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''Ficus'' ( or ) is a
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial n ...
of about 850
species In biology, a species is the basic unit of Taxonomy (biology), classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of ...
of woody
tree In botany, a tree is a perennial plant with an elongated stem, or trunk, usually supporting branches and leaves. In some usages, the definition of a tree may be narrower, including only woody plants with secondary growth, plants that are ...
s, shrubs,
vine A vine ( Latin ''vīnea'' "grapevine", "vineyard", from ''vīnum'' "wine") is any plant with a growth habit of trailing or scandent (that is, climbing) stems, lianas or runners. The word ''vine'' can also refer to such stems or runners thems ...
s, epiphytes and
hemiepiphyte A hemiepiphyte is a plant that spends part of its life cycle as an epiphyte. The seeds of primary hemiepiphytes germinate in the canopy and initially live epiphytically. They send roots downward, and these roots eventually make contact with t ...
s in the
family Family (from la, familia) is a group of people related either by consanguinity (by recognized birth) or affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its members and of society. Idea ...
Moraceae The Moraceae — often called the mulberry family or fig family — are a family of flowering plants comprising about 38 genera and over 1100 species. Most are widespread in tropical and subtropical regions, less so in temperate climates; however ...
. Collectively known as fig trees or figs, they are native throughout the
tropics The tropics are the regions of Earth surrounding the Equator. They are defined in latitude by the Tropic of Cancer in the Northern Hemisphere at N and the Tropic of Capricorn in the Southern Hemisphere at S. The tropics are also referred ...
with a few species extending into the semi-warm
temperate In geography, the temperate climates of Earth occur in the middle latitudes (23.5° to 66.5° N/S of Equator), which span between the tropics and the polar regions of Earth. These zones generally have wider temperature ranges throughout t ...
zone. The
common fig The fig is the edible fruit of ''Ficus carica'', a species of small tree in the flowering plant family Moraceae. Native to the Mediterranean and western Asia, it has been cultivated since ancient times and is now widely grown throughout the wor ...
(''F. carica'') is a temperate species native to southwest Asia and the
Mediterranean The Mediterranean Sea is a sea connected to the Atlantic Ocean, surrounded by the Mediterranean Basin and almost completely enclosed by land: on the north by Western and Southern Europe and Anatolia, on the south by North Africa, and on th ...
region (from Afghanistan to Portugal), which has been widely cultivated from ancient times for its
fruit In botany, a fruit is the seed-bearing structure in flowering plants that is formed from the ovary after flowering. Fruits are the means by which flowering plants (also known as angiosperms) disseminate their seeds. Edible fruits in partic ...
, also referred to as figs. The fruit of most other species are also edible though they are usually of only local economic importance or eaten as
bushfood Bush tucker, also called bush food, is any food native to Australia and used as sustenance by Indigenous Australians, the Aboriginal and Torres Strait Islander peoples, but it can also describe any native flora or fauna used for culinary or m ...
. However, they are extremely important food resources for wildlife. Figs are also of considerable cultural importance throughout the tropics, both as objects of worship and for their many practical uses.


Description

''Ficus'' is a
pantropical A pantropical ("all tropics") distribution is one which covers tropical regions of both hemispheres. Examples of species include caecilians, modern sirenians and the plant genera ''Acacia'' and '' Bacopa''. '' Neotropical'' is a zoogeographic t ...
genus of trees, shrubs, and vines occupying a wide variety of
ecological niche In ecology, a niche is the match of a species to a specific environmental condition. Three variants of ecological niche are described by It describes how an organism or population responds to the distribution of resources and competitors (fo ...
s; most are
evergreen In botany, an evergreen is a plant which has foliage that remains green and functional through more than one growing season. This also pertains to plants that retain their foliage only in warm climates, and contrasts with deciduous plants, which ...
, but some deciduous species are found in areas outside of the tropics and to higher elevations. Fig species are characterized by their unique
inflorescence An inflorescence is a group or cluster of flowers arranged on a stem that is composed of a main branch or a complicated arrangement of branches. Morphologically, it is the modified part of the shoot of seed plants where flowers are formed ...
and distinctive
pollination syndrome Pollination syndromes are suites of flower traits that have evolved in response to natural selection imposed by different pollen vectors, which can be abiotic (wind and water) or biotic, such as birds, bees, flies, and so forth through a process c ...
, which uses wasp species belonging to the family
Agaonidae The family Agaonidae is a group of pollinating and nonpollinating fig wasps. They spend their larval stage inside the fruits of figs. The pollinating wasps ( Agaoninae, Kradibiinae, and Tetrapusiinae) are the mutualistic partners of the fig tr ...
for pollination. Specific identification of many of the species can be difficult, but members of the genus ''Ficus'' are relatively easy to recognize. Many have
aerial root Aerial roots are roots above the ground. They are almost always adventitious. They are found in diverse plant species, including epiphytes such as orchids (''Orchidaceae''), tropical coastal swamp trees such as mangroves, banyan figs ('' ...
s and a distinctive shape or habit, and their fruits distinguish them from other plants. The fruit of ''Ficus'' is an
inflorescence An inflorescence is a group or cluster of flowers arranged on a stem that is composed of a main branch or a complicated arrangement of branches. Morphologically, it is the modified part of the shoot of seed plants where flowers are formed ...
enclosed in an urn-like structure called a
syconium Syconium (plural ''syconia'') is the type of inflorescence borne by figs (genus ''Ficus''), formed by an enlarged, fleshy, hollow receptacle with multiple ovaries on the inside surface. In essence, it is really a fleshy stem with a number of flow ...
, which is lined on the inside with the fig's tiny flowers that develop into multiple ovaries on the inside surface. In essence, the fig fruit is a fleshy stem with multiple tiny flowers that fruit and coalesce. The unique fig
pollination Pollination is the transfer of pollen from an anther of a plant to the stigma of a plant, later enabling fertilisation and the production of seeds, most often by an animal or by wind. Pollinating agents can be animals such as insects, birds ...
system, involving tiny, highly specific wasps, known as
fig wasp Fig wasps are wasps of the superfamily Chalcidoidea which spend their larval stage inside figs. Most are pollinators but others simply feed off the plant. The non-pollinators belong to several groups within the superfamily Chalcidoidea, while ...
s that enter via
ostiole An ''ostiole'' is a small hole or opening through which algae or fungi release their mature spores. The word is a diminutive of "ostium", "opening". The term is also used in higher plants, for example to denote the opening of the involuted ...
these subclosed inflorescences to both pollinate and lay their own eggs, has been a constant source of inspiration and wonder to biologists. Notably, three vegetative traits together are unique to figs. All figs present a white to yellowish
latex Latex is an emulsion (stable dispersion) of polymer microparticles in water. Latexes are found in nature, but synthetic latexes are common as well. In nature, latex is found as a milky fluid found in 10% of all flowering plants (angiosper ...
, some in copious quantities; the twig shows paired
stipules In botany, a stipule is an outgrowth typically borne on both sides (sometimes on just one side) of the base of a leafstalk (the petiole). Stipules are considered part of the anatomy of the leaf of a typical flowering plant, although in many speci ...
—or circular scars if the stipules have fallen off; the lateral veins at the base of the leaf are steep, forming a tighter angle with the midrib than the other lateral veins, a feature referred to as "triveined". No unambiguous older
fossil A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserved ...
s of ''Ficus'' are known. However, current
molecular clock The molecular clock is a figurative term for a technique that uses the mutation rate of biomolecules to deduce the time in prehistory when two or more life forms diverged. The biomolecular data used for such calculations are usually nucleo ...
estimates indicate that ''Ficus'' is a relatively ancient genus being at least 60 million years old, and possibly as old as 80 million years. The main radiation of
extant Extant is the opposite of the word extinct. It may refer to: * Extant hereditary titles * Extant literature, surviving literature, such as ''Beowulf'', the oldest extant manuscript written in English * Extant taxon, a taxon which is not extinct, ...
species, however, may have taken place more recently, between 20 and 40 million years ago. Some better-known species that represent the diversity of the genus include the
common fig The fig is the edible fruit of ''Ficus carica'', a species of small tree in the flowering plant family Moraceae. Native to the Mediterranean and western Asia, it has been cultivated since ancient times and is now widely grown throughout the wor ...
, a small, temperate deciduous tree whose fingered fig leaf is well known in art and
iconography Iconography, as a branch of art history, studies the identification, description and interpretation of the content of images: the subjects depicted, the particular compositions and details used to do so, and other elements that are distinct fro ...
; the weeping fig (''F. benjamina''), a
hemiepiphyte A hemiepiphyte is a plant that spends part of its life cycle as an epiphyte. The seeds of primary hemiepiphytes germinate in the canopy and initially live epiphytically. They send roots downward, and these roots eventually make contact with t ...
with thin, tough leaves on pendulous stalks adapted to its
rain forest Rainforests are characterized by a closed and continuous tree canopy, moisture-dependent vegetation, the presence of epiphytes and lianas and the absence of wildfire. Rainforest can be classified as tropical rainforest or temperate rainforest ...
habitat; the rough-leaved sandpaper figs from Australia; and the creeping fig (''F. pumila''), a vine whose small, hard leaves form a dense carpet of foliage over rocks or garden walls. Moreover, figs with different plant habits have undergone
adaptive radiation In evolutionary biology, adaptive radiation is a process in which organisms diversify rapidly from an ancestral species into a multitude of new forms, particularly when a change in the environment makes new resources available, alters biotic int ...
in different
biogeographic Biogeography is the study of the distribution of species and ecosystems in geographic space and through geological time. Organisms and biological communities often vary in a regular fashion along geographic gradients of latitude, elevation, ...
regions, leading to very high levels of
alpha diversity In ecology, alpha diversity (α-diversity) is the mean species diversity in a site at a local scale. The term was introduced by R. H. WhittakerWhittaker, R. H. (1960) Vegetation of the Siskiyou Mountains, Oregon and California. Ecological Monograph ...
. In the tropics, ''Ficus'' commonly is the most species-rich plant genus in a particular forest. In Asia, as many as 70 or more species can co-exist. ''Ficus''
species richness Species richness is the number of different species represented in an ecological community, landscape or region. Species richness is simply a count of species, and it does not take into account the abundances of the species or their relative ab ...
declines with an increase in latitude in both hemispheres. A description of fig tree cultivation is set out in
Ibn al-'Awwam Ibn al-'Awwam ( ar, ابن العوام), also called Abu Zakariya Ibn al-Awwam ( ar, أبو زكريا بن العوام), was a Muslim Arab agriculturist who flourished at Seville (modern-day southern Spain) in the later 12th century. He wrote ...
's 12th-century agricultural work entitled, ''Book on Agriculture''.


