''Exbucklandia'' is a

genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...

flowering plant Flowering plants are plants that bear flowers and fruits, and form the clade Angiospermae (). The term angiosperm is derived from the Ancient Greek, Greek words (; 'container, vessel') and (; 'seed'), meaning that the seeds are enclosed with ...

s in the

family Family (from ) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). It forms the basis for social order. Ideally, families offer predictabili ...

Hamamelidaceae Hamamelidaceae, commonly referred to as the witch-hazel family, is a family of flowering plants in the order Saxifragales. The clade consists of shrubs and small trees positioned within the woody clade of the core Saxifragales. An earlier system ...

.Peter K. Endress. 1993. "Hamamelidaceae". pages 322-331. In: Klaus Kubitzki (editor); Jens G. Rohwer, and Volker Bittrich (volume editors). ''The Families and Genera of Vascular Plants'' volume II. Springer-Verlag: Berlin; Heidelberg, Germany. (Berlin) (New York) They are medium to large trees whose

natural range Species distribution, or species dispersion, is the manner in which a biological taxon is spatially arranged. The geographic limits of a particular taxon's distribution is its range, often represented as shaded areas on a map. Patterns of distr ...

is from eastern

India India, officially the Republic of India, is a country in South Asia. It is the List of countries and dependencies by area, seventh-largest country by area; the List of countries by population (United Nations), most populous country since ...

through southern

China China, officially the People's Republic of China (PRC), is a country in East Asia. With population of China, a population exceeding 1.4 billion, it is the list of countries by population (United Nations), second-most populous country after ...

and southward through the

Malay Peninsula The Malay Peninsula is located in Mainland Southeast Asia. The landmass runs approximately north–south, and at its terminus, it is the southernmost point of the Asian continental mainland. The area contains Peninsular Malaysia, Southern Tha ...

.Jingyu Wu, Bainian Sun, Yu-Sheng (Christopher) Liu, Sanping Xie, and Zhicheng Lin. 2009. "A new species of ''Exbucklandia'' (Hamamelidaceae) from the Pliocene of China and its paleoclimatic significance". ''Review of Palaeobotany and Palynology'' 155(1-2):32-41. In India and China, they are widely cultivated for their impressive foliage and valuable

lumber Lumber is wood that has been processed into uniform and useful sizes (dimensional lumber), including beams and planks or boards. Lumber is mainly used for construction framing, as well as finishing (floors, wall panels, window frames). ...

.Robert Scott Troupe. 1984. "Exbucklandia". pages 13-19. In: ''The Silviculture of Indian Trees'' volume V. (revised and enlarged by H.B. Joshi). Government of India Press. A few have been grown in the southernmost parts of the

United States The United States of America (USA), also known as the United States (U.S.) or America, is a country primarily located in North America. It is a federal republic of 50 U.S. state, states and a federal capital district, Washington, D.C. The 48 ...

.Arthur Lee Jacobson. 2009. "Plant of the Month: January 2009". At: Jacobson's Plant of the Month series. (see ''External links'' below). To speakers of English, ''Exbucklandia'' is generally known as the Pipli tree, from the

Bengal Bengal ( ) is a Historical geography, historical geographical, ethnolinguistic and cultural term referring to a region in the Eastern South Asia, eastern part of the Indian subcontinent at the apex of the Bay of Bengal. The region of Benga ...

i name for the species '' Exbucklandia populnea''.

Cultivation

Propagation Propagation can refer to: *Chain propagation in a chemical reaction mechanism *Crack propagation, the growth of a crack during the fracture of materials *Propaganda, non-objective information used to further an agenda *Reproduction, and other forms ...

is usually by seed, but cuttings have been successful under favorable conditions.

