''Dichotomyctere ocellatus'' (
syn.
The Botanical and Zoological Codes of nomenclature treat the concept of synonymy differently.
* In botanical nomenclature, a synonym is a scientific name that applies to a taxon that (now) goes by a different scientific name. For example, Linna ...
''Tetraodon biocellatus''), commonly the figure 8 puffer or eyespot puffer, is a
pufferfish
Tetraodontidae is a family of primarily marine and estuarine fish of the order Tetraodontiformes. The family includes many familiar species variously called pufferfish, puffers, balloonfish, blowfish, blowies, bubblefish, globefish, swellfi ...
found in
freshwater
Fresh water or freshwater is any naturally occurring liquid or frozen water containing low concentrations of dissolved salts and other total dissolved solids. Although the term specifically excludes seawater and brackish water, it does in ...
in
Southeast Asia
Southeast Asia, also spelled South East Asia and South-East Asia, and also known as Southeastern Asia, South-eastern Asia or SEA, is the geographical United Nations geoscheme for Asia#South-eastern Asia, south-eastern region of Asia, consistin ...
. It is known from the lower reaches of the
Mekong
The Mekong or Mekong River is a trans-boundary river in East Asia and Southeast Asia. It is the world's twelfth longest river and the third longest in Asia. Its estimated length is , and it drains an area of , discharging of water annuall ...
(Cambodia), the
Peninsular Malaysia
Peninsular Malaysia ( ms, Semenanjung Malaysia; Jawi: سمننجڠ مليسيا), or the States of Malaya ( ms, Negeri-negeri Tanah Melayu; Jawi: نڬري-نڬري تانه ملايو), also known as West Malaysia or the Malaysian Peninsula, ...
as well as
Borneo
Borneo (; id, Kalimantan) is the List of islands by area, third-largest island in the world and the largest in Asia. At the geographic centre of Maritime Southeast Asia, in relation to major Indonesian islands, it is located north of Java Isl ...
(
Sarawak
Sarawak (; ) is a state of Malaysia. The largest among the 13 states, with an area almost equal to that of Peninsular Malaysia, Sarawak is located in northwest Borneo Island, and is bordered by the Malaysian state of Sabah to the northeast, ...
,
Kalimantan
Kalimantan () is the Indonesian portion of the island of Borneo. It constitutes 73% of the island's area. The non-Indonesian parts of Borneo are Brunei and East Malaysia. In Indonesia, "Kalimantan" refers to the whole island of Borneo.
In 2019, ...
).
Characteristics
Figure 8 puffers grow to about
total length
Fish measurement is the measuring of individual fish and various parts of their anatomies. These data are used in many areas of ichthyology, including taxonomy and fisheries biology.
Overall length
* Standard length (SL) is the length of a fish ...
(TL).
[ They are colourful fish, with greenish yellow patterns on their backs. These patterns vary greatly from fish to fish, but the markings either side of the caudal fin resemble the number eight, or eye-spots (earning the species another common name as "eyespot puffer"). Figure 8 puffers are relatively peaceful among ]Tetraodontidae
Tetraodontidae is a family of primarily marine and estuarine fish of the order Tetraodontiformes. The family includes many familiar species variously called pufferfish, puffers, balloonfish, blowfish, blowies, bubblefish, globefish, swellfi ...
, and have been kept successfully with other fish such as bumblebee gobies and mollies, but as with all pufferfish there is a risk that tankmates will not be tolerated.
Like all members of its family, the figure 8 puffer is capable of inflating itself with water or air when stressed or otherwise frightened.
In freshwater
Like the closely related green pufferfish, the figure 8 exhibits euryhalinity, and undergoes epithelial
Epithelium or epithelial tissue is one of the four basic types of animal tissue, along with connective tissue, muscle tissue and nervous tissue. It is a thin, continuous, protective layer of compactly packed cells with a little intercellu ...
remodeling and changes in the kidneys
The kidneys are two reddish-brown bean-shaped organs found in vertebrates. They are located on the left and right in the retroperitoneal space, and in adult humans are about in length. They receive blood from the paired renal arteries; bloo ...
and gills
A gill () is a respiratory organ that many aquatic organisms use to extract dissolved oxygen from water and to excrete carbon dioxide. The gills of some species, such as hermit crabs, have adapted to allow respiration on land provided they are ...
when exposed to changes in salinity. Accordingly, the figure 8 puffer is often mistakenly regarded as an entirely freshwater fish, and has considerable resistance to freshwater exposure. In a marine environment, it exhibits typical saltwater fish morphological features in its gills, allowing it to secrete sodium chloride
Sodium chloride , commonly known as salt (although sea salt also contains other chemical salts), is an ionic compound with the chemical formula NaCl, representing a 1:1 ratio of sodium and chloride ions. With molar masses of 22.99 and 35 ...
across the gill epithelium like a typical saltwater fish. However, acclimated to freshwater, the figure 8 puffer will exhibit specialized cells that do not match any gill ion
An ion () is an atom or molecule with a net electrical charge.