Ecology

Figs are
keystone species A keystone species is a species which has a disproportionately large effect on its natural environment relative to its abundance, a concept introduced in 1969 by the zoologist Robert T. Paine. Keystone species play a critical role in maintaini ...
in many
tropical forest Tropical forests (a.k.a. jungle) are forested landscapes in tropical regions: ''i.e.'' land areas approximately bounded by the tropic of Cancer and Capricorn, but possibly affected by other factors such as prevailing winds. Some tropical fo ...
ecosystem An ecosystem (or ecological system) consists of all the organisms and the physical environment with which they interact. These biotic and abiotic components are linked together through nutrient cycles and energy flows. Energy enters the syst ...
s. Their fruit are a key resource for some frugivores including fruit bats, and primates including:
capuchin monkey The capuchin monkeys () are New World monkeys of the subfamily Cebinae. They are readily identified as the " organ grinder" monkey, and have been used in many movies and television shows. The range of capuchin monkeys includes some tropical fores ...
s,
langurs The Colobinae or leaf-eating monkeys are a subfamily of the Old World monkey family that includes 61 species in 11 genera, including the black-and-white colobus, the large-nosed proboscis monkey, and the gray langurs. Some classifications split ...
,
gibbons Gibbons may refer to: * The plural of gibbon, an ape in the family Hylobatidae * Gibbons (surname) * Gibbons, Alberta * Gibbons (automobile), a British light car of the 1920s * Gibbons P.C., a leading American law firm headquartered in New Jerse ...
and
mangabey Mangabeys are West-African Old World monkeys, with species in three of the six genera of tribe Papionini. The more typical representatives of '' Cercocebus'', also known as the white-eyelid mangabeys, are characterized by their bare, upper eye ...
s. They are even more important for
bird Birds are a group of warm-blooded vertebrates constituting the class Aves (), characterised by feathers, toothless beaked jaws, the laying of hard-shelled eggs, a high metabolic rate, a four-chambered heart, and a strong yet lightweig ...
s such as Asian barbets,
pigeons Columbidae () is a bird family consisting of doves and pigeons. It is the only family in the order Columbiformes. These are stout-bodied birds with short necks and short slender bills that in some species feature fleshy ceres. They primarily ...
,
hornbill Hornbills (Bucerotidae) are a family of bird found in tropical and subtropical Africa, Asia and Melanesia. They are characterized by a long, down-curved bill which is frequently brightly coloured and sometimes has a casque on the upper mandib ...
s, fig-parrots and
bulbul The bulbuls are members of a family, Pycnonotidae, of medium-sized passerine songbirds, which also includes greenbuls, brownbuls, leafloves, and bristlebills. The family is distributed across most of Africa and into the Middle East, tropical ...
s, which may almost entirely subsist on figs when these are in plenty. Many Lepidoptera
caterpillar Caterpillars ( ) are the larva, larval stage of members of the order Lepidoptera (the insect order comprising butterfly, butterflies and moths). As with most common names, the application of the word is arbitrary, since the larvae of sawfly ...
s feed on fig leaves, for example several '' Euploea'' species (crow butterflies), the plain tiger (''Danaus chrysippus''), the giant swallowtail (''Papilio cresphontes''), the
brown awl ''Badamia exclamationis'', commonly known as the brown awl or narrow-winged awl,Markku Savela's website on Lepidoptera. Page on genuTOL web page on genu''Badamia''/ref> is a butterfly belonging to the family Hesperiidae. It is found in south an ...
(''Badamia exclamationis''), and '' Chrysodeixis eriosoma'', Choreutidae and
Copromorphidae Copromorphidae, the "tropical fruitworm moths", is a family of insects in the lepidopteran order. These moths have broad, rounded forewings, and well-camouflaged scale patterns. Unlike Carposinidae the mouthparts include "labial palps" with the ...
moth Moths are a paraphyletic group of insects that includes all members of the order Lepidoptera that are not butterflies, with moths making up the vast majority of the order. There are thought to be approximately 160,000 species of moth, many of ...
s. The citrus long-horned beetle (''Anoplophora chinensis''), for example, has larvae that feed on
wood Wood is a porous and fibrous structural tissue found in the stems and roots of trees and other woody plants. It is an organic materiala natural composite of cellulose fibers that are strong in tension and embedded in a matrix of ligni ...
, including that of fig trees; it can become a
pest Pest or The Pest may refer to: Science and medicine * Pest (organism), an animal or plant deemed to be detrimental to humans or human concerns ** Weed, a plant considered undesirable * Infectious disease, an illness resulting from an infection ** ...
in fig plantations. Similarly, the sweet potato whitefly (''Bemisia tabaci'') is frequently found as a pest on figs grown as potted plants and is spread through the export of these plants to other localities. For a list of other diseases common to fig trees, see
List of foliage plant diseases (Moraceae) This is a list of diseases affecting species of the genus '' Ficus''. Plant Species Bacterial diseases Fungal diseases Nematodes, parasitic ReferencesCommon Names of Diseases, The American Phytopathological Society {{DEFAULTSORT:List ...
.


