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''Ficus grossularioides'', the white-leaved fig, is a
species In biology, a species is the basic unit of Taxonomy (biology), classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of ...
of
flowering plant Flowering plants are plants that bear flowers and fruits, and form the clade Angiospermae (), commonly called angiosperms. They include all forbs (flowering plants without a woody stem), grasses and grass-like plants, a vast majority of ...
that belongs to
Moraceae The Moraceae — often called the mulberry family or fig family — are a family of flowering plants comprising about 38 genera and over 1100 species. Most are widespread in tropical and subtropical regions, less so in temperate climates; however ...
, the fig or mulberry
family Family (from la, familia) is a group of people related either by consanguinity (by recognized birth) or affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its members and of society. Idea ...
, it is native to
Southeast Asia Southeast Asia, also spelled South East Asia and South-East Asia, and also known as Southeastern Asia, South-eastern Asia or SEA, is the geographical United Nations geoscheme for Asia#South-eastern Asia, south-eastern region of Asia, consistin ...
.


Description

It is a small
dioecious Dioecy (; ; adj. dioecious , ) is a characteristic of a species, meaning that it has distinct individual organisms (unisexual) that produce male or female gametes, either directly (in animals) or indirectly (in seed plants). Dioecious reproducti ...
evergreen In botany, an evergreen is a plant which has foliage that remains green and functional through more than one growing season. This also pertains to plants that retain their foliage only in warm climates, and contrasts with deciduous plants, which ...
tree or shrub with a small trunk of only 15 cm in diameter. The bark of the trunk is thorn-less with a light brown reflection, with a smooth texture. It exudes a milky white latex exudate profusely. It can grow up to a height of 13 m (43 ft) in natural conditions and has glabrous twigs around 0.2-0.5 cm thick that are reddish brown. The leaves are elliptic to obovate in shape and are serrated from the mid margin to the tip, thin-coriaceous, glaucous, and velvety hairy on the underside, with very rough hairs on the topside. They are 19–11.5 cm long and 5.5–3.5 cm broad, with a 1.5–11.5 cm petiole. They are simple to trilobed and are spirally arranged on branches. The leaves have 4- 6 pairs of lateral veins, sunken on the top side, raised on the underside, basal pair reaching about half the length of the blade, with an oil gland in each axil on the underside. The
fruit In botany, a fruit is the seed-bearing structure in flowering plants that is formed from the ovary after flowering. Fruits are the means by which flowering plants (also known as angiosperms) disseminate their seeds. Edible fruits in partic ...
s are small ellipsoid to subglobose figs that grow along the twigs, 0.8–1 cm in diameter, light green initially, ripening to a yellowish brown, but sometimes to orange or red. The figs closely resemble
Gooseberries Gooseberry ( or (American and northern British) or (southern British)) is a common name for many species of ''Ribes'' (which also includes currants), as well as a large number of plants of similar appearance. The berries of those in the genu ...
.