The charge of an electron is considered to be negative by convention and this charge is equal and opposite to the charge of a proton, which is considered to be positive by conven ...
-regulating cell of a freshwater fish. The other fish known to have these types of cells is the Mozambique tilapia
The Mozambique tilapia (''Oreochromis mossambicus'') is an oreochromine cichlid fish native to southeastern Africa. Dull colored, the Mozambique tilapia often lives up to a decade in its native habitats. It is a popular fish for aquaculture. Due ...
(''O. mossambicus''), but this is a saltwater fish.
In the aquarium
In captivity, figure 8 puffers require a 15 US gallon tank with temperatures between 24 °C and 28 °C. They are sensitive to nitrites
The nitrite ion has the chemical formula . Nitrite (mostly sodium nitrite) is widely used throughout chemical and pharmaceutical industries. The nitrite anion is a pervasive intermediate in the nitrogen cycle in nature. The name nitrite also r ...
and nitrates
Nitrate is a polyatomic ion with the chemical formula . Salts containing this ion are called nitrates. Nitrates are common components of fertilizers and explosives. Almost all inorganic nitrates are soluble in water. An example of an insol ...
and must be introduced into a fully cycled aquarium. Over-filtration is recommended as well as frequent partial water changes. Water pH will reflect the brackish environment to which these fish are best suited; the addition of marine salt
Marine is an adjective meaning of or pertaining to the sea or ocean.
Marine or marines may refer to:
Ocean
* Maritime (disambiguation)
* Marine art
* Marine biology
* Marine debris
* Marine habitats
* Marine life
* Marine pollution
Military
* ...
will buffer the pH to a range between 7.8 and 8.3. With a salinity
Salinity () is the saltiness or amount of salt dissolved in a body of water, called saline water (see also soil salinity). It is usually measured in g/L or g/kg (grams of salt per liter/kilogram of water; the latter is dimensionless and equal ...
of between 1.005 and 1.008 specific gravity
Relative density, or specific gravity, is the ratio of the density (mass of a unit volume) of a substance to the density of a given reference material. Specific gravity for liquids is nearly always measured with respect to water (molecule), wa ...
(S.G.), the lifespan of this species has been as long as 15 years.[The Puffer Forum, http://www.thepufferforum.com/forum/ug.php/v/PufferPedia/Brackish/T_Biocellatus/ (Retrieved 25 July 2007)] As with all puffer fish, they require a complex aquarium set up consisting of many nooks and crannies. This helps figure 8 puffers to behave as they would in nature, and reduce boredom.
Diet
Pufferfish are classed as molluscivore A molluscivore is a carnivorous animal that specialises in feeding on molluscs such as gastropods, bivalves, brachiopods and cephalopods. Known molluscivores include numerous predatory (and often cannibalistic) molluscs, (e.g.octopuses, murexes, de ...
s and feed mainly on benthic
The benthic zone is the ecological region at the lowest level of a body of water such as an ocean, lake, or stream, including the sediment surface and some sub-surface layers. The name comes from ancient Greek, βένθος (bénthos), meaning "t ...
organisms which may include mussels
Mussel () is the common name used for members of several families of bivalve molluscs, from saltwater and freshwater habitats. These groups have in common a shell whose outline is elongated and asymmetrical compared with other edible clams, which ...
, cockles, oysters
Oyster is the common name for a number of different families of salt-water bivalve molluscs that live in marine or brackish habitats. In some species, the valves are highly calcified, and many are somewhat irregular in shape. Many, but not al ...
and krill
Krill are small crustaceans of the order Euphausiacea, and are found in all the world's oceans. The name "krill" comes from the Norwegian word ', meaning "small fry of fish", which is also often attributed to species of fish.
Krill are consid ...
, though some puffers will take other foods, such as flake, it is not recommended that their primary food is flake. Their teeth, a beak formed from two plates, are capable of crushing shells in order to feed on prey. In captivity many fishkeepers feed snails as a substitute for the aforementioned foods as snails help keep the beak trim; it would otherwise be susceptible to overgrowth.
References
Bibliography
Ebert, K. (2001), ''The Puffers of Fresh and Brackish Water'', p 19. Aqualog, .
{{Taxonbar, from=Q28599195
Tetraodontidae
Taxa named by Franz Steindachner
Fish described in 1885
Fish of the Mekong Basin
Freshwater fish of Borneo
Fish of Cambodia
Freshwater fish of Indonesia
Freshwater fish of Malaysia