Eysarcorini

Pentatominae is a subfamily of Pentatomidae, a family of shield bugs. This subfamily is the largest one within the Pentatomidae, having 4937 species classified in 938 genera. Species in this subfamily are phytophages and several of them are considered agricultural pests. Some invasive pentatomines such as '' Halyomorpha halys'' and '' Bagrada hilaris'' have been considered household pests. Higher systematics of the group have been revised by Rider et al.

Tribes and genera

Tribes within this subfamily include:

* Aelini Douglas & Scott, 1865
# '' Aelia'' Fabricius, 1803
# '' Aeliopsis'' Bergevin, 1931
# '' Neottiglossa'' Kirby, 1837
* Aeptini Stål, 1871
# '' Aeliosoma'' Baehr, 1985
# '' Aeptus'' Dallas, 1851
# '' Caridillus'' Bergroth, 1909
# '' Eribotes (bug)'' Stål, 1867
# '' Hillieria'' Distant, 1910
# '' Mariomella'' Faúndez & Rider, 2019
# '' Menestheus (bug)'' Stål, 1867
# '' Paramenestheus'' Breddin, 1900
# '' Pseudaelia'' Distant, 1910
* Aeschrocorini Distant, 1902
# '' Aeschrocoris'' Bergroth, 1887
# '' Aeschrus'' Spinola, 1850
# '' Chraesus'' Cachan, 1952
# ''Geomorpha (bug)'' Bergroth, 1893
# '' Risbecella'' Schouteden, 1942
# '' Scylax (bug)'' Distant, 1887
# '' Tyoma'' Spinola, 1850
# '' Tyomana'' Miller, 1952
* Agaeini Cachan, 1952
# '' Agaeus'' Dallas, 1851
* Agonoscelidini Atkinson, 1888
# '' Agonoscelis'' Spinola, 1837
* Amyntorini Distant, 1902
# '' Ambohicorypha'' Cachan, 1952
# '' Belopis'' Distant, 1879
# ''Bolaca'' Walker, 1867
# '' Halyabbas'' Distant, 1900
# '' Madecorypha'' Cachan, 1952
# '' Sennertus'' Distant, 1900
* Antestiini Distant, 1902 - selected genera:
** '' Antestia'' Stål, 1865
** '' Antestiopsis'' Leston, 1952
** '' Dryadocoris'' Kirkaldy, 1909
** '' Plautia'' Stål, 1864
* Aulacetrini Mulsant & Rey, 1866
# '' Holcogaster'' Fieber, 1860
* Axiagastini Atkinson, 1888
# '' Acesines'' Stål, 1876
# '' Axiagastus'' Dallas, 1851
# '' Indrapura (bug)'' China, 1928
# '' Oncotropis'' Stål, 1871
* Bathycoeliini Atkinson, 1888
# '' Bathycoelia'' Amyot & Serville, 1843
* Cappaeini Atkinson, 1888
# '' Adelolcus'' Bergroth, 1893
# '' Benia (bug)'' Schouteden, 1916
# '' Boerias'' Kirkaldy, 1909
# '' Cappaea'' Ellenrieder, 1862
# '' Caura (bug)'' Stål, 1865
# '' Cauromorpha'' Jeannel, 1913
# '' Erlangerella'' Schouteden, 1905
# '' Halycorypha'' Jeannel, 1913
# '' Halydicoris'' Jeannel, 1913
# '' Halyoides'' Cachan, 1952
# '' Halyomorpha'' Mayr, 1864
# '' Homalogonia'' Jakovlev, 1876
# '' Hymenomaga'' Karsch, 1892
# '' Lerida (bug)'' Karsch, 1894
# '' Leridella'' Jeannel, 1913
# '' Lokaia'' Linnavuori, 1975
# '' Mabusana'' Distant, 1912
# '' Massocephalus'' Dallas, 1851
# '' Paralerida'' Linnavuori, 1982
# '' Prytanicoris'' Gross, 1978
# '' Tolumnia'' Stål, 1867
# '' Tripanda'' Berg, 1899
# '' Tropicorypha'' Mayr, 1864
# '' Veterna'' Stål, 1865
* Carpocorini Mulsant & Rey, 1866 selected genera:
** '' Antheminia'' Mulsant & Rey, 1866
** '' Carpocoris'' Kolenati, 1846
** '' Chlorochroa'' Stål, 1872
** '' Dolycoris'' Mulsant & Rey, 1866
** '' Graphorn''
** '' Peribalus'' Mulsant & Rey, 1866
** '' Rubiconia'' Dohrn, 1860
** '' Thestral''
* Catacanthini Atkinson, 1888
# '' Arocera'' Spinola, 1837
# '' Boea (bug)'' Walker, 1867
# '' Catacanthus'' Spinola, 1837
# '' Chalcocoris'' Dallas, 1851
# '' Rhyssocephala'' Rider, 1992
# '' Runibia'' Stål, 1861
# '' Vitruvius (bug)'' Distant, 1901
# '' Vulsirea'' Spinola, 1837
* Caystrini Ahmad & Afzal, 1979