Fig fruit and reproduction system

Many fig species are grown for their fruits, though only ''
Ficus carica The fig is the edible fruit of ''Ficus carica'', a species of small tree in the flowering plant family Moraceae. Native to the Mediterranean and western Asia, it has been cultivated since ancient times and is now widely grown throughout the w ...
'' is cultivated to any extent for this purpose. A fig "fruit" is a type of multiple fruit known as a
syconium Syconium (plural ''syconia'') is the type of inflorescence borne by figs (genus ''Ficus''), formed by an enlarged, fleshy, hollow receptacle with multiple ovaries on the inside surface. In essence, it is really a fleshy stem with a number of flow ...
, derived from an arrangement of many small flowers on an inverted, nearly closed receptacle. The many small flowers are unseen unless the fig is cut open. The fruit typically has a bulbous shape with a small opening (the
ostiole An ''ostiole'' is a small hole or opening through which algae or fungi release their mature spores. The word is a diminutive of "ostium", "opening". The term is also used in higher plants, for example to denote the opening of the involuted ...
) at the outward end that allows access to pollinators. The flowers are pollinated by very small wasps that crawl through the opening in search of a suitable place to lay eggs. Without this pollinator service fig trees could not reproduce by seed. In turn, the flowers provide a safe haven and nourishment for the next generation of wasps. This accounts for the frequent presence of wasp larvae in the fruit, and has led to a
coevolution In biology, coevolution occurs when two or more species reciprocally affect each other's evolution through the process of natural selection. The term sometimes is used for two traits in the same species affecting each other's evolution, as well ...
ary relationship. Technically, a fig fruit proper would be only one of the many tiny matured, seed-bearing gynoecia found inside one fig – if you cut open a fresh fig, individual fruit will appear as fleshy "threads", each bearing a single seed inside. The genus ''
Dorstenia ''Dorstenia'' is a genus within the mulberry family, Moraceae. Depending on the author, there are said to be 100 to 170 species within this genus, second only in number to the genus ''Ficus'' within Moraceae. ''Dorstenia'' species are mainly kn ...
'', also in the fig family (Moraceae), exhibits similar tiny flowers arranged on a receptacle but in this case the receptacle is a more or less flat, open surface. Fig plants can be
monoecious Monoecy (; adj. monoecious ) is a sexual system in seed plants where separate male and female cones or flowers are present on the same plant. It is a monomorphic sexual system alongside gynomonoecy, andromonoecy and trimonoecy. Monoecy i ...
(
hermaphrodite In reproductive biology, a hermaphrodite () is an organism that has both kinds of reproductive organs and can produce both gametes associated with male and female sexes. Many taxonomic groups of animals (mostly invertebrates) do not have ...
) or gynodioecious (hermaphrodite and female). Nearly half of fig species are gynodioecious, and therefore have some plants with inflorescences (syconium) with long styled pistillate flowers, and other plants with staminate flowers mixed with short styled pistillate flowers. The long-styled flowers tend to prevent wasps from laying their eggs within the ovules, while the short-styled flowers are accessible for egg laying. All the native fig trees of the American continent are hermaphrodites, as well as species like Indian banyan (''F. benghalensis''), weeping fig (''F. benjamina''), Indian rubber plant (''F. elastica''), fiddle-leaved fig (''F. lyrata''),
Moreton Bay fig ''Ficus macrophylla'', commonly known as the Moreton Bay fig or Australian banyan, is a large evergreen banyan tree of the family Moraceae native to eastern Australia, from the Wide Bay–Burnett region in the north to the Illawarra in New So ...
(''F. macrophylla''),
Chinese banyan ''Ficus microcarpa'', also known as Chinese banyan, Malayan banyan, Indian laurel, curtain fig, or , is a tree in the fig family Moraceae. It is native in a range from China through tropical Asia and the Caroline Islands to Australia. It is wide ...
(''F. microcarpa''),
sacred fig ''Ficus religiosa'' or sacred fig is a species of fig native to the Indian subcontinent and Indochina that belongs to Moraceae, the fig or mulberry family. It is also known as the bodhi tree, pippala tree, peepul tree, peepal tree, pipal tree ...
(''F. religiosa'') and
sycamore fig ''Ficus sycomorus'', called the sycamore fig or the fig-mulberry (because the leaves resemble those of the mulberry), sycamore, or sycomore, is a fig species that has been cultivated since ancient times. The term ''sycamore'' spelled with an ...
(''F. sycomorus''). The common fig (''Ficus carica'') is a gynodioecious plant, as well as lofty fig or clown fig ('' F. aspera''), Roxburgh fig ('' F. auriculata''), mistletoe fig ('' F. deltoidea''), '' F. pseudopalma'', creeping fig (''F. pumila'') and related species. The hermaphrodite common figs are called "inedible figs" or "caprifigs"; in traditional culture in the
Mediterranean region In biogeography, the Mediterranean Basin (; also known as the Mediterranean Region or sometimes Mediterranea) is the region of lands around the Mediterranean Sea that have mostly a Mediterranean climate, with mild to cool, rainy winters and ...
they were considered food for
goat The goat or domestic goat (''Capra hircus'') is a domesticated species of goat-antelope typically kept as livestock. It was domesticated from the wild goat (''C. aegagrus'') of Southwest Asia and Eastern Europe. The goat is a member of ...
s (''Capra aegagrus''). In the female fig trees, the male flower parts fail to develop; they produce the "'edible figs".
Fig wasp Fig wasps are wasps of the superfamily Chalcidoidea which spend their larval stage inside figs. Most are pollinators but others simply feed off the plant. The non-pollinators belong to several groups within the superfamily Chalcidoidea, while ...
s grow in common fig caprifigs but not in the female syconiums because the female flower is too long for the wasp to successfully lay her eggs in them. Nonetheless, the wasp pollinates the flower with pollen from the caprifig it grew up in. When the wasp dies, it is broken down by enzymes ( Ficain) inside the fig. Fig wasps are not known to transmit any diseases harmful to humans. When a caprifig ripens, another caprifig must be ready to be pollinated. In temperate climes, wasps hibernate in figs, and there are distinct crops. Caprifigs have three crops per year; common figs have two. The first crop ( breba) is larger and juicier, and usually eaten fresh. In cold climates the breba crop is often destroyed by spring frosts. Some
parthenocarpic In botany and horticulture, parthenocarpy is the natural or artificially induced production of fruit without fertilisation of ovules, which makes the fruit seedless. Stenospermocarpy may also produce apparently seedless fruit, but the seeds are ac ...
cultivar A cultivar is a type of cultivated plant that people have selected for desired traits and when propagated retain those traits. Methods used to propagate cultivars include: division, root and stem cuttings, offsets, grafting, tissue culture ...
s of common figs do not require pollination at all, and will produce a crop of figs (albeit
sterile Sterile or sterility may refer to: *Asepsis, a state of being free from biological contaminants * Sterile (archaeology), a sediment deposit which contains no evidence of human activity *Sterilization (microbiology), any process that eliminates or ...
) in the absence of caprifigs and fig wasps. Depending on the species, each fruit can contain hundreds or even thousand of seeds. Figs can be propagated by seeds, cuttings, air-layering or grafting. However, as with any plant, figs grown from seed are not necessarily genetically identical to the parent and are only propagated this way for breeding purposes.


