Eyles's Harrier
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Eyles's harrier (''Circus teauteensis'') () is an
extinct Extinction is the termination of an organism by the death of its Endling, last member. A taxon may become Functional extinction, functionally extinct before the death of its last member if it loses the capacity to Reproduction, reproduce and ...
bird of prey Birds of prey or predatory birds, also known as (although not the same as) raptors, are hypercarnivorous bird species that actively predation, hunt and feed on other vertebrates (mainly mammals, reptiles and smaller birds). In addition to speed ...
that was
endemic Endemism is the state of a species being found only in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also foun ...
to
New Zealand New Zealand () is an island country in the southwestern Pacific Ocean. It consists of two main landmasses—the North Island () and the South Island ()—and List of islands of New Zealand, over 600 smaller islands. It is the List of isla ...
.


Name

This species was named after Jim Eyles, paleontologist and former director of the
Nelson Provincial Museum The Nelson Provincial Museum, ''Pupuri Taonga O Te Tai Ao'' is a regional museum in the city of Nelson, New Zealand, Nelson, New Zealand. The museum showcases the Nelson and Tasman regions' history, from geological origins to the stories of indiv ...
and the West Coast Museum.


Description

It was an example of
island gigantism Island gigantism, or insular gigantism, is a biological phenomenon in which the size of an animal species isolated on an island increases dramatically in comparison to its mainland relatives. Island gigantism is one aspect of the more general "is ...
, as an adult female weighed around , over twice as much as a
swamp harrier The swamp harrier (''Circus approximans''), also known as the Australasian marsh harrier or Australasian harrier, is a large, slim bird of prey widely distributed across Australasia. In New Zealand, it is also known by the Māori name . It arriv ...
. Its shape differed from that of most other harriers, and it was initially mistaken for a huge
hawk Hawks are birds of prey of the family Accipitridae. They are very widely distributed and are found on all continents, except Antarctica. The subfamily Accipitrinae includes goshawks, sparrowhawks, sharp-shinned hawks, and others. This ...
, possibly a giant ''
Accipiter ''Accipiter'' () is a genus of birds of prey in the family Accipitridae. Some species are called sparrowhawks, but there are many sparrowhawks in other genera such as '' Tachyspiza''. These birds are slender with short, broad, rounded wings and ...
''.


Ecology

It was a generalist predator, taking prey of the same size as small
eagle Eagle is the common name for the golden eagle, bald eagle, and other birds of prey in the family of the Accipitridae. Eagles belong to several groups of Genus, genera, some of which are closely related. True eagles comprise the genus ''Aquila ( ...
species do: land animals weighing one or a few
kilogram The kilogram (also spelled kilogramme) is the base unit of mass in the International System of Units (SI), equal to one thousand grams. It has the unit symbol kg. The word "kilogram" is formed from the combination of the metric prefix kilo- (m ...
s. In its hunting strategy, however, it was more adapted to avian prey, as aside from bats, mammals were entirely absent from New Zealand. Presumably, it hunted diurnal birds in a manner similar to
goshawk Goshawk may refer to several species of birds of prey in the family Accipitridae: Palearctic * Eurasian goshawk, ''Astur gentilis'', often referred to simply as the goshawk, since it is the only goshawk found in much of its range (in Europe and ...
s.


