Franz Josef Maria Werner (15 August 1867 in

Vienna Vienna ( ; ; ) is the capital city, capital, List of largest cities in Austria, most populous city, and one of Federal states of Austria, nine federal states of Austria. It is Austria's primate city, with just over two million inhabitants. ...

– 28 February 1939 in Vienna) was an

Austrian Austrian may refer to: * Austrians, someone from Austria or of Austrian descent ** Someone who is considered an Austrian citizen * Austrian German dialect * Something associated with the country Austria, for example: ** Austria-Hungary ** Austria ...

zoologist Zoology ( , ) is the scientific study of animals. Its studies include the structure, embryology, classification, habits, and distribution of all animals, both living and extinct, and how they interact with their ecosystems. Zoology is one ...

and

explorer Exploration is the process of exploring, an activity which has some Expectation (epistemic), expectation of Discovery (observation), discovery. Organised exploration is largely a human activity, but exploratory activity is common to most organis ...

. Specializing as a

herpetologist Herpetology (from Ancient Greek ἑρπετόν ''herpetón'', meaning "reptile" or "creeping animal") is a branch of zoology concerned with the study of amphibians (including frogs, salamanders, and caecilians (Gymnophiona)) and reptiles (in ...

and

entomologist Entomology (from Ancient Greek ἔντομον (''éntomon''), meaning "insect", and -logy from λόγος (''lógos''), meaning "study") is the branch of zoology that focuses on insects. Those who study entomology are known as entomologists. In ...

, Werner described numerous species and other taxa of frogs, snakes, insects, and other organisms. His father introduced him at age six to

reptile Reptiles, as commonly defined, are a group of tetrapods with an ectothermic metabolism and Amniotic egg, amniotic development. Living traditional reptiles comprise four Order (biology), orders: Testudines, Crocodilia, Squamata, and Rhynchocepha ...

s and

amphibian Amphibians are ectothermic, anamniote, anamniotic, tetrapod, four-limbed vertebrate animals that constitute the class (biology), class Amphibia. In its broadest sense, it is a paraphyletic group encompassing all Tetrapod, tetrapods, but excl ...

s. A brilliant student, he corresponded often with

George Albert Boulenger George Albert Boulenger (19 October 1858 – 23 November 1937) was a Belgian-British zoologist who described and gave scientific names to over 2,000 new animal species, chiefly fish, reptiles, and amphibians. Boulenger was also an active botani ...

(1858–1937) and

Oskar Boettger Oskar Boettger (; 31 March 1844 – 25 September 1910) was a German zoologist who was a native of Frankfurt am Main. He was an uncle of the noted malacologist Caesar Rudolf Boettger (1888–1976). From 1863 to 1866 he studied at the Bergakademie ...

(1844–1910) who encouraged his studies with these animals. Werner obtained his doctorate in Vienna in 1890 and then after spending a year in

Leipzig Leipzig (, ; ; Upper Saxon: ; ) is the most populous city in the States of Germany, German state of Saxony. The city has a population of 628,718 inhabitants as of 2023. It is the List of cities in Germany by population, eighth-largest city in Ge ...

, began to teach at the Vienna Institute of Zoology. In 1919, he became tenured as a professor, maintaining this title until his retirement in 1933. Although working close to the Vienna Natural History Museum, he could not use their herpetological collections, after the death of its director,

Franz Steindachner Franz Steindachner (11 November 1834 in Vienna – 10 December 1919 in Vienna) was an Austrian zoologist, ichthyologist, and herpetologist. He published over 200 papers on fishes and over 50 papers on reptiles and amphibians. Steindachner des ...

(1834–1919), who did not like Werner, and had barred him from accessing the collections. Werner succeeded in constituting an immense personal collection, and published more than 550 publications principally on herpetology. He named many new species of reptiles, amphibians, and

arthropod Arthropods ( ) are invertebrates in the phylum Arthropoda. They possess an arthropod exoskeleton, exoskeleton with a cuticle made of chitin, often Mineralization (biology), mineralised with calcium carbonate, a body with differentiated (Metam ...

s, of which he specialized in

orthoptera Orthoptera () is an order of insects that comprises the grasshoppers, locusts, and crickets, including closely related insects, such as the bush crickets or katydids and wētā. The order is subdivided into two suborders: Caelifera – gras ...

ns and

scorpion Scorpions are predatory arachnids of the Order (biology), order Scorpiones. They have eight legs and are easily recognized by a pair of Chela (organ), grasping pincers and a narrow, segmented tail, often carried in a characteristic forward cur ...

s. He published in 1931, ''Dritte Klasse der Craniota: dritte und zugleich letzte Klasse der Ichthyopsida: Amphibia, Lurche: allgemeine Einleitung in said Naturgeschichte der Amphibia.'' His book, ''Amphibien und Reptilien'' (1910), contributed to the popularization of terraphilia, or raising pet reptiles and amphibians in

terraria ''Terraria'' ( ) is a 2011 action-adventure sandbox game developed by Re-Logic and published by 505 Games. The game features exploration, combat, crafting, building, and mining inside a procedurally generated 2D computer graphics, 2D world. ...