Mutualism with the pollinating fig wasps

Each species of fig is pollinated by one or a few specialised wasp species, and therefore plantings of fig species outside of their native range results in effectively sterile individuals. For example, in
Hawaii Hawaii ( ; haw, Hawaii or ) is a state in the Western United States, located in the Pacific Ocean about from the U.S. mainland. It is the only U.S. state outside North America, the only state that is an archipelago, and the only ...
, some 60 species of figs have been introduced, but only four of the wasps that fertilize them, so only those species of figs produce viable seeds there and can become
invasive species An invasive species otherwise known as an alien is an introduced organism that becomes overpopulated and harms its new environment. Although most introduced species are neutral or beneficial with respect to other species, invasive species adv ...
. This is an example of mutualism, in which each organism (fig plant and
fig wasp Fig wasps are wasps of the superfamily Chalcidoidea which spend their larval stage inside figs. Most are pollinators but others simply feed off the plant. The non-pollinators belong to several groups within the superfamily Chalcidoidea, while ...
) benefit each other, in this case reproductively. The intimate association between fig species and their wasp pollinators, along with the high incidence of a one-to-one plant-pollinator ratio have long led scientists to believe that figs and wasps are a clear example of
coevolution In biology, coevolution occurs when two or more species reciprocally affect each other's evolution through the process of natural selection. The term sometimes is used for two traits in the same species affecting each other's evolution, as well ...
. Morphological and reproductive behavior evidence, such as the correspondence between fig and wasp larvae maturation rates, have been cited as support for this hypothesis for many years. Additionally, recent genetic and molecular dating analyses have shown a very close correspondence in the character evolution and
speciation Speciation is the evolutionary process by which populations evolve to become distinct species. The biologist Orator F. Cook coined the term in 1906 for cladogenesis, the splitting of lineages, as opposed to anagenesis, phyletic evolution withi ...
phylogenies of these two clades. According to meta-analysis of molecular data for 119 fig species 35% (41) have multiple pollinator wasp species. The real proportion is higher because not all wasp species were detected. On the other hand, species of wasps pollinate multiple host fig species. Molecular techniques, like
microsatellite A microsatellite is a tract of repetitive DNA in which certain DNA motifs (ranging in length from one to six or more base pairs) are repeated, typically 5–50 times. Microsatellites occur at thousands of locations within an organism's genome. ...
markers and mitochondrial sequence analysis, allowed a discovery of multiple genetically distinct, cryptic wasp species. Not all these cryptic species are sister taxa and thus must have experienced a host fig shift at some point. These cryptic species lacked evidence of genetic
introgression Introgression, also known as introgressive hybridization, in genetics is the transfer of genetic material from one species into the gene pool of another by the repeated backcrossing of an interspecific hybrid with one of its parent species. Intro ...
or backcrosses indicating limited fitness for
hybrid Hybrid may refer to: Science * Hybrid (biology), an offspring resulting from cross-breeding ** Hybrid grape, grape varieties produced by cross-breeding two ''Vitis'' species ** Hybridity, the property of a hybrid plant which is a union of two diff ...
s and effective
reproductive isolation The mechanisms of reproductive isolation are a collection of evolutionary mechanisms, behaviors and physiological processes critical for speciation. They prevent members of different species from producing offspring, or ensure that any offsprin ...
and
speciation Speciation is the evolutionary process by which populations evolve to become distinct species. The biologist Orator F. Cook coined the term in 1906 for cladogenesis, the splitting of lineages, as opposed to anagenesis, phyletic evolution withi ...
. The existence of cryptic species suggests that neither the number of symbionts nor their evolutionary relationships are necessarily fixed ecologically. While the morphological characteristics that facilitate the fig-wasp mutualisms are likely to be shared more fully in closer relatives, the absence of unique pairings would make it impossible to do a one-to-one tree comparison and difficult to determine cospeciation.


Systematics

With 800 species, ''Ficus'' is by far the largest genus in the Moraceae, and is one of the largest genera of flowering plants currently described. The species currently classified within ''Ficus'' were originally split into several genera in the mid-1800s, providing the basis for a subgeneric classification when reunited into one genus in 1867. This classification put functionally
dioecious Dioecy (; ; adj. dioecious , ) is a characteristic of a species, meaning that it has distinct individual organisms (unisexual) that produce male or female gametes, either directly (in animals) or indirectly (in seed plants). Dioecious reproducti ...
species into four subgenera based on floral characters. In 1965, E. J. H. Corner reorganized the genus on the basis of breeding system, uniting these four dioecious subgenera into a single dioecious subgenus ''Ficus''.
Monoecious Monoecy (; adj. monoecious ) is a sexual system in seed plants where separate male and female cones or flowers are present on the same plant. It is a monomorphic sexual system alongside gynomonoecy, andromonoecy and trimonoecy. Monoecy i ...
figs were classified within the subgenera ''
Urostigma A banyan, also spelled "banian", is a fig that develops accessory trunks from adventitious prop roots, allowing the tree to spread outwards indefinitely. This distinguishes banyans from other trees with a strangler habit that begin life as a ...
'', '' Pharmacosycea'' and '' Sycomorus''. This traditional classification has been called into question by recent
phylogenetic In biology, phylogenetics (; from Greek φυλή/ φῦλον [] "tribe, clan, race", and wikt:γενετικός, γενετικός [] "origin, source, birth") is the study of the evolutionary history and relationships among or within groups o ...
studies employing genetic methods to investigate the relationships between representative members of the various sections of each subgenus. Of Corner's original subgeneric divisions of the genus, only ''Sycomorus'' is supported as monophyletic in the majority of phylogenetic studies. Notably, there is no clear split between dioecious and monoecious lineages. One of the two sections of ''Pharmacosycea'', a monoecious group, form a monophyletic clade basal to the rest of the genus, which includes the other section of ''Pharmacosycea'', the rest of the monoecious species, and all of the dioecious species. These remaining species are divided into two main monophyletic lineages (though the statistical support for these lineages isn't as strong as for the monophyly of the more derived clades within them). One consists of all sections of ''Urostigma'' except for section ''Urostigma s. s.''. The other includes section ''Urostigma s. s.'', subgenus ''Sycomorus'', and the species of subgenus ''Ficus'', though the relationships of the sections of these groups to one another are not well resolved.


Selected species

There are 875 accepted ''Ficus'' species, as of March 2021, according to
Plants of the World Online Plants of the World Online (POWO) is an online database published by the Royal Botanic Gardens, Kew. It was launched in March 2017 with the ultimate aim being "to enable users to access information on all the world's known seed-bearing plants ...
.


