Fanning (bees)
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The Nasonov (alternatively, Nasanov) pheromone is released by
worker Worker may refer to: * Worker, a person who performs work for a living * Laborer, a person who performs unskilled physical labour, especially in construction * Worker, a member of the working class * Worker, a member of the workforce ** Designati ...
bees Bees are winged insects closely related to wasps and ants, known for their roles in pollination and, in the case of the best-known bee species, the western honey bee, for producing honey. Bees are a monophyletic lineage within the superfamil ...
to orient returning forager bees back to the colony. To broadcast this scent, bees raise their
abdomen The abdomen (colloquially called the gut, belly, tummy, midriff, tucky, or stomach) is the front part of the torso between the thorax (chest) and pelvis in humans and in other vertebrates. The area occupied by the abdomen is called the abdominal ...
s, which contain the Nasonov
gland A gland is a Cell (biology), cell or an Organ (biology), organ in an animal's body that produces and secretes different substances that the organism needs, either into the bloodstream or into a body cavity or outer surface. A gland may also funct ...
s, and fan their
wing A wing is a type of fin that produces both Lift (force), lift and drag while moving through air. Wings are defined by two shape characteristics, an airfoil section and a planform (aeronautics), planform. Wing efficiency is expressed as lift-to-d ...
s vigorously. Nasonov includes a number of different
terpenoid The terpenoids, also known as isoprenoids, are a class of naturally occurring organic compound, organic chemicals derived from the 5-carbon compound isoprene and its derivatives called terpenes, diterpenes, etc. While sometimes used interchangeabl ...
s including
geraniol Geraniol is a monoterpenoid and an alcohol. It is the primary component of citronella oil and is a primary component of rose oil and palmarosa oil. It is a colorless oil, although commercial samples can appear yellow. It has low solubility i ...
,
nerolic acid Nerolic acid, also known as (''Z'')-3,7-dimethyl-2,6-octadienoic acid, is an organically-derived chemical. In nature Nerolic acid is found in the Nasonov scent gland of honey-bees along with geraniol, geranic acid, citral, farnesol, and nerol ...
,
citral Citral is an acyclic monoterpene aldehyde. Being a monoterpene, it is made of two isoprene units. Citral is a collective term which covers two geometric isomers that have their own separate names; the ''E''-isomer is named geranial (''trans''- ...
and
geranic acid Geranic acid, or 3,7-dimethyl-2,6-octadienoic acid, is a pheromone A pheromone () is a secreted or excreted chemical factor that triggers a social response in members of the same species. Pheromones are chemicals capable of acting like hormo ...
. Bees use these to find the entrance to their colony or hive, and they release them on flowers so other bees know which flowers have nectar. Once the foraging bee leaves the nest it uses its sense from special sensing cells on the antennae to locate and distinguish forage plants, which each give off a unique blend of odour chemicals. When a beekeeper lifts out frames from a hive they disturb the balance of smells within the hive. It can take up to 48 hours for the colony to re-establish its scent equilibrium. A synthetically produced Nasonov
pheromone A pheromone () is a secreted or excreted chemical factor that triggers a social response in members of the same species. Pheromones are chemicals capable of acting like hormones outside the body of the secreting individual, to affect the behavio ...
can be used to attract a honey bee swarm to an unoccupied hive or a swarm-catching box. Synthetically produced Nasonov consists of
citral Citral is an acyclic monoterpene aldehyde. Being a monoterpene, it is made of two isoprene units. Citral is a collective term which covers two geometric isomers that have their own separate names; the ''E''-isomer is named geranial (''trans''- ...
and
geraniol Geraniol is a monoterpenoid and an alcohol. It is the primary component of citronella oil and is a primary component of rose oil and palmarosa oil. It is a colorless oil, although commercial samples can appear yellow. It has low solubility i ...
in a 2:1 ratio. The Nasonov gland was first described in 1882 by the Russian zoologist Nikolai Viktorovich Nasonov. Nasonov thought that the gland performed perspiration; it was Frederick William Lambert Sladen (May 30, 1876 - 1921) of England who in 1901 first proposed that the gland produced a pheromone.See: * (Snodgrass, 1925), p. 116. * * Republished in condensed form in: * Reprinted in:


See also

* Nasonov's gland


References

Beekeeping Bee ecology Insect pheromones Olfaction {{Bee-stub