Plantation Plantations are farms specializing in cash crops, usually mainly planting a single crop, with perhaps ancillary areas for vegetables for eating and so on. Plantations, centered on a plantation house, grow crops including cotton, cannabis, tob ...

s of the young trees must be fenced against

cattle Cattle (''Bos taurus'') are large, domesticated, bovid ungulates widely kept as livestock. They are prominent modern members of the subfamily Bovinae and the most widespread species of the genus '' Bos''. Mature female cattle are calle ...

and

deer A deer (: deer) or true deer is a hoofed ruminant ungulate of the family Cervidae (informally the deer family). Cervidae is divided into subfamilies Cervinae (which includes, among others, muntjac, elk (wapiti), red deer, and fallow deer) ...

which eat whatever leaves they can reach. If ''Exbucklandia'' is grown in the open, the trunk forks and

branch A branch, also called a ramus in botany, is a stem that grows off from another stem, or when structures like veins in leaves are divided into smaller veins. History and etymology In Old English, there are numerous words for branch, includ ...

es are retained close to the ground. In the

forest A forest is an ecosystem characterized by a dense ecological community, community of trees. Hundreds of definitions of forest are used throughout the world, incorporating factors such as tree density, tree height, land use, legal standing, ...

, where it usually grows, the trunk is single, straight, and free of branches for 9 to 18 metres.

Sapling In botany, a tree is a perennial plant with an elongated stem, or trunk, usually supporting branches and leaves. In some usages, the definition of a tree may be narrower, e.g., including only woody plants with secondary growth, only pla ...

s grow slowly for the first few years, then more quickly later on.

Species

Only three species names have ever been published in ''Exbucklandia''. These are '' Exbucklandia populnea'', '' Exbucklandia tonkinensis'', and '' Exbucklandia longipetala''.The International Plant Names Index. "Exbucklandia". (see ''External links'' below). The

specific epithet In Taxonomy (biology), taxonomy, binomial nomenclature ("two-term naming system"), also called binary nomenclature, is a formal system of naming species of living things by giving each a name composed of two parts, both of which use Latin gramm ...

s refer to poplars,

Tonkin Tonkin, also spelled Tongkin, Tonquin or Tongking, is an exonym referring to the northern region of Vietnam. During the 17th and 18th centuries, this term referred to the domain '' Đàng Ngoài'' under Trịnh lords' control, including both the ...

, and long petals. ''Exbucklandia populnea'' is found throughout the entire

range Range may refer to: Geography * Range (geographic), a chain of hills or mountains; a somewhat linear, complex mountainous or hilly area (cordillera, sierra) ** Mountain range, a group of mountains bordered by lowlands * Range, a term used to i ...

of the genus. ''Exbucklandia tonkinensis'' is

native Native may refer to: People * '' Jus sanguinis'', nationality by blood * '' Jus soli'', nationality by location of birth * Indigenous peoples, peoples with a set of specific rights based on their historical ties to a particular territory ** Nat ...

to southeastern

Laos Laos, officially the Lao People's Democratic Republic (LPDR), is the only landlocked country in Southeast Asia. It is bordered by Myanmar and China to the northwest, Vietnam to the east, Cambodia to the southeast, and Thailand to the west and ...

, and northern

Vietnam Vietnam, officially the Socialist Republic of Vietnam (SRV), is a country at the eastern edge of mainland Southeast Asia, with an area of about and a population of over 100 million, making it the world's List of countries and depende ...

. ''Exbucklandia longipetala'' has a much more restricted range, being known only from the

Guangxi Guangxi,; officially the Guangxi Zhuang Autonomous Region, is an Autonomous regions of China, autonomous region of the China, People's Republic of China, located in South China and bordering Vietnam (Hà Giang Province, Hà Giang, Cao Bằn ...

and

Guizhou ) , image_skyline = , image_caption = , image_map = Guizhou in China (+all claims hatched).svg , mapsize = 275px , map_alt = Map showing the location of Guizhou Province , map_caption = Map s ...