Subgenus ''Ficus''

*'' Ficus amplissima'' Sm. – bat fig *''
Ficus carica The fig is the edible fruit of ''Ficus carica'', a species of small tree in the flowering plant family Moraceae. Native to the Mediterranean and western Asia, it has been cultivated since ancient times and is now widely grown throughout the w ...
'' L. – common fig *'' Ficus daimingshanensis'' Chang *'' Ficus deltoidea'' Jack – mistletoe fig *'' Ficus erecta'' Thunb. – Japanese fig *'' Ficus fulva'' Reinw. ex Blume *'' Ficus grossularioides'' Burman f. – white-leaved fig *'' Ficus neriifolia'' Sm. *'' Ficus palmata'' Forssk. *'' Ficus pandurata'' Hance *'' Ficus simplicissima'' Lour. (synonym ''
Ficus hirta ''Ficus simplicissima'' is an Asian species of fig tree in the family Moraceae. This species is similar to ''Ficus triloba ''Ficus triloba''Voigt JO (1845) In: ''Hort. Suburb. Calcutt.'': 284. is an Asian species of fig tree in the family Mo ...
'' Vahl) *''
Ficus triloba ''Ficus triloba''Voigt JO (1845) In: ''Hort. Suburb. Calcutt.'': 284. is an Asian species of fig tree in the family Moraceae. It is dioecious Dioecy (; ; adj. dioecious , ) is a characteristic of a species, meaning that it has distinct indivi ...
'' Buch.-Ham. ex Voigt


Subgenus ''Pharmacosycea''

*''
Ficus crassiuscula ''Ficus crassiuscula'' is a species of flowering plant in the family Moraceae, native to Central America (Guatemala, Honduras, Belize, Nicaragua, Costa Rica, Panama) and north-western parts of South America (Colombia, Ecuador, Peru and Venezuela) ...
'' Standl. *'' Ficus gigantosyce'' Dugand *''
Ficus insipida ''Ficus insipida'' is a common tropical tree in the fig genus of the family Moraceae growing in forest habitats along rivers. It ranges from Mexico to northern South America. Taxonomy The tree was described in 1806 under the scientific na ...
'' Willd. *'' Ficus lacunata'' Kvitvik *'' Ficus maxima'' Mill. *'' Ficus mutabilis'' Bureau *'' Ficus nervosa'' Heyne ex Roth *'' Ficus pulchella'' Schott *'' Ficus yoponensis'' Desv.


Subgenus ''Sycidium''

*'' Ficus andamanica'' Corner *'' Ficus aspera'' G.Forst. *'' Ficus assamica'' Miq. *'' Ficus bojeri'' Baker *'' Ficus capreifolia'' Delile *'' Ficus coronata'' Spin – creek sandpaper fig *'' Ficus fraseri'' Miq. – shiny sandpaper fig *'' Ficus heterophylla'' L.f. *'' Ficus lateriflora'' Vahl *'' Ficus montana'' Burm.f. – oakleaf fig *'' Ficus opposita'' Miq. – sweet sandpaper fig *'' Ficus phaeosyce'' K.Schum. & Lauterb. *''
Ficus tinctoria ''Ficus tinctoria'', also known as dye fig, or humped fig is a hemiepiphytic tree of genus ''Ficus''. It is also one of the species known as ''strangler fig''. It is found in Asia, Malesia, northern Australia, and the South Pacific islands. It ...
'' G.Forst. – dye fig *'' Ficus ulmifolia'' Lam. *''
Ficus wassa ''Ficus wassa'' is a species of fig in the family Moraceae found in Malesia Malesia is a biogeographical region straddling the Equator and the boundaries of the Indomalayan and Australasian realms, and also a phytogeographical floristic ...
'' Roxb. *'' Ficus parietalis'' *'' Ficus sinuata'' *'' Ficus hampelas''


Subgenus ''Sycomorus''

*'' Ficus auriculata'' Lour. – Roxburgh fig *'' Ficus bernaysii'' King *'' Ficus dammaropsis'' Diels – highland breadfruit, ''kapiak'' *'' Ficus fistulosa'' Blume *'' Ficus hispida'' L. *'' Ficus nota'' Merr. – ''tibig'' *'' Ficus pseudopalma'' Blanco *''
Ficus racemosa ''Ficus racemosa'', the cluster fig, red river fig or gular, is a species of plant in the family Moraceae. It is native to Australia and tropical Asia. It is a fast-growing plant with large, very rough leaves, usually attaining the size of a lar ...
'' L. – cluster fig *'' Ficus septica'' Burm.f. – hauli tree *''
Ficus sycomorus ''Ficus sycomorus'', called the sycamore fig or the fig-mulberry (because the leaves resemble those of the mulberry), sycamore, or sycomore, is a fig species that has been cultivated since ancient times. The term ''sycamore'' spelled with an ...
'' L., 1753 – sycamore fig (Africa) *'' Ficus variegata'' Blume


Subgenus ''Synoecia''

The following species are typically spreading or climbing
liana A liana is a long- stemmed, woody vine that is rooted in the soil at ground level and uses trees, as well as other means of vertical support, to climb up to the canopy in search of direct sunlight. The word ''liana'' does not refer to a ...
s: *'' Ficus hederacea'' Roxb. *'' Ficus pantoniana'' King – climbing fig *'' Ficus pumila'' L. – creeping fig ** ''Ficus pumila'' var. ''awkeotsang'' (Makino) Corner – jelly fig *'' Ficus punctata'' Thunb. *'' Ficus sagittata'' J. König ex Vahl *'' Ficus sarmentosa'' Buch.-Ham. ex Sm. *'' Ficus trichocarpa'' Blume *''
Ficus villosa ''Ficus villosa'', known as the shaggy-leaf fig or villous fig, is a species of ''Ficus'' native to South East Asia. Etymology The species epithet "villosa" is derived from Latin ''villōsa'' "hairy", from ''villus'' "hair" and the adjective ...
'' Blume


Subgenus ''

Urostigma A banyan, also spelled "banian", is a fig that develops accessory trunks from adventitious prop roots, allowing the tree to spread outwards indefinitely. This distinguishes banyans from other trees with a strangler habit that begin life as a ...
''

