, are the reproductive structures of flowering plants ( angiosperms). Typically, they are structured in four circular levels, called whorls, around the end of a stalk. These whorls include: calyx, m ...
. It shows the number of floral organs, their arrangement and fusion. Different parts of the flower are represented by their respective symbols. Floral diagrams are useful for flower identification or can help in understanding
angiosperm Flowering plants are plants that bear flowers and fruits, and form the clade Angiospermae (). The term angiosperm is derived from the Greek words (; 'container, vessel') and (; 'seed'), meaning that the seeds are enclosed within a fruit ...
evolution Evolution is the change in the heritable Phenotypic trait, characteristics of biological populations over successive generations. It occurs when evolutionary processes such as natural selection and genetic drift act on genetic variation, re ...
. They were introduced in the late 19th century and are generally attributed to A. W. Eichler. They are typically used with the
floral formula A floral formula is a notation for representing the structure of particular types of flowers. Such notations use numbers, letters and various symbols to convey significant information in a compact form. They may represent the floral form of a part ...
of that flower to study its
morphology Morphology, from the Greek and meaning "study of shape", may refer to: Disciplines *Morphology (archaeology), study of the shapes or forms of artifacts *Morphology (astronomy), study of the shape of astronomical objects such as nebulae, galaxies, ...
.


History

In the 19th century, two contrasting methods of describing the flower were introduced: the textual
floral formula A floral formula is a notation for representing the structure of particular types of flowers. Such notations use numbers, letters and various symbols to convey significant information in a compact form. They may represent the floral form of a part ...
e and pictorial floral diagrams. Floral diagrams are credited to A. W. Eichler, whose extensive work ''Blüthendiagramme'' (1875, 1878) remains a valuable source of information on floral morphology. Eichler inspired later generations of scientists, including John Henry Schaffner. Diagrams were included e.g. in ''Types of Floral Mechanism'' by Arthur Church (1908). They were used in different textbooks, e.g. ''Organogenesis of Flowers'''' ''by Rolf Sattler (1973), ''Botanische Bestimmungsübungen'' by Stützel (2006) or ''Plant Systematics'' by Simpson (2010). ''Floral Diagrams'' (2010) by Ronse De Craene followed Eichler’s approach using the contemporary APG II was contemporary at the time of book’s writing.
APG II system The APG II system (Angiosperm Phylogeny Group II system) of plant classification is the second, now obsolete, version of a modern, mostly Molecular phylogenetics, molecular-based, list of systems of plant taxonomy, system of plant taxonomy that ...
.


Basic characteristics and significance

A floral diagram is a schematic cross-section through a young flower. It may be also defined as “projection of the flower perpendicular to its axis”. It usually shows the number of floral parts,It also shows organs that are not part of the flower, but may be closely associated with it, such as bracts and bracteoles. their sizes, relative positions and fusion. Different organs are represented by distinguishable symbols, which may be uniform for one organ type, or may reflect concrete
morphology Morphology, from the Greek and meaning "study of shape", may refer to: Disciplines *Morphology (archaeology), study of the shapes or forms of artifacts *Morphology (astronomy), study of the shape of astronomical objects such as nebulae, galaxies, ...
. The diagram may also include symbols that don’t represent physical structures, but carry additional information (e.g.
symmetry Symmetry () in everyday life refers to a sense of harmonious and beautiful proportion and balance. In mathematics, the term has a more precise definition and is usually used to refer to an object that is Invariant (mathematics), invariant und ...
plane orientation). There is no agreement on how floral diagrams should be drawn, it depends on the author whether it is just a rough representation, or whether structural details of the flower are included. Diagrams can describe the
ontogeny Ontogeny (also ontogenesis) is the origination and development of an organism (both physical and psychological, e.g., moral development), usually from the time of fertilization of the ovum, egg to adult. The term can also be used to refer to t ...
of flowers, or can show evolutionary relationships. They can be generalized to show the typical floral structure of a
taxon In biology, a taxon (back-formation from ''taxonomy''; : taxa) is a group of one or more populations of an organism or organisms seen by taxonomists to form a unit. Although neither is required, a taxon is usually known by a particular name and ...
. It is also possible to represent (partial)
inflorescence In botany, an inflorescence is a group or cluster of flowers arranged on a plant's Plant stem, stem that is composed of a main branch or a system of branches. An inflorescence is categorized on the basis of the arrangement of flowers on a mai ...
s by diagrams. A substantial amount of information may be included in a good diagram. It can be useful for flower identification or comparison between
angiosperm Flowering plants are plants that bear flowers and fruits, and form the clade Angiospermae (). The term angiosperm is derived from the Greek words (; 'container, vessel') and (; 'seed'), meaning that the seeds are enclosed within a fruit ...
taxa. Paleontologists can take advantage of diagrams for reconstruction of
fossil A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserve ...
flowers. Floral diagrams are also of didactic value.


Orientation

Diagrams are usually depicted with the subtending
bract In botany, a bract is a modified or specialized leaf, associated with a reproductive structure such as a flower, inflorescence axis or cone scale. Bracts are usually different from foliage leaves in size, color, shape or texture. They also lo ...
''below'' and the axis ''above'' the flower itself, both in the median line. The axis corresponds to the position of the main
stem Stem or STEM most commonly refers to: * Plant stem, a structural axis of a vascular plant * Stem group * Science, technology, engineering, and mathematics Stem or STEM can also refer to: Language and writing * Word stem, part of a word respon ...
relative to a lateral flower. When a terminal flower is depicted, the axis is not present and therefore cannot be shown. Bracteoles, if they are present, are usually drawn on the sides of the diagram.