provinces A province is an administrative division within a country or state. The term derives from the ancient Roman , which was the major territorial and administrative unit of the Roman Empire's territorial possessions outside Italy. The term ''provi ...

of China.Zhang Zhiyun, Zhang Hongda (Chang Hung-ta), and Peter K. Endress. 2003. "Hamamelidaceae" pages 18-42. In: Wu Zhengyi, Peter H. Raven, and Hong Deyuan (editors). ''Flora of China'' volume 9. Science Press: Beijing, China; Missouri Botanical Garden Press: St. Louis, Missouri, USA. Further

field work Field research, field studies, or fieldwork is the empirical research, collection of raw data outside a laboratory, library, or workplace setting. The approaches and methods used in field research vary across branches of science, disciplines. ...

will be needed to determine whether these are three distinct entities and whether the

specimen Specimen may refer to: Science and technology * Sample (material), a limited quantity of something which is intended to be similar to and represent a larger amount * Biological specimen or biospecimen, an organic specimen held by a biorepository f ...

s assigned to ''Exbucklandia populnea'' represent a single species or two. Consequently, estimates of the number of species range from two David J. Mabberley. 2008. ''Mabberley's Plant-Book''. Cambridge University Press: UK. to four.

Description

The following description of ''Exbucklandia'' applies only to

extant Extant or Least-concern species, least concern is the opposite of the word extinct. It may refer to: * Extant hereditary titles * Extant literature, surviving literature, such as ''Beowulf'', the oldest extant manuscript written in English * Exta ...

species and does not necessarily hold for species known only from the

fossil record A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserved ...

. It is adapted from the description in ''Flora of China'' volume 9, with the exceptions noted. Exbucklandias are

evergreen In botany, an evergreen is a plant which has Leaf, foliage that remains green and functional throughout the year. This contrasts with deciduous plants, which lose their foliage completely during the winter or dry season. Consisting of many diffe ...

tree In botany, a tree is a perennial plant with an elongated stem, or trunk, usually supporting branches and leaves. In some usages, the definition of a tree may be narrower, e.g., including only woody plants with secondary growth, only ...

s. ''Exbucklandia populnea'' and ''Exbucklandia tonkinensis'' are usually 16 to 20 meters in height, occasionally reaching 30 meters. The largest known individual of ''Exbucklandia populnea'' grew to 45 meters in the Darjeeling Hills of India. The usual height of ''Exbucklandia longipetala'' is not known. The

twigs A twig is a thin, often short, branch of a tree or bush. The buds on the twig are an important diagnostic characteristic, as are the abscission scars where the leaves have fallen away. The color, texture, and patterning of the twig bark are ...

have conspicuous nodes. The

leaves A leaf (: leaves) is a principal appendage of the stem of a vascular plant, usually borne laterally above ground and specialized for photosynthesis. Leaves are collectively called foliage, as in "autumn foliage", while the leaves, stem, ...

are attractively reddish when immature. They are arranged alternately on the stems, an arrangement unusual for

. The petioles are long and the leaves flutter in even a light breeze, like the leaves of poplars. The poplar petiole is flattened, and in

cross section Cross section may refer to: * Cross section (geometry) ** Cross-sectional views in architecture and engineering 3D *Cross section (geology) * Cross section (electronics) * Radar cross section, measure of detectability * Cross section (physics) **A ...

, it is long in the vertical direction. Whether ''Exbucklandia'' has the same sort of petiole has not been recorded. The leaf

blade A blade is the Sharpness (cutting), sharp, cutting portion of a tool, weapon, or machine, specifically designed to puncture, chop, slice, or scrape surfaces or materials. Blades are typically made from materials that are harder than those they a ...

simple Simple or SIMPLE may refer to: *Simplicity, the state or quality of being simple Arts and entertainment * ''Simple'' (album), by Andy Yorke, 2008, and its title track * "Simple" (Florida Georgia Line song), 2018 * "Simple", a song by John ...