*'' Ficus abutilifolia'' Miq. *'' Ficus albert-smithii'' Standl. *'' Ficus altissima'' Blume *'' Ficus amazonica'' Miq. *'' Ficus americana'' Aubl. *'' Ficus aripuanensis'' Berg & Kooy *'' Ficus arpazusa'' Carauta and Diaz – Brazil *''
Ficus aurea ''Ficus aurea'', commonly known as the Florida strangler fig (or simply strangler fig), golden fig, or ''higuerón'', is a tree in the family Moraceae that is native to the U.S. state of Florida, the northern and western Caribbean, southern Me ...
'' Nutt. – Florida strangler fig *''
Ficus beddomei ''Ficus'' ( or ) is a genus of about 850 species of woody trees, shrubs, vines, epiphytes and hemiepiphytes in the family Moraceae. Collectively known as fig trees or figs, they are native throughout the tropics with a few species exten ...
'' King – ''thavital'' *''
Ficus benghalensis ''Ficus benghalensis'', commonly known as the banyan, banyan fig and Indian banyan, is a tree native to the Indian Subcontinent. Specimens in India are among the largest trees in the world by canopy coverage. It also known as the "strangler fig" ...
'' L. – Indian banyan *''
Ficus benjamina ''Ficus benjamina'', commonly known as weeping fig, benjamin fig or ficus tree, and often sold in stores as just ficus, is a species of flowering plant in the family Moraceae, native to Asia and Australia. It is the official tree of Bangkok. The ...
'' L. – weeping fig *'' Ficus binnendijkii'' Miq. *'' Ficus bizanae'' Hutch. & Burtt-Davy *'' Ficus blepharophylla'' Vázquez Avila *'' Ficus broadwayi'' Urb. *'' Ficus burtt-davyi'' Hutch. *'' Ficus calyptroceras'' Miq. *'' Ficus castellviana'' Dugand *'' Ficus catappifolia'' Kunth & Bouché *'' Ficus citrifolia'' Mill. – short-leaved fig *'' Ficus consociata'' Bl. *'' Ficus cordata'' Thunb. *'' Ficus costata'' Ait. *'' Ficus crassipes'' F.M.Bailey – round-leaved banana fig *'' Ficus craterostoma'' Mildbr. & Burret *'' Ficus cyathistipula'' Warb. *'' Ficus cyclophylla'' (Miq.) Miq. *'' Ficus dendrocida'' Kunth *'' Ficus depressa'' Bl. *'' Ficus destruens'' F.White *'' Ficus drupacea'' Thunb. *''
Ficus elastica ''Ficus elastica'', the rubber fig, rubber bush, rubber tree, rubber plant, or Indian rubber bush, Indian rubber tree, is a species of flowering plant in the family Moraceae, native to eastern parts of South and Southeast Asia. It has become n ...
'' Hornem. – rubber plant *'' Ficus exasperata'' Vahl. *'' Ficus faulkneriana'' Berg *'' Ficus fergusonii'' (King) T.B.Worth. ex Corner *'' Ficus glaberrima'' Blume *'' Ficus glumosa'' Delile *'' Ficus greiffiana'' Dugand *'' Ficus hirsuta'' Schott *'' Ficus ilicina'' Miq. *'' Ficus kerkhovenii'' Valeton – Johore fig *'' Ficus kurzii'' King *'' Ficus luschnathiana'' Miq. *''
Ficus ingens ''Ficus ingens'', the red-leaved fig, is a fig species with an extensive range in the subtropical to dry tropical regions of Africa and southern Arabia. Despite its specific name, which means "huge", or "vast", it is usually a shrub or tree of mo ...
'' Miq. *'' Ficus krukovii'' Standl. *''
Ficus lacor ''Ficus lacor'' is a large evergreen tree of the family Moraceae. It is the city tree of Chongqing Chongqing ( or ; ; Sichuanese pronunciation: , Standard Mandarin pronunciation: ), alternately romanized as Chungking (), is a municip ...
'' Buch.-Ham. *'' Ficus lapathifolia'' Miq. *'' Ficus lauretana'' Vázquez Avila *'' Ficus lutea'' Vahl *''
Ficus lyrata ''Ficus lyrata'', commonly known as the fiddle-leaf fig, is a species of flowering plant in the mulberry and fig family Moraceae. It is native to western Africa, from Cameroon west to Sierra Leone, where it grows in lowland tropical rainforest ...
'' Warb. – fiddle-leaved fig *'' Ficus maclellandii'' King – Alii fig *''
Ficus macrophylla ''Ficus macrophylla'', commonly known as the Moreton Bay fig or Australian banyan, is a large evergreen banyan tree of the family Moraceae native to eastern Australia, from the Wide Bay–Burnett region in the north to the Illawarra in New ...
'' Desf. ex Pers. – Moreton Bay fig *'' Ficus malacocarpa'' Standl. *'' Ficus mariae'' Berg, Emygdio & Carauta *'' Ficus mathewsii'' Miq. *'' Ficus matiziana'' Dugand *'' Ficus mexiae'' Standl. *''
Ficus microcarpa ''Ficus microcarpa'', also known as Chinese banyan, Malayan banyan, Indian laurel, curtain fig, or , is a tree in the fig family Moraceae. It is native in a range from China through tropical Asia and the Caroline Islands to Australia. It is wide ...
'' L. – Chinese banyan *'' Ficus muelleriana'' Berg *''
Ficus natalensis ''Ficus natalensis'' is a tree in the family Moraceae. It is commonly known as the natal fig in South Africa. In central and western Uganda, where it has an important cultural value, it is known as ''omutuba'' to the Baganda people and ''omutoma' ...
'' Hochst. – Natal fig *'' Ficus obliqua'' G.Forst. – small-leaved fig *'' Ficus obtusifolia'' Kunth *'' Ficus pakkensis'' Standl. *'' Ficus pallida'' Vahl *'' Ficus panurensis'' Standl. *'' Ficus pertusa'' L.f. *'' Ficus petiolaris'' Kunth *'' Ficus pisocarpa'' Bl. *''
Ficus platypoda ''Ficus platypoda'', commonly known as the desert fig or rock fig, is a fig that is endemic to central and northern Australia. It is a lithophytic plant that grows on rocky outcrops, reaching 10 m in height. Taxonomy Dutch botanist Friedrich A ...
'' Cunn. – desert fig *'' Ficus pleurocarpa'' DC. – banana fig *'' Ficus polita'' Vahl *''
Ficus religiosa ''Ficus religiosa'' or sacred fig is a species of fig native to the Indian subcontinent and Indochina that belongs to Moraceae, the fig or mulberry family. It is also known as the bodhi tree, pippala tree, peepul tree, peepal tree, pipal tree ...
'' L. – sacred fig *'' Ficus roraimensis'' Berg *'' Ficus rubiginosa'' Desf. – Port Jackson fig *''
Ficus rumphii ''Ficus rumphii''von Blume CL (1825) In: ''Bijdragen tot de Flora van Nederlandsch Indie'' 437. is a banyan fig species in the family Moraceae. No subspecies are listed in the Catalogue of Life. The species can be found in: India, southern Chin ...
'' Blume *'' Ficus salicifolia'' Vahl – willow-leaved fig *'' Ficus sansibarica'' Warb. *'' Ficus schippii'' Standl. *'' Ficus schultesii'' Dugand *'' Ficus schumacheri'' Griseb. *'' Ficus sphenophylla'' Standl. *'' Ficus stuhlmannii'' Warb. *''
Ficus subcordata ''Ficus subcordata''von Blume CL (1825) In: ''Bijdragen tot de Flora van Nederlandsch Indie'' 440. is a banyan fig species in the family Moraceae. No subspecies are listed in the Catalogue of Life. The species can be found in Indo-china, Ma ...
'' Bl. *'' Ficus subpisocarpa'' Gagnep. *'' Ficus subpuberula'' Corner *'' Ficus sumatrana'' Miq. *'' Ficus superba'' Miq. ** ''Ficus superba'' var. ''henneana'' (Miq.) Corner *'' Ficus thonningii'' Blume *'' Ficus trichopoda'' Baker *'' Ficus trigona'' L.f. *'' Ficus trigonata'' L. *'' Ficus triradiata'' Corner – red-stipule fig *'' Ficus ursina'' Standl. *'' Ficus velutina'' Willd. *'' Ficus verruculosa'' Warb. *''
Ficus virens ''Ficus virens'' is a plant of the genus '' Ficus'' found in Pakistan, India, southeast Asia, through Malaysia and into Northern Australia. Its common name is white fig; it is locally known as ''pilkhan'' and in the Kunwinjku language it is cal ...
'' Aiton – white fig ** ''Ficus virens'' var. ''sublanceolata'' (Miq.) Corner – sour fig *''
Ficus watkinsiana ''Ficus watkinsiana'', commonly known as strangler fig, Watkins' fig, nipple fig or the green-leaved Moreton Bay fig is a hemiepiphytic fig that is endemic to Australia. The species exists in three populations—one in northeast Queensland an ...
'' F.M.Bailey – Watkins's fig


Unknown subgenus

*'' Ficus bibracteata'' *'' Ficus callosa'' Willd. *'' Ficus cristobalensis'' *'' Ficus hebetifolia'' *'' Ficus tsjahela'' Burm.f. * '' Ficus nymphaeifolia'' Mill.