Symbols used in diagrams

Not only the information contained within diagrams, but also their appearance commonly varies between authors. Some publications include an overview of symbols used.


Bracts and bracteoles, axes

Bracts and bracteoles are commonly shown as arcs. In ''Floral Diagrams'' by Ronse De Craene they consistently have a black fill and a little triangle on the outer side to distinguish them from the perianth. In Eichler’s ''Blüthendiagramme'' their representation alters between diagrams. The axis relative to the flower is shown as black circle in ''Floral Diagrams''. When inflorescence is depicted, the position of its main stem is illustrated by a crossed circle. Eichler’s depiction of axes alternates between diagrams.


Perianth

Perianth parts are also shown as arcs. They may be colored according to their type. In ''Blüthendiagramme'' the
tepal A tepal is one of the outer parts of a flower (collectively the perianth). The term is used when these parts cannot easily be classified as either sepals or petals. This may be because the parts of the perianth are undifferentiated (i.e. of very ...
s are usually white with black stroke,
sepal A sepal () is a part of the flower of angiosperms (flowering plants). Usually green, sepals typically function as protection for the flower in bud, and often as support for the petals when in bloom., p. 106 Etymology The term ''sepalum'' ...
s are hatched and
petal Petals are modified leaves that form an inner whorl surrounding the reproductive parts of flowers. They are often brightly coloured or unusually shaped to attract pollinators. All of the petals of a flower are collectively known as the ''corol ...
s are black. Ronse De Craene implies that it may be sometimes impossible to classify the organs, he shows green perianth parts as black and pigmented as white. Estivation can be accurately shown in the diagram.


Androecium

Stamen The stamen (: stamina or stamens) is a part consisting of the male reproductive organs of a flower. Collectively, the stamens form the androecium., p. 10 Morphology and terminology A stamen typically consists of a stalk called the filament ...
s are represented by a cross-section through anthers. In case there are many stamens in the flower, they can be simplified and drawn as circles.
Staminode In botany, a staminode is an often rudimentary, sterile or abortive stamen, which means that it does not produce pollen.Jackson, Benjamin, Daydon; ''A Glossary of Botanic Terms with their Derivation and Accent''; Published by Gerald Duckworth & Co ...
s have a small black circle inside or are painted black in ''Floral Diagrams'', Eichler also fills them black.


Gynoecium

The
pistil Gynoecium (; ; : gynoecia) is most commonly used as a collective term for the parts of a flower that produce ovules and ultimately develop into the fruit and seeds. The gynoecium is the innermost whorl (botany), whorl of a flower; it consists ...
is shown as a sectional view of the
ovary The ovary () is a gonad in the female reproductive system that produces ova; when released, an ovum travels through the fallopian tube/ oviduct into the uterus. There is an ovary on the left and the right side of the body. The ovaries are end ...
. Ovary position is highlighted by small triangles in ''Floral Diagrams''. Ronse De Craene also incorporates ovule morphology or shows the position of stigmatic lobes by white shapes.


Nectaries

In ''Floral Diagrams'',
nectar Nectar is a viscous, sugar-rich liquid produced by Plant, plants in glands called nectaries, either within the flowers with which it attracts pollination, pollinating animals, or by extrafloral nectaries, which provide a nutrient source to an ...
ies are filled by grey color, Eichler fills them by hatching.


Other

Fusion can be shown in diagrams by full connecting lines between organs. Lost organs can be represented by a star (✶), lost perianth parts or bracts/bracteoles can be shown with dashed stroke. It is possible to show the direction of monosymmetry by a large arrow.
Resupination Resupination is derived from the Latin word ''resupinus'', meaning "bent back with the face upward" or "on the back". " Resupination" is the noun form of the adjective "resupine" which means "being upside-down, supine or facing upward". The word ...
may be illustrated by a curved arrow. Floral parts can be accompanied by numbers to show their sequence of initialization.


Floral diagrams and floral formulae

Each of these two concepts is better in expressing some information. Floral diagrams can show the size and relative position of the organs. On the other hand,
floral formula A floral formula is a notation for representing the structure of particular types of flowers. Such notations use numbers, letters and various symbols to convey significant information in a compact form. They may represent the floral form of a part ...
e are capable of broader generalization. Prenner ''et al.'' view them as complementary methods and state they make an “identikit” flower when utilized together. Ronse De Craene also approves of their combined use.


Examples


See also

*
Floral formula A floral formula is a notation for representing the structure of particular types of flowers. Such notations use numbers, letters and various symbols to convey significant information in a compact form. They may represent the floral form of a part ...
e *
Flower Flowers, also known as blooms and blossoms, are the reproductive structures of flowering plants ( angiosperms). Typically, they are structured in four circular levels, called whorls, around the end of a stalk. These whorls include: calyx, m ...
* Floral organs *
Dicot The dicotyledons, also known as dicots (or, more rarely, dicotyls), are one of the two groups into which all the flowering plants (angiosperms) were formerly divided. The name refers to one of the typical characteristics of the group: namely, ...
flower *
Monocot Monocotyledons (), commonly referred to as monocots, (Lilianae ''sensu'' Chase & Reveal) are flowering plants whose seeds contain only one Embryo#Plant embryos, embryonic leaf, or cotyledon. A monocot taxon has been in use for several decades, but ...
flower *
inflorescence In botany, an inflorescence is a group or cluster of flowers arranged on a plant's Plant stem, stem that is composed of a main branch or a system of branches. An inflorescence is categorized on the basis of the arrangement of flowers on a mai ...


Notes


References


Bibliography

*


External links


Floral diagram generator
{{botany Flowers Plant morphology