Distribution

The species can be found in
Thailand Thailand ( ), historically known as Siam () and officially the Kingdom of Thailand, is a country in Southeast Asia, located at the centre of the Indochinese Peninsula, spanning , with a population of almost 70 million. The country is b ...
, Sumatra,
Peninsular Malaysia Peninsular Malaysia ( ms, Semenanjung Malaysia; Jawi: سمننجڠ مليسيا), or the States of Malaya ( ms, Negeri-negeri Tanah Melayu; Jawi: نڬري-نڬري تانه ملايو), also known as West Malaysia or the Malaysian Peninsula, ...
,
Singapore Singapore (), officially the Republic of Singapore, is a sovereign island country and city-state in maritime Southeast Asia. It lies about one degree of latitude () north of the equator, off the southern tip of the Malay Peninsula, borde ...
,
Borneo Borneo (; id, Kalimantan) is the List of islands by area, third-largest island in the world and the largest in Asia. At the geographic centre of Maritime Southeast Asia, in relation to major Indonesian islands, it is located north of Java Isl ...
, and
Java Java (; id, Jawa, ; jv, ꦗꦮ; su, ) is one of the Greater Sunda Islands in Indonesia. It is bordered by the Indian Ocean to the south and the Java Sea to the north. With a population of 151.6 million people, Java is the world's mo ...
. It is common throughout most of
Kalimantan Kalimantan () is the Indonesian portion of the island of Borneo. It constitutes 73% of the island's area. The non-Indonesian parts of Borneo are Brunei and East Malaysia. In Indonesia, "Kalimantan" refers to the whole island of Borneo. In 2019, ...
(In the mountains of Kalimantan it is replaced by the much rarer ''F. tricolor'') and
Sarawak Sarawak (; ) is a state of Malaysia. The largest among the 13 states, with an area almost equal to that of Peninsular Malaysia, Sarawak is located in northwest Borneo Island, and is bordered by the Malaysian state of Sabah to the northeast, ...
, growing in most secondary forests. In
Brunei Brunei ( , ), formally Brunei Darussalam ( ms, Negara Brunei Darussalam, Jawi alphabet, Jawi: , ), is a country located on the north coast of the island of Borneo in Southeast Asia. Apart from its South China Sea coast, it is completely sur ...
however, it is uncommon and cannot be found in
Sabah Sabah () is a state of Malaysia located in northern Borneo, in the region of East Malaysia. Sabah borders the Malaysian state of Sarawak to the southwest and the North Kalimantan province of Indonesia to the south. The Federal Territory ...
where its ecological niche is occupied by ''
Ficus septica ''Ficus septica'' (called Hauili in the Philippines, 稜果榕 in Taiwan) is a shrub or tree of the family Moraceae living at low altitudes from northeast India to north Australia (Queensland), and throughout Malesia. It lives on the edge of t ...
''. The non-overlapping ranges of these two common species in Borneo's secondary forest figs is quite possibly related to the basic
competitive exclusion principle In ecology Ecology () is the study of the relationships between living organisms, including humans, and their physical environment. Ecology considers organisms at the individual, population, community, ecosystem, and biosphere level. ...
, but this is currently unconfirmed as studies on the ecology of the two species are currently lacking.


Local names

* Called Kukan by the Bidayuh ethnic group * Called Kupan or Kumpan by the Iban ethnic group * also called Lenakan, Lengkan or Lelingkan by the Iban


Ecology


Habitat

It can be found in various habitats, from dry disturbed sandy soils,
kerangas forest The Sundaland heath forest, also known as ''Kerangas'' forest, is a type of tropical moist forest found on the island of Borneo, which is divided between Brunei, Indonesia, and Malaysia, as well as on the Indonesian islands of Belitung and Bangk ...
, swamps, paddy and oil palm plantations, riverbanks, sandy coastal forests, and up to 400 m in elevation. The species is commonly found on the edges of forests as it is usually among the first species to re-establish in a disturbed habitat.


Reproduction

''Ficus grossularioides'' is associated with a
fig wasp Fig wasps are wasps of the superfamily Chalcidoidea which spend their larval stage inside figs. Most are pollinators but others simply feed off the plant. The non-pollinators belong to several groups within the superfamily Chalcidoidea, while ...
, an agaonid wasp that acts as its sole pollinator as this wasp lays its
egg An egg is an organic vessel grown by an animal to carry a possibly fertilized egg cell (a zygote) and to incubate from it an embryo within the egg until the embryo has become an animal fetus that can survive on its own, at which point the a ...
s only on trees of this species. All fig trees have this unique form of
fertilization Fertilisation or fertilization (see spelling differences), also known as generative fertilisation, syngamy and impregnation, is the fusion of gametes to give rise to a new individual organism or offspring and initiate its development. Pro ...
, with each species relying on a single specialized species of pollinating wasp that relies on the fig tree to reproduce. The fruits are then eaten by birds, bats, and other mammals, later distributing the seeds via their fecal matter


Uses

The species is not usually cultivated as it is a commonly found plant and all products from it are collected from wild plants, traditional medicine uses latex against scorpion stings and leaf extract (
decoction Decoction is a method of extraction by boiling herbal or plant material (which may include stems, roots, bark and rhizomes) to dissolve the chemicals of the material. It is the most common preparation method in various herbal-medicine systems. De ...
) is used against kidney complaints. Young shoots are reported to be edible and can be eaten raw.


References

{{Taxonbar, from=Q15473447 grossularioides Flora of Southeast Asia