, and sometimes has three pointed lobes, or rarely, five. It is thickly leathery and its

margin Margin may refer to: Physical or graphical edges *Margin (typography), the white space that surrounds the content of a page * Continental margin, the zone of the ocean floor that separates the thin oceanic crust from thick continental crust *Leaf ...

entire Entire may refer to: * Entire function, a function that is holomorphic on the whole complex plane * Entire (animal), an indication that an animal is not neutered * Entire (botany) This glossary of botanical terms is a list of definitions o ...

. The

venation Venation may refer to: * Venation (botany), the arrangement of veins in leaves * Wing venation, the arrangement of veins in insect wings See also * * Vernation Vernation or leafing is the formation of new leaves or fronds. In plant anatomy, it ...

is palmate, with the secondary veins radiating from the

apex The apex is the highest point of something. The word may also refer to: Arts and media Fictional entities * Apex (comics) A-Bomb Abomination Absorbing Man Abraxas Abyss Abyss is the name of two characters appearing in Ameri ...

of the petiole. The

stipule In botany, a stipule is an outgrowth typically borne on both sides (sometimes on just one side) of the base of a leafstalk (the petiole (botany), petiole). They are primarily found among dicots and rare among monocots. Stipules are considered part ...

s are large and

coherent Coherence is, in general, a state or situation in which all the parts or ideas fit together well so that they form a united whole. More specifically, coherence, coherency, or coherent may refer to the following: Physics * Coherence (physics ...

; soon falling away. Each

inflorescence In botany, an inflorescence is a group or cluster of flowers arranged on a plant's Plant stem, stem that is composed of a main branch or a system of branches. An inflorescence is categorized on the basis of the arrangement of flowers on a mai ...

has 7 to 16

flower Flowers, also known as blooms and blossoms, are the reproductive structures of flowering plants ( angiosperms). Typically, they are structured in four circular levels, called whorls, around the end of a stalk. These whorls include: calyx, m ...

s and is located in the

axil A leaf (: leaves) is a principal appendage of the stem of a vascular plant, usually borne laterally above ground and specialized for photosynthesis. Leaves are collectively called foliage, as in "autumn foliage", while the leaves, stem, fl ...

of a leaf. The flowers are small and bisexual.

Sepal A sepal () is a part of the flower of angiosperms (flowering plants). Usually green, sepals typically function as protection for the flower in bud, and often as support for the petals when in bloom., p. 106 Etymology The term ''sepalum'' ...

s are completely lacking.

Petal Petals are modified leaves that form an inner whorl surrounding the reproductive parts of flowers. They are often brightly coloured or unusually shaped to attract pollinators. All of the petals of a flower are collectively known as the ''corol ...

s are often absent, but are small and white when present. Some authors have interpreted what appear to be petals as petaloid

staminode In botany, a staminode is an often rudimentary, sterile or abortive stamen, which means that it does not produce pollen.Jackson, Benjamin, Daydon; ''A Glossary of Botanic Terms with their Derivation and Accent''; Published by Gerald Duckworth & Co ...

s. The

stamen The stamen (: stamina or stamens) is a part consisting of the male reproductive organs of a flower. Collectively, the stamens form the androecium., p. 10 Morphology and terminology A stamen typically consists of a stalk called the filament ...

s are 10 to 15 in number. The

anther The stamen (: stamina or stamens) is a part consisting of the male reproductive organs of a flower. Collectively, the stamens form the androecium., p. 10 Morphology and terminology A stamen typically consists of a stalk called the filament ...

s are basifixed, as in all of Hamamelidaceae. Each

theca In biology, a theca (: thecae) is a sheath or a covering. Botany In botany, the theca is related to plant's flower anatomy. The theca of an angiosperm consists of a pair of microsporangia that are adjacent to each other and share a common ar ...

has one

sporangium A sporangium (from Late Latin, ; : sporangia) is an enclosure in which spores are formed. It can be composed of a unicellular organism, single cell or can be multicellular organism, multicellular. Virtually all plants, fungus, fungi, and many ot ...