Uses

The wood of fig trees is often soft and the
latex Latex is an emulsion (stable dispersion) of polymer microparticles in water. Latexes are found in nature, but synthetic latexes are common as well. In nature, latex is found as a milky fluid found in 10% of all flowering plants (angiosper ...
precludes its use for many purposes. It was used to make
mummy A mummy is a dead human or an animal whose soft tissues and organs have been preserved by either intentional or accidental exposure to chemicals, extreme cold, very low humidity, or lack of air, so that the recovered body does not decay fu ...
casket A casket jewelry box is a container that is usually smaller than a chest, and in the past were typically decorated. Whereas cremation jewelry is a small container, usually in the shape of a pendant or bracelet, to hold a small amount of ashes. ...
s in Ancient Egypt. Certain fig species (mainly ''F. cotinifolia'', '' F. insipida'' and '' F. padifolia'') are traditionally used in
Mesoamerica Mesoamerica is a historical region and cultural area in southern North America and most of Central America. It extends from approximately central Mexico through Belize, Guatemala, El Salvador, Honduras, Nicaragua, and northern Costa Rica. W ...
to produce '' papel amate'' (
Nahuatl Nahuatl (; ), Aztec, or Mexicano is a language or, by some definitions, a group of languages of the Uto-Aztecan language family. Varieties of Nahuatl are spoken by about Nahua peoples, most of whom live mainly in Central Mexico and have small ...
: ''āmatl''). ''Mutuba'' ('' F. natalensis'') is used to produce
barkcloth Barkcloth or bark cloth is a versatile material that was once common in Asia, Africa, and the Pacific. Barkcloth comes primarily from trees of the family Moraceae, including '' Broussonetia papyrifera'', ''Artocarpus altilis'', '' Artocarpus ...
in Uganda. ''Pou'' ('' F. religiosa'') leaves' shape inspired one of the standard '' kbach rachana'', decorative elements in Cambodian architecture. Indian banyan (''F. benghalensis'') and the Indian rubber plant, as well as other species, have use in
herbalism Herbal medicine (also herbalism) is the study of pharmacognosy and the use of medicinal plants, which are a basis of traditional medicine. With worldwide research into pharmacology, some herbal medicines have been translated into modern remedie ...
. The inner bark of an unknown type of wild fig, locally known as ''urú'', was once used by the of Bolivia to produce a fibrous cloth used for clothing. Figs have figured prominently in some human cultures. There is evidence that figs, specifically the
common fig The fig is the edible fruit of ''Ficus carica'', a species of small tree in the flowering plant family Moraceae. Native to the Mediterranean and western Asia, it has been cultivated since ancient times and is now widely grown throughout the wor ...
(''F. carica'') and sycamore fig (''
Ficus sycomorus ''Ficus sycomorus'', called the sycamore fig or the fig-mulberry (because the leaves resemble those of the mulberry), sycamore, or sycomore, is a fig species that has been cultivated since ancient times. The term ''sycamore'' spelled with an ...
''), were among the first – if not the very first – plant species that were deliberately bred for agriculture in the Middle East, starting more than 11,000 years ago. Nine
subfossil A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserved ...
''F. carica'' figs dated to about 9400–9200 BCE were found in the early
Neolithic The Neolithic period, or New Stone Age, is an Old World archaeological period and the final division of the Stone Age. It saw the Neolithic Revolution, a wide-ranging set of developments that appear to have arisen independently in several part ...
village Gilgal I (in the
Jordan Valley The Jordan Valley ( ar, غور الأردن, ''Ghor al-Urdun''; he, עֵמֶק הַיַרְדֵּן, ''Emek HaYarden'') forms part of the larger Jordan Rift Valley. Unlike most other river valleys, the term "Jordan Valley" often applies just to ...
, 13 km, or 8.1 mi, north of Jericho). These were a
parthenogenetic Parthenogenesis (; from the Greek grc, παρθένος, translit=parthénos, lit=virgin, label=none + grc, γένεσις, translit=génesis, lit=creation, label=none) is a natural form of asexual reproduction in which growth and developmen ...
type and thus apparently an early cultivar. This find predates the first known cultivation of
grain A grain is a small, hard, dry fruit ( caryopsis) – with or without an attached hull layer – harvested for human or animal consumption. A grain crop is a grain-producing plant. The two main types of commercial grain crops are cereals and legu ...
in the Middle East by many hundreds of years. Fig is a popular species in the practice of bonsai. It is particularly popular with beginners, as it is widely available and survives well as an indoor plant.


Cultivation

Numerous species of fig are found in cultivation in domestic and office environments, including: *'' F. carica'', common fig – hardy to . Shrub or small tree which can be grown outdoors in mild temperate regions, producing substantial harvests of fruit. Many cultivars are available. *'' F. benjamina'', weeping fig, ficus – hardy to . Widely used as an indoor plant for the home or the office. It benefits from the dry, warm atmosphere of centrally-heated interiors, and can grow to substantial heights in a favoured position. Several variegated cultivars are available. *'' F. elastica'', rubber plant – hardy to : widely cultivated as a houseplant; several cultivars with variegated leaves *'' F. lyrata'', fiddle-leaf fig – hardy to *'' F. maclellandii'' – hardy to *'' F. microcarpa'', Indian laurel – hardy to *'' F. pumila'', creeping fig – hardy to *'' F. rubiginosa'', Port Jackson fig – hardy to