. ''

Nota bene ( ; plural: ) is the Latin language, Latin phrase meaning ''note well''. In manuscripts, ''nota bene'' is abbreviated in upper-case as NB and N.B., and in lower-case as n.b. and nb; the editorial usages of ''nota bene'' and ''notate bene'' fi ...

'': Franz Werner should not be confused with Israeli herpetologist Yehudah L. Werner (born 1931).

List of taxa described by Franz Werner

higher order taxa

* Family:

Leptodactylidae The southern frogs form the Leptodactylidae, a name that comes from Greek meaning a bird or other animal having slender toes. They are a diverse family of frogs that most likely diverged from other hyloids during the Cretaceous. The family has u ...

(Werner, 1896) (49 genera of Southern frogs or tropical frogs) * Subfamily:

Leptodactylinae The Leptodactylinae are a subfamily of leptodactylid frogs distributed between southern North America (Texas, USA, and Sonora, Mexico) and South America to Brazil. Their sister taxon is the clade comprising the Leiuperinae and Paratelmatobiina ...

(Werner, 1896) * Genus:

Astylosternus ''Astylosternus'' (commonly known as night frogs) is a genus of frogs in the family Arthroleptidae. It contains 12 species found in an area spanning from Sierra Leone in West Africa to the Democratic Republic of Congo in Central Africa, with a ga ...

(Werner, 1898)

Species

Fishes

* ''

Himantura schmardae ''Himantura'' is a genus of stingray in the family Dasyatidae that is native to the Indo-Pacific. In a 2016 taxonomic revision, many of the species formerly assigned to ''Himantura'' were reassigned to other genera ('' Brevitrygon'', '' Fluvitry ...

'' (Werner, 1904) (Chupare Stingray) * ''

Neobola nilotica ''Neobola'' is a genus of freshwater ray-finned fishes belonging to the family Danionidae, the danios or danionins. The fishes in this genus are found mostly in East Africa. There are currently five species in this genus. Species ''Neobola'' ...

'' Werner, 1919

Amphibians

=Frogs and Toads

= *

Turkish Frog The long-legged wood frog (''Rana macrocnemis''), also known as Caucasus frog, Brusa frog, or Uludağ frog, is a species of frog in the family Ranidae found in Armenia, Azerbaijan, Georgia, Iran, Russia, Turkey, and Turkmenistan. Its natural h ...

, ''

Rana holtzi Rana may refer to: Astronomy * Rana (crater), a crater on Mars * Delta Eridani or Rana, a star Films * Rana (2012 film), an Indian Kannada-language action drama * Rana, a 1998 Telugu-language action film directed by A. Kodandarami Reddy * R ...

'' (Werner, 1898) * ''

Rana leporipes ''Odorrana leporipes'' is a species of frog in the family Ranidae. It is endemic to Guangdong province of China and only known from its type locality, "Lung Tao Shan" (=Mount Longdao) in northern Guangdong. It is only known from the original sp ...

'' (Werner, 1930) *

Nikkō Frog The montane brown frog or Nikkō frog (''Rana ornativentris'') is a species of frog in the family Ranidae. It is endemic to Japan. Its natural habitats are subtropical or tropical dry forests, rivers, swamps, freshwater marshes, arable land ...

, '' Rana ornativentris'' (Werner, 1903) * Garman New Guinea Tree Frog, ''

Litoria jeudii The Garman New Guinea tree frog ("''Litoria''" ''jeudii'') is a species of frog in the subfamily Pelodryadinae. This species is endemic to Papua New Guinea, and the type Type may refer to: Science and technology Computing * Typing, producing ...

'' (Werner, 1901) * Humming Frog, '' Neobatrachus pelobatoides'' (Werner, 1914) * '' Eleutherodactylus affinis'' (Werner, 1899) * '' Eleutherodactylus appendiculatus'' (Werner, 1894) * '' Eleutherodactylus bisignatus'' (Werner, 1899) * ''

Eleutherodactylus frater ''Pristimantis frater'' is a species of frog in the family Strabomantidae. It is endemic to Colombia and occurs on the eastern slopes of the Cordillera Oriental in the Norte de Santander, Boyacá, Casanare, Cundinamarca, and Meta Departments, ...