# '' Caystrus'' Stål, 1861
# '' Cephaloplatus'' White, 1842
# '' Heissocoris'' Ahmad & Afzal, 1979
# '' Hippotiscus'' Bergroth, 1906
# '' Kyrtalus'' Van Duzee, 1929
# '' Lyalocoris'' Zahid, 2006
# '' Mcdonaldiana'' Zahid, 2006
# '' Neodius'' (Stål, 1867)
# '' Neoparodius'' Zahid, 2006
# '' Neoriazocoris'' Zahid, 2006
# '' Paramecocoris'' Stål, 1853
# '' Parodius (bug)'' Distant, 1900
# '' Praetextatus (bug)'' Distant, 1901
# '' Riazocoris'' Ahmad & Afzal, 1979
# '' Valescus'' Distant, 1901
* Chlorocorini Rider, Greve, Schwertner & Grazia, 2018
# '' Arvelius'' Spinola, 1837
# '' Chlorocoris'' Spinola, 1837
# '' Chloropepla'' Stål, 1867
# '' Eludocoris'' Thomas, 1992
# '' Fecelia'' Stål, 1872
# '' Mayrinia'' Horváth, 1925
# '' Rhyncholepta'' Bergroth, 1911
* Coquereliini Cachan, 1952
# '' Cleoqueria'' Cachan, 1952
# '' Coquerelia'' Signoret, 1861
# '' Coquerelidea'' Reuter, 1887
# '' Neocoquerelidea'' Cachan, 1952
* Degonetini Azim & Shafee, 1984
# '' Degonetus'' Distant, 1902
* Diemeniini Kirkaldy, 1909 - selected genera:
** '' Commius (bug)'' Stål, 1876
** '' Diemenia (bug)'' Spinola, 1850
* Diplostirini Distant, 1902
# '' Ambiorix (bug)'' Stål, 1876
# '' Diplostira'' Dallas, 1851
* Diploxyini Atkinson, 1888
# '' Acoloba'' Spinola, 1850
# '' Aeladria'' Cachan, 1952
# '' Diploxys'' Amyot & Serville, 1843
* Eurysaspidini Atkinson, 1888
# '' Eurysaspis'' Signoret, 1851
# '' Flaminia (bug)'' Stål, 1865
* Eysarcorini Mulsant & Rey, 1866
# '' Adria (bug)'' Stål, 1876
# '' Aspavia'' Stål, 1865
# '' Cachanocoris'' Rider, 1998
# '' Carbula'' Stål, 1864
# '' Corisseura'' Cachan, 1952
# '' Cratonotus'' Distant, 1879
# '' Durmia'' Stål, 1865
# '' Dymantiscus'' Hsiao, 1981
# '' Empiesta'' Linnavuori, 1982
# '' Eysarcoris'' Hahn, 1834
# '' Hermolaus'' Distant, 1902
# '' Mulungua'' Linnavuori, 1982
# '' Pseudolerida'' Schouteden, 1958
# '' Sepontia'' Stål, 1865
# '' Sepontiella'' Miyamoto, 1990
# '' Spermatodes'' Bergroth, 1914
# '' Stagonomus'' Gorski, 1852
* Halyini Amyot & Serville, 1843 - selected genera:
** '' Brochymena'' Amyot & Serville, 1843
* Hoplistoderini Atkinson, 1888
# '' Alcimocoris'' Bergroth, 1891
# '' Glottaspis'' Bergroth, 1918
# '' Hoplistodera'' Westwood, 1837
# '' Paracritheus'' Bergroth, 1891
# '' Paralcimocoris'' Zhang & Lin, 1988
# '' Stachyomia'' Stål, 1871
# '' Taurodes'' Dallas, 1851
* Lestonocorini Ahmad & Mohammad, 1980
# '' Dardjilingia'' Yang, 1936
# '' Gynenica'' Dallas, 1851
# '' Lestonocoris'' Ahmad & Mohammad, 1980
# '' Neogynenica'' Yang, 1935
# '' Umgababa (bug)'' Leston, 1953
* Mecideini Distant, 1902
# ''Mecidea'' Dallas, 1851
* Memmiini Cachan, 1952
# '' Memmia'' Stål, 1865
* Menidini Atkinson, 1888
# '' Actuarius (bug)'' Distant, 1901
# '' Aegaleus (bug)'' Stål, 1876
# '' Amphimachus (bug)'' Stål, 1867
# '' Anchesmus'' Stål, 1876
# '' Apines'' Dallas, 1851
# '' Aspideurus'' Signoret, 1880
# '' Banya (bug)'' Schouteden, 1916
# '' Brachycoris'' Stål, 1871
# '' Carvalhocoris'' Leston, 1953
# '' Cresphontes (bug)'' Stål, 1867
# '' Dabessus'' Distant, 1902
# '' Decellella'' Schouteden, 1964
# '' Desertomenida'' Kiritshenko, 1914
# '' Dunnius'' Distant, 1902
# '' Elanela'' Rolston, 1980
# '' Eurymenida'' Bergroth, 1917
# '' Gwea'' Schouteden, 1958
# '' Keleacoris'' Rider & Rolston, 1995
# '' Lathraedoeus'' Breddin, 1900
# '' Menida'' Motschulsky, 1861
# '' Neostrachia'' Saunders, 1877
# '' Phavorinus'' Distant, 1902
# '' Rio (bug)'' Kirkaldy, 1909
# '' Saceseurus'' Breddin, 1900
# '' Sciomenida'' Gross, 1976
# '' Solenocoris'' Oshanin, 1922
# '' Udonga'' Distant, 1921
* Myrocheini Stål, 1871