, whereas for most

angiosperm Flowering plants are plants that bear flowers and fruits, and form the clade Angiospermae (). The term angiosperm is derived from the Greek words (; 'container, vessel') and (; 'seed'), meaning that the seeds are enclosed within a fruit ...

s, there are two. The thecae

open Open or OPEN may refer to: Music * Open (band), Australian pop/rock band * The Open (band), English indie rock band * ''Open'' (Blues Image album), 1969 * ''Open'' (Gerd Dudek, Buschi Niebergall, and Edward Vesala album), 1979 * ''Open'' (Go ...

by one

valve A valve is a device or natural object that regulates, directs or controls the flow of a fluid (gases, liquids, fluidized solids, or Slurry, slurries) by opening, closing, or partially obstructing various passageways. Valves are technically Pip ...

. The

ovary The ovary () is a gonad in the female reproductive system that produces ova; when released, an ovum travels through the fallopian tube/ oviduct into the uterus. There is an ovary on the left and the right side of the body. The ovaries are end ...

is half- inferior and, as in the rest of the family, consists of two

carpel Gynoecium (; ; : gynoecia) is most commonly used as a collective term for the parts of a flower that produce ovules and ultimately develop into the fruit and seeds. The gynoecium is the innermost whorl of a flower; it consists of (one or more ...

s. The number of

ovule In seed plants, the ovule is the structure that gives rise to and contains the female reproductive cells. It consists of three parts: the ''integument'', forming its outer layer, the ''nucellus'' (or remnant of the sporangium, megasporangium), ...

s in each carpel has been reported as five or six and as six to eight. What some authors have loosely called styles are actually styluli. These are separate by definition, channeling the

pollen tube A pollen tube is a tubular structure produced by the male gametophyte of seed plants when it germinates. Pollen tube elongation is an integral stage in the plant life cycle. The pollen tube acts as a conduit to transport the male gamete cells fr ...

s that invade them into only one carpel. Each stigma is somewhat

decurrent ''Decurrent'' (sometimes decurring) is a term used in botany and mycology to describe plant or fungal parts that extend downward. In botany, the term is most often applied to leaf blades that partly wrap or have wings around the stem or petio ...

down one side of its stylulus. The

fruit In botany, a fruit is the seed-bearing structure in flowering plants (angiosperms) that is formed from the ovary after flowering. Fruits are the means by which angiosperms disseminate their seeds. Edible fruits in particular have long propaga ...

is a 4-valved capsule. The two

locule A locule (: locules) or loculus (; : loculi) is a small cavity or compartment within an organ or part of an organism (animal, plant, or fungus). In angiosperms (flowering plants), the term ''locule'' usually refers to a chamber within an ovary ...

s separate, and each splits into two valves. Each locule contains five to seven seeds. The upper four or five are sterile and wingless. The lowest one or two are fertile and narrowly winged. The seeds are light and can travel far in a strong wind.

Affinities

Exbucklandia oregonensis as Ficus washingtonensis USNM P36980 img3

The

phylogeny A phylogenetic tree or phylogeny is a graphical representation which shows the evolutionary history between a set of species or Taxon, taxa during a specific time.Felsenstein J. (2004). ''Inferring Phylogenies'' Sinauer Associates: Sunderland, M ...

of Hamamelidaceae has not been resolved with much certainty, but in one recent

molecular phylogenetic Molecular phylogenetics () is the branch of phylogeny that analyzes genetic, hereditary molecular differences, predominantly in DNA sequences, to gain information on an organism's evolutionary relationships. From these analyses, it is possible to ...

study, ''Exbucklandia'' and '' Rhodoleia'' formed the most basal

clade In biology, a clade (), also known as a Monophyly, monophyletic group or natural group, is a group of organisms that is composed of a common ancestor and all of its descendants. Clades are the fundamental unit of cladistics, a modern approach t ...