'' (Werner, 1899) * '' Eleutherodactylus laevissimus'' (Werner, 1896) * '' Eleutherodactylus rostralis'' (Werner, 1896) * '' Rheobates palmatus'' (Werner, 1899) * '' Astylosternus diadematus'' (Werner, 1898) * '' Leptopelis boulengeri'' (Werner, 1898) * '' Leptopelis brevirostris'' (Werner, 1898) * '' Leptopelis modestus'' (Werner, 1898) * '' Craugastor laevissimus'' (Werner, 1896) * '' Craugastor rostralis'' (Werner, 1896) * '' Eleutherodactylus affinis'' (Werner, 1899) * '' Eleutherodactylus appendiculatus'' (Werner, 1894) * '' Eleutherodactylus bisignatus'' (Werner, 1899) * ''

'' (Werner, 1899) * '' Phrynopus columbianus'' (Werner, 1899) * ''

Atelopus subornatus ''Atelopus subornatus'' is a species of toad in the family Bufonidae. It is endemic to Colombia and is restricted to the Cordillera Oriental in the Cundinamarca Department. Common name Bogota stubfoot toad has been coined for this species. Des ...

'' (Werner, 1899) * ''

Chaunus limensis ''Rhinella limensis'' is a species of toad in the family Bufonidae. It is endemic to Peru. Its natural habitats are rivers, hot deserts, sandy shores, arable land, and rural garden A garden is a planned space, usually outdoors, set aside for ...

'' (Werner, 1901) * '' Chaunus schneideri'' (Werner, 1894) * '' Rhaebo nasicus'' (Werner, 1903) * '' Wolterstorffina parvipalmata'' (Werner, 1898) * '' Telmatobius verrucosus'' (Werner, 1899) * '' Cycloramphus asper'' (Werner, 1899) * '' Cycloramphus bolitoglossus'' (Werner, 1897) * ''

Fejervarya schlueteri The frog family (biology), family Dicroglossidae occurs in tropical and subtropical regions of Asia and Africa, with most genera and species being found in Asia. The common name of the family is fork-tongued frogs. The Dicroglossidae were previou ...

'' (Werner, 1893) * '' Hypsiboas pellucens'' (Werner, 1901) * '' Osteocephalus verruciger'' (Werner, 1901) * '' Scinax dolloi'' (Werner, 1903) * ''

Sphaenorhynchus platycephalus ''Sphaenorhynchus platycephalus'', commonly known as the South American lime treefrog or Lutz's lime treefrog, is a species of frog in the family Hylidae. It is endemic to south-eastern Brazil where it occurs in the Serra do Mar and Serra da ...

'' (Werner, 1894) * ''

'' (Werner, 1901) * '' Afrixalus quadrivittatus'' (Werner, 1908) * Incertae sedis: '' Hyperolius papyri'' (Werner, 1908) * '' Hyperolius balfouri'' (Werner, 1908) * '' Neobatrachus pelobatoides'' (Werner, 1914) * '' Oreophryne brachypus'' (Werner, 1898) * '' Oreophryne wolterstorffi'' (Werner, 1901) * '' Plethodontohyla angulifera'' (Werner, 1903) * ''

Phrynomantis annectens The marbled rubber frog (''Phrynomantis annectens'') is a species of frog in the family Microhylidae. It is native to Angola, Namibia, and South Africa. Its natural habitats are dry savanna, subtropical or tropical dry shrubland, intermittent f ...

'' (Werner, 1910) * '' Ptychadena aequiplicata'' (Werner, 1898) * '' Ptychadena schillukorum'' (Werner, 1908) * Incertae Sedis: '' Rana temporaria var. nigromaculata'' (Werner, 1897) * '' Huia leporipes'' (Werner, 1930) * ''

'' (Werner, 1898) * '' Rana ornativentris'' (Werner, 1903) * '' Buergeria pollicaris'' (Werner, 1914)

=Salamanders

= * '' Bolitoglossa dofleini'' (Werner, 1903) * '' Bolitoglossa palmata'' (Werner, 1897)

=Caecilians

= * '' Crotaphatrema bornmuelleri'' (Werner, 1899)

Reptiles

=Snakes

= * '' Vipera bornmuelleri'' (Werner, 1898) * '' Daboia palaestinae'' (Werner, 1938) * '' Bothrops oligolepis'' (Werner, 1901) * ''

Atheris ceratophora :''Common names: Usambara bush viper,Spawls S, Branch B. 1995. The Dangerous Snakes of Africa. Ralph Curtis Books. Dubai: Oriental Press. 192 pp. . horned bush viper, eyelash bush viper, more.'' ''Atheris ceratophora'' is a venomous viper speci ...

'' (Werner, 1895) * '' Macrovipera schweizeri'' (Werner, 1935) * ''

Paranaja multifasciata The many-banded snake (''Naja multifasciata''), also known commonly as the burrowing cobra, is a species of venomous snake in the family Elapidae. The species is native to Central Africa. There are three recognized subspecies. Geographic range ...