# '' Aednulus'' Breddin, 1901
# '' Aednus'' Dallas, 1851
# '' Arniscus'' Distant, 1899
# '' Delegorguella'' Spinola, 1850
# '' Dictyotus'' Dallas, 1851
# '' Dictytotus'' Dallas, 1851
# '' Dissocolpus'' Bergroth, 1906
# '' Dollingiana'' Ahmad & Kamaluddin, 1986
# '' Dymantis'' Stål, 1861
# '' Ennius (bug)'' Stål, 1861
# '' Erachtheus'' Stål, 1861
# '' Humria'' Linnavuori, 1975
# '' Jitka (bug)'' Kment, 2015
# '' Laprius'' Stål, 1861
# '' Lodosia'' Ahmad & Önder, 1996
# '' Munshiana'' Ahmad & Kamaluddin, 1987
# '' Myrochea'' Amyot & Serville, 1843
# '' Neococalus'' Bergroth, 1891
# '' Neodorpius'' Ahmad & Afzal, 1989
# '' Paradictyotus'' Gross, 1975
# '' Pretorius (bug)'' Distant, 1898
# '' Stysicoris'' Ahmad & Kamaluddin, 1985
# '' Tholosanus'' Distant, 1899
# '' Utheria'' Gross, 1975
* Nealeriini Cachan, 1952
# '' Nealeria'' Bergroth, 1893
# '' Paraleria'' Reuter, 1887
* Nezarini Atkinson, 1888 - selected genera:
** '' Nezara'' Amyot et Serville, 1843
** '' Palomena'' Mulsant et Rey, 1866
* Opsitomini Cachan, 1952
# '' Opsitoma'' Cachan, 1952
* Pentamyrmexini Rider & Brailovsky, 2014 (monotypic, Thailand)
** '' Pentamyrmex'' Rider & Brailovsky, 2014
* Pentatomini Leach, 1815 - selected genera:
** '' Pentatoma'' Olivier, 1789
* Phricodini Cachan, 1952
# '' Phricodus'' Spinola, 1839
* Piezodorini Atkinson, 1888
# '' Anaximenes (bug)'' Stål, 1876
# '' Chaubattiana'' Distant, 1912
# '' Pausias (bug)'' Jakovlev, 1905
# '' Piezodorus'' Fieber, 1860
* Procleticini Pennington, 1920
# '' Aleixus'' McDonald, 1981
# '' Brepholoxa'' Van Duzee, 1904
# '' Dendrocoris'' Bergroth, 1891
# '' Lobepomis'' Berg, 1891
# '' Neoderoploa'' Pennington, 1922
# '' Odmalea'' Bergroth, 1914
# '' Parodmalea'' Rider, 1994
# '' Procleticus'' Berg, 1891
# '' Pseudobebaeus'' Distant, 1911
# '' Terania'' Pirán, 1963
# '' Thoreyella'' Spinola, 1850
* Rhynchocorini Stål, 1871
# '' Araducta'' Walker, 1867
# '' Avicenna (bug)'' Distant, 1901
# '' Biprorulus'' Breddin, 1900
# '' Cuspicona'' Dallas, 1851
# '' Diaphyta'' Bergroth, 1891
# '' Everardia (bug)'' Gross, 1975
# '' Hoffmanseggiella'' Spinola, 1850
# '' Koogobatha'' Distant, 1910
# ''Morna (bug)'' Stål, 1867
# '' Mygoodano'' Distant, 1910
# '' Ocirrhoe'' Stål, 1876
# '' Parocirrhoe'' Gross, 1975
# '' Pegala'' Stål, 1867
# '' Petalaspis'' Bergroth, 1916
# '' Pugione'' Stål, 1867
# '' Rhynchocoris'' Westwood, 1837
# '' Utana'' Distant, 1900
# ''Vitellus'' Stål, 1865
* Rolstoniellini Rider, 1997
# '' Agathocles (bug)'' Stål, 1876
# '' Amasenus'' Stål, 1863
# '' Amatembuna'' Distant, 1910
# '' Critheus'' Stål, 1867
# '' Exithemus'' Distant, 1902
# '' Rolstoniellus'' Rider, 1997
* Sciocorini Amyot & Serville, 1843 selected genera:
** '' Dyroderes'' Spinola, 1837
** '' Sciocoris'' Fallén, 1829
* Strachiini Mulsant & Rey, 1866 selected genera:
** '' Afrania'' Stål, 1865
** '' Bagrada'' Stål, 1862
** '' Eurydema'' Laporte de Castelnau, 1833
** '' Murgantia (bug)'' Stål, 1862
** '' Stenozygum'' Fieber, 1861
** '' Trochiscocoris'' Reuter, 1890
* Triplatyxini Cachan, 1952
# '' Anoano'' Cachan, 1952
# '' Nene (bug)'' Kment, 2015
# '' Tricompastes'' Cachan, 1952
# '' Triplatyx'' Horváth, 1904