Taxonomy

Eyles' harrier was presumably somewhat similar to the living
spotted harrier The spotted harrier (''Circus assimilis'') also known as the smoke hawk, is a large Australasian bird of prey belonging to the family Accipitridae. Taxonomy The spotted harrier belongs to the family Accipitridae. Accipitridae is the largest fami ...
, its closest living relative, from which it diverged around 2.4 million years ago. The
nomenclature Nomenclature (, ) is a system of names or terms, or the rules for forming these terms in a particular field of arts or sciences. (The theoretical field studying nomenclature is sometimes referred to as ''onymology'' or ''taxonymy'' ). The principl ...
and
taxonomy image:Hierarchical clustering diagram.png, 280px, Generalized scheme of taxonomy Taxonomy is a practice and science concerned with classification or categorization. Typically, there are two parts to it: the development of an underlying scheme o ...
of ''C. teauteensis'' are quite convoluted. As ''Circus eylesi'', it was only described in the mid-20th century, from remains found in Pyramid Valley in the South Island of New Zealand. However, essentially identical bones had been found about 100 years earlier by
Augustus Hamilton Augustus Hamilton (1 March 1853 – 12 October 1913) was a New Zealand ethnologist, biologist and museum director. He was born in Poole, Dorset, England on 1 March 1853. He wrote on the fishing and seafoods of the ancient Māori people. He also w ...
and discussed subsequently by
Henry Ogg Forbes Henry Ogg Forbes (30 January 1851 – 27 October 1932) was a Scottish explorer, ornithologist, and botanist. He also described a new species of spider, '' Thomisus decipiens''. Biography Forbes was the son of Rev Alexander Forbes M.A. (1821 ...
; this material probably comes from the Te Aute region in the North Island. Forbes gave it the names ''Circus hamiltoni'' and ''Circus teauteensis'', but since it seems no
holotype A holotype (Latin: ''holotypus'') is a single physical example (or illustration) of an organism used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of s ...
was ever formally named nor can be deduced from the minimal description; these names are generally considered ''
nomina nuda Nomen may refer to: *Nomen gentilicium, the middle part of Ancient Roman names ** ''Nomen est omen'', a Latin quote about nominative determinism *Nomen (ancient Egypt), the personal name of Ancient Egyptian pharaohs *Jaume Nomen (born 1960), Catal ...
''. ''C. teauteensis'' is considered valid, based on the argument that as with many New Zealand birds, it is reasonable to assume that North and South Island populations were at least distinct
subspecies In Taxonomy (biology), biological classification, subspecies (: subspecies) is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (Morphology (biology), morpholog ...
, if not
species A species () is often defined as the largest group of organisms in which any two individuals of the appropriate sexes or mating types can produce fertile offspring, typically by sexual reproduction. It is the basic unit of Taxonomy (biology), ...
. But even though
Kálmán Lambrecht Kálmán Lambrecht (1889–1936) was a Hungarian palaeontologist, best known for his work on fossil birds. He authored the “Handbuch der Palaeornithologie”, an exhaustive review of fossil birds published in 1933. Positions held include libra ...
mentioned a putative holotype
tibiotarsus The tibiotarsus is the large bone between the femur and the tarsometatarsus in the leg of a bird. It is the fusion of the proximal part of the tarsus with the tibia. A similar structure also occurred in the Mesozoic Heterodontosauridae. These ...
(which still exists in the BMNH) he explicitly stated that Forbes' names were both invalid. In addition, harrier bones of comparatively recent age in the collection of
Walter Mantell Walter Baldock Durrant Mantell (11 March 1820 – 7 September 1895) was a 19th-century New Zealand naturalist, politician, and land purchase commissioner. He was a founder and first secretary of the New Zealand Institute, and a collector of moa ...
, assigned to ''C. gouldi'' (an obsolete name of ''C. approximans'') by
Richard Lydekker Richard Lydekker (; 25 July 1849 – 16 April 1915) was a British naturalist, geologist and writer of numerous books on natural history. He was known for his contributions to zoology, paleontology, and biogeography. He worked extensively in cata ...
, seem to be of a more robust bird judging from the published descriptions; this material may now be lost, but all things considered it is not unlikely to be the first remains of Eyles' harrier known to science.Worthy, T. H. and R. N. Holdaway. (2002): ''The lost world of the Moa: Prehistoric Life of New Zealand''.
Indiana University Indiana University (IU) is a state university system, system of Public university, public universities in the U.S. state of Indiana. The system has two core campuses, five regional campuses, and two regional centers under the administration o ...
Press, Bloomington.


Extinction

The modern
swamp harrier The swamp harrier (''Circus approximans''), also known as the Australasian marsh harrier or Australasian harrier, is a large, slim bird of prey widely distributed across Australasia. In New Zealand, it is also known by the Māori name . It arriv ...
occurs all over New Zealand today. Even though the bones of its females can be mistaken for those of Eyles's harrier's males (as they differ little except in stoutness), all swamp harrier remains from the islands that have been studied in detail are some 1000 years old at most. That means that they post-date human settlement, i.e. there is no evidence for widespread coexistence of swamp and Eyles' harriers on New Zealand. Apparently, despite their considerable differences, the two harriers were still ecologically similar enough to competitively exclude one another, and only when the endemic Eyles' harrier became extinct could ''C. approximans'' become established.


References


External links


Wingspan Birds of Prey TrustZealand Birds OnlineForbes' Harrier. Circus teauteensis. From the series: Extinct Birds of New Zealand., Te Papa's collections Online (2006-0010-1/35)
{{Taxonbar, from=Q760860 Eyles's harrier Extinct birds of New Zealand Bird extinctions since 1500 Late Quaternary prehistoric birds Eyles's harrier