'' (Werner, 1902) * ''

Elaphe persica The Persian ratsnake (''Zamenis persicus'') is a species of medium-sized nonvenomous snake in the family Colubridae. The species is endemic to Western Asia. Geographic range ''Z. persicus'' is found in temperate northwestern Iran and Azerbaijan, ...

'' (Werner, 1913) * '' Elapsoidea laticincta'' (Werner, 1919) * '' Coluber andreanus'' (Werner, 1917) * '' Lampropeltis triangulum arcifera'' (Werner, 1903) * '' Leptotyphlops rubrolineatus'' (Werner, 1901) * '' Micrurus diastema alienus'' (Werner, 1903) * '' Micrurus diastema sapperi'' (Werner, 1903) * '' Micrurus frontifasciatus'' (Werner, 1927) * '' Micrurus lemniscatus frontifasciatus'' (Werner, 1927) * '' Micrurus multifasciatus hertwigi'' (Werner, 1897) * '' Micrurus steindachneri steindachneri'' (Werner, 1901) * '' Morelia spilota macrospila'' (Werner, 1910) * '' Chrysopelea ornata ornatissima'' (Werner, 1925)

=Lizards

= * Werner's Leaf-toed Gecko, ''

Asaccus elisae ''Asaccus elisae'', also known as Elisa's leaf-toed gecko or Werner's leaf-toed gecko, is a species of gecko, a lizard in the Family (biology), family Phyllodactylidae. The species is native to the Middle East. Etymology The Specific name (zoolo ...

'' * Ebner's Skink, '' Chalcides ebneri'' ( Werner, 1931) * '' Gastropholis prasina'' ( Werner, 1904)

=Amphibaenians

= * '' Monopeltis leonhardi''

Insects

* '' Creobroter fasciatus'' (Werner, 1927) (a west Asian

mantid Mantidae is one of the largest families in the order of praying mantises, based on the type species ''Mantis religiosa''; most genera are tropical or subtropical. Historically, this was the only family in the order, and many references still ...

) * '' Hebardiella karnyi'' (Werner, 1924) (a west Asian

) * '' Hebardiella rehni'' (Werner, 1924) (a west Asian

)

Taxa named in honor of Franz Werner

*'' Achalinus werneri'' (snake species)Beolens, Bo; Watkins, Michael; Grayson, Michael (2011). ''The Eponym Dictionary of Reptiles''. Baltimore: Johns Hopkins University Press. xiii + 296 pp. . ("Werner, F.", p. 282). *'' Aparallactus werneri'' (snake species) *'' Arthroseps werneri'' (lizard species) = ''

Ecpleopus gaudichaudii ''Ecpleopus'' is a genus of lizard in the family Gymnophthalmidae. The genus contains only one species, ''Ecpleopus gaudichaudii'', which is endemic to Brazil. Etymology The specific name, ''gaudichaudii'', is in honor of French botanist Charl ...

'' *'' Atractus werneri'' (snake species) *''

Chamaelycus werneri ''Chamaelycus'' is a genus of snakes, commonly referred to as banded snakes, in the family Lamprophiidae. The genus is endemic to Central Africa. Species The following three species are recognized as being valid.. www.reptile-database.org. *'' ...

'' (snake species) *'' Clarias werneri'' Boulenger, 1906 Walking Catfish from Africa. *'' Dendropsophus werneri'' (frog species) *'' Dipsadoboa werneri'' (snake species) *''

Firmicus werneri Firmicus may refer to: *The lunar crater Firmicus. *The Christian astrologer Julius Firmicus Maternus __NOTOC__ Julius Firmicus Maternus was a Roman Latin writer and astrologer, who received a pagan classical education that made him conversant w ...

'' (spider species) *'' Iguanognathus werneri'' (snake species) *'' Olios werneri'' (spider species) *'' Pachydactylus werneri'' (gecko species) *'' Stenaelurillus werneri'' (spider species) *'' Trioceros werneri'' (chameleon species) *''

Werneria ''Werneria'', also known as the torrent toads or smalltongue toads, is a small genus of "true toads", family Bufonidae. They are found in western Central Africa (Equatorial Guinea, Gabon, and Cameroon), with the greatest species richness in the ...

'' (genus of toads) *'' Xenodon werneri'' (snake species)

References

External links

Amphibian Species of the World
from the

American Museum of Natural History The American Museum of Natural History (AMNH) is a natural history museum on the Upper West Side of Manhattan in New York City. Located in Theodore Roosevelt Park, across the street from Central Park, the museum complex comprises 21 interconn ...

. {{DEFAULTSORT:Werner, Franz Austrian entomologists Austrian herpetologists 1867 births 1939 deaths Scientists from Vienna Zoologists from Austria-Hungary