Genera ''

incertae sedis''

# '' Amphidexius'' Bergroth, 1918
# '' Bachesua'' Gross, 1975
# '' Birna'' McDonald, 2006
# '' Brizocoris'' Gross, 1975
# '' Buthumka'' Gross, 1975
# '' Cooperocoris'' Gross, 1975
# '' Dysnoetus'' Bergroth, 1918
# '' Eremophilacoris'' Gross, 1975
# '' Gadarscama'' Reuter, 1887
# '' Ippatha'' Distant, 1910
# '' Kitsonia'' Gross, 1975
# '' Kitsoniocoris'' Rider, 1998
# '' Kumbutha'' Distant, 1910
# '' Lakhonia'' Yang, 1936
# '' Macrocarenoides'' Gross, 1975
# '' Macrocarenus'' Stål, 1873
# '' Mcphersonarcys'' Thomas, 2012
# '' Muscanda'' Walker, 1868
# '' Ochisme'' Kirkaldy, 1904
# '' Ooldeon'' Gross, 1975
# '' Patanius'' Rolston, 1987
# '' Pirricoris'' Gross, 1975
# '' Poecilotoma'' Dallas, 1851
# '' Riaziana'' Hasan, Afzal & Ahmad, 1989
# '' Senectius'' Rolston, 1987
# '' Tepperocoris'' Gross, 1975
# '' Thryptomenecoris'' Gross, 1975
# '' Trachyops'' Dallas, 1851
# '' Turrubulana'' Distant, 1910
# '' Xiengia'' Distant, 1921
# '' Zouicoris'' Zheng, 1986

Notable species

*'' Acrostemum hilare'' – green stink bug
*'' Bagrada hilaris'' – bagrada bug
*'' Bathycoelia thalassina'' - African cocoa shield bug
*'' Halyomorpha halys'' – brown marmorated stink bug
*''Nezara viridula'' - green vegetable bug
*'' Pentatoma rufipes'' – forest bug
*'' Piezodorus lituratus'' – gorse shield bug

References

External links

*
*
{{Taxonbar, from=Q1949348

Hemiptera subfamilies