Cultural and spiritual significance

Fig trees have profoundly influenced culture through several religious traditions. Among the more famous species are the
sacred fig ''Ficus religiosa'' or sacred fig is a species of fig native to the Indian subcontinent and Indochina that belongs to Moraceae, the fig or mulberry family. It is also known as the bodhi tree, pippala tree, peepul tree, peepal tree, pipal tree ...
tree (Pipal, bodhi, bo, or po, ''Ficus religiosa'') and other
banyan A banyan, also spelled "banian", is a fig that develops accessory trunks from adventitious prop roots, allowing the tree to spread outwards indefinitely. This distinguishes banyans from other trees with a strangler habit that begin life as a ...
figs such as ''Ficus benghalensis''. The oldest living plant of known planting date is a ''Ficus religiosa'' tree known as the Sri Maha Bodhi planted in the temple at
Anuradhapura Anuradhapura ( si, අනුරාධපුරය, translit=Anurādhapuraya; ta, அனுராதபுரம், translit=Aṉurātapuram) is a major city located in north central plain of Sri Lanka. It is the capital city of North Central P ...
, Sri Lanka by King Tissa in 288 BCE. The common fig is one of two significant trees in Islam, and there is a
sura A ''surah'' (; ar, سورة, sūrah, , ), is the equivalent of "chapter" in the Qur'an. There are 114 ''surahs'' in the Quran, each divided into '' ayats'' (verses). The chapters or ''surahs'' are of unequal length; the shortest surah ('' Al-K ...
in Quran named "The Fig" or At-Tin (سوره تین). In Asia, figs are important in
Buddhism Buddhism ( , ), also known as Buddha Dharma and Dharmavinaya (), is an Indian religion or philosophical tradition based on teachings attributed to the Buddha. It originated in northern India as a -movement in the 5th century BCE, and ...
and
Hinduism Hinduism () is an Indian religion or ''dharma'', a religious and universal order or way of life by which followers abide. As a religion, it is the world's third-largest, with over 1.2–1.35 billion followers, or 15–16% of the global po ...
. In
Jainism Jainism ( ), also known as Jain Dharma, is an Indian religion. Jainism traces its spiritual ideas and history through the succession of twenty-four tirthankaras (supreme preachers of ''Dharma''), with the first in the current time cycle being ...
, the consumption of any fruit belonging to this genus is prohibited. The
Buddha Siddhartha Gautama, most commonly referred to as the Buddha, was a wandering ascetic and religious teacher who lived in South Asia during the 6th or 5th century BCE and founded Buddhism. According to Buddhist tradition, he was born in ...
is traditionally held to have found ''
bodhi The English term enlightenment is the Western translation of various Buddhist terms, most notably bodhi and vimutti. The abstract noun ''bodhi'' (; Sanskrit: बोधि; Pali: ''bodhi''), means the knowledge or wisdom, or awakened intellect, ...
'' (enlightenment) while meditating for 49 days under a
sacred fig ''Ficus religiosa'' or sacred fig is a species of fig native to the Indian subcontinent and Indochina that belongs to Moraceae, the fig or mulberry family. It is also known as the bodhi tree, pippala tree, peepul tree, peepal tree, pipal tree ...
. The same species was '' Ashvattha'', the "
world tree The world tree is a motif present in several religions and mythologies, particularly Indo-European religions, Siberian religions, and Native American religions. The world tree is represented as a colossal tree which supports the heavens, thereb ...
" of Hinduism. The '' Plaksa Pra-sravana'' was said to be a fig tree between the roots of which the
Sarasvati River The Sarasvati River () is a deified river first mentioned in the Rigveda and later in Vedic and post-Vedic texts. It played an important role in the Vedic religion, appearing in all but the fourth book of the Rigveda. As a physical river, i ...
sprang forth; it is usually held to be a sacred fig but more probably is ''
Ficus virens ''Ficus virens'' is a plant of the genus '' Ficus'' found in Pakistan, India, southeast Asia, through Malaysia and into Northern Australia. Its common name is white fig; it is locally known as ''pilkhan'' and in the Kunwinjku language it is cal ...
''. According to the
Kikuyu people The Kikuyu (also ''Agĩkũyũ/Gĩkũyũ'') are a Bantu ethnic group native to Central Kenya. At a population of 8,148,668 as of 2019, they account for 17.13% of the total population of Kenya, making them Kenya's largest ethnic group. The t ...
, sacrifices to Ngai were performed under a sycomore tree (Mũkũyũ) and if one was not available, a fig tree (Mũgumo) would be used. The common fig tree is cited in the
Bible The Bible (from Koine Greek , , 'the books') is a collection of religious texts or scriptures that are held to be sacred in Christianity, Judaism, Samaritanism, and many other religions. The Bible is an anthologya compilation of texts o ...
, where in Genesis 3:7,
Adam and Eve Adam and Eve, according to the creation myth of the Abrahamic religions, were the first man and woman. They are central to the belief that humanity is in essence a single family, with everyone descended from a single pair of original ancestors ...
cover their nakedness with fig leaves. The fig fruit is also one of the traditional crops of Israel, and is included in the list of food found in the Promised Land, according to the Torah ( Deut. 8). Jesus cursed a fig tree for bearing no fruit (). The fig tree was sacred in ancient
Greece Greece,, or , romanized: ', officially the Hellenic Republic, is a country in Southeast Europe. It is situated on the southern tip of the Balkans, and is located at the crossroads of Europe, Asia, and Africa. Greece shares land borders wit ...
and
Cyprus Cyprus ; tr, Kıbrıs (), officially the Republic of Cyprus,, , lit: Republic of Cyprus is an island country located south of the Anatolian Peninsula in the eastern Mediterranean Sea. Its continental position is disputed; while it is ...
, where it was a symbol of
fertility Fertility is the capability to produce offspring through reproduction following the onset of sexual maturity. The fertility rate is the average number of children born by a female during her lifetime and is quantified demographically. Ferti ...
. File:Ficus religiosa Bo.jpg, Leaves of the sacred fig (''
Ficus religiosa ''Ficus religiosa'' or sacred fig is a species of fig native to the Indian subcontinent and Indochina that belongs to Moraceae, the fig or mulberry family. It is also known as the bodhi tree, pippala tree, peepul tree, peepal tree, pipal tree ...
'') File:เศียรพระพุทธรูปในรากโพธิ์.jpg, Fig tree roots overgrowing a sandstone Buddha statue, near Wat Maha That in Ayutthaya province, Thailand Image:Sarkaradevi Temple Ficus Tree.jpg, Ficus tree in front of Sarkaradevi Temple,
Kerala Kerala ( ; ) is a state on the Malabar Coast of India. It was formed on 1 November 1956, following the passage of the States Reorganisation Act, by combining Malayalam-speaking regions of the erstwhile regions of Cochin, Malabar, South C ...
, India


List of famous fig trees

* '' Ashvattha'' – the
world tree The world tree is a motif present in several religions and mythologies, particularly Indo-European religions, Siberian religions, and Native American religions. The world tree is represented as a colossal tree which supports the heavens, thereb ...
of
Hinduism Hinduism () is an Indian religion or ''dharma'', a religious and universal order or way of life by which followers abide. As a religion, it is the world's third-largest, with over 1.2–1.35 billion followers, or 15–16% of the global po ...
, held to be a supernatural ''F. religiosa'' * Bodhi tree – a ''F. religiosa'' *
Charybdis Fig Tree Charybdis (; grc, Χάρυβδις, Khárybdis, ; la, Charybdis, ) is a sea monster in Greek mythology. She, with the sea monster Scylla, appears as a challenge to epic characters such as Odysseus, Jason, and Aeneas. Scholarship locates her in t ...
of
Homer Homer (; grc, Ὅμηρος , ''Hómēros'') (born ) was a Greek poet who is credited as the author of the '' Iliad'' and the '' Odyssey'', two epic poems that are foundational works of ancient Greek literature. Homer is considered one of ...
's ''
Odyssey The ''Odyssey'' (; grc, Ὀδύσσεια, Odýsseia, ) is one of two major Ancient Greek literature, ancient Greek Epic poetry, epic poems attributed to Homer. It is one of the oldest extant works of literature still widely read by moder ...
'', presumably a ''F. carica'' *
Curtain Fig Tree Curtain Fig Tree is a heritage-listed tree at Curtain Fig Tree Road, Yungaburra, Tablelands Region, Queensland, Australia. It is one of the largest trees in Tropical North Queensland, Australia, and one of the best known attractions on t ...
– a ''F. virens'' *
Ficus Ruminalis The ''Ficus Ruminalis'' was a wild fig tree that had religious and mythological significance in ancient Rome. It stood near the small cave known as the Lupercal at the foot of the Palatine Hill and was the spot where according to tradition th ...
– a ''F. carica'' * '' Plaksa'' – another supernatural fig in Hinduism; usually identified as ''F. religiosa'' but is probably ''F. virens'' * Santa Barbara's Moreton Bay Fig Tree – a ''F. macrophylla'' * Sri Maha Bodhi – another ''F. religiosa'', planted in 288 BCE, the oldest human-planted tree on record * '' The Barren Fig Tree'' – Matthew 21:19 of
The Holy Bible The Bible (from Koine Greek , , 'the books') is a collection of religious texts or scriptures that are held to be sacredness, sacred in Christianity, Judaism, Samaritanism, and many other religions. The Bible is an anthologya compilation of ...
, Jesus put a curse on the tree and used this as an example for believers of the promise of the power faith in the only true God. '' *
The Great Banyan The Great Banyan is a banyan tree (''Ficus benghalensis'') located in Acharya Jagadish Chandra Bose Indian Botanic Garden, Shibpur, Howrah, near Kolkata, India. The great banyan tree draws more visitors to the garden than its collection of ex ...
– a ''F. benghalensis'', a
clonal colony A clonal colony or genet is a group of genetically identical individuals, such as plants, fungi, or bacteria, that have grown in a given location, all originating vegetatively, not sexually, from a single ancestor. In plants, an individual in ...
and once the largest organism known * Vidurashwatha – "
Vidura Vidura (Sanskrit: विदुर, lit. ''skilled'', ''intelligent'' or ''wise''), also known as Kshatri, plays a key role in the Hindu epic ''Mahabharata''. He is described as the prime minister of the Kuru kingdom and is the paternal uncle o ...
's Sacred Fig Tree", a village in India named after a famous ''F. religiosa'' that until recently stood there * Wonderboom – the largest fig tree in Pretoria, South Africa


Citations


General references

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Supporting Online Material
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External links



��Major reference site for the genus ''Ficus''
World checklist of Ficus species from the Catalogue of Life
845 species supplied by M. Hassler's World Plants.

��Multi-award-winning documentary


BBC: Fig fossil clue to early farming
Video
How the fig tree strangles other plants for survival in the rainforest
{{Authority control Moraceae genera Taxa named by Carl Linnaeus