in the family.Susana Magallón. 2007. "From Fossils to Molecules: Phylogeny and the Core Eudicot Floral Groundplan in Hamamelidoideae (Hamamelidaceae, Saxifragales)". ''Systematic Botany'' 32(2):317-347. '' Chunia bucklandioides'', a rare tree from

Hainan Hainan is an island provinces of China, province and the southernmost province of China. It consists of the eponymous Hainan Island and various smaller islands in the South China Sea under the province's administration. The name literally mean ...

which has never been sampled for

DNA Deoxyribonucleic acid (; DNA) is a polymer composed of two polynucleotide chains that coil around each other to form a double helix. The polymer carries genetic instructions for the development, functioning, growth and reproduction of al ...

, might also be a member of this clade. Vegetatively, it is hard to distinguish from ''Exbucklandia'', while florally, it is intermediate between ''Exbucklandia'' and '' Mytilaria''.Hung-Ta Chang (Hongda Zhang). 1948. "Chunia". page 63. In: "Additions to the Hamamelidaceous flora of China". ''Sunyatsenia'' (1-2):63-74. The

morphology Morphology, from the Greek and meaning "study of shape", may refer to: Disciplines *Morphology (archaeology), study of the shapes or forms of artifacts *Morphology (astronomy), study of the shape of astronomical objects such as nebulae, galaxies, ...

of ''Chunia'' indicates that if it is not in the clade with ''Exbucklandia'' and ''Rhodoleia'', then it either forms its own clade or it is

sister A sister is a woman or a girl who shares parents or a parent with another individual; a female sibling. The male counterpart is a brother. Although the term typically refers to a familial relationship, it is sometimes used endearingly to ref ...

to ''Mytilaria''.

Evolution

''Exbucklandia'' was once more widely

distributed Distribution may refer to: Mathematics *Distribution (mathematics), generalized functions used to formulate solutions of partial differential equations *Probability distribution, the probability of a particular value or value range of a varia ...

than it is today. Four species are known only from

fossil A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserve ...

s. '' Exbucklandia oregonensis'' grew in the northwestern

during the

Oligocene The Oligocene ( ) is a geologic epoch (geology), epoch of the Paleogene Geologic time scale, Period that extends from about 33.9 million to 23 million years before the present ( to ). As with other older geologic periods, the rock beds that defin ...

and

Miocene The Miocene ( ) is the first epoch (geology), geological epoch of the Neogene Period and extends from about (Ma). The Miocene was named by Scottish geologist Charles Lyell; the name comes from the Greek words (', "less") and (', "new") and mea ...

epochs In chronology and periodization, an epoch or reference epoch is an instant in time chosen as the origin of a particular calendar era. The "epoch" serves as a reference point from which time is measured. The moment of epoch is usually decided by ...

.Herb Meyer. 1973. "The Oligocene Lyons Flora of northwestern Oregon". ''The Ore Bin'' 35(3):37-51. (see ''External links'' below).Kathleen B. Pigg and Wesley C. Wehr. 2002. "Tertiary Flowers, Fruits, and Seeds of Washington State and Adjacent Areas - Part III". ''Washington Geology'' 30(3-4):3-20. (see ''External links'' below). ''

Exbucklandia microdictya ''Exbucklandia'' is a genus of flowering plants in the family Hamamelidaceae.Peter K. Endress. 1993. "Hamamelidaceae". pages 322-331. In: Klaus Kubitzki (editor); Jens G. Rohwer, and Volker Bittrich (volume editors). ''The Families and Genera of ...

'' is known from

Paleocene The Paleocene ( ), or Palaeocene, is a geological epoch (geology), epoch that lasted from about 66 to 56 mya (unit), million years ago (mya). It is the first epoch of the Paleogene Period (geology), Period in the modern Cenozoic Era (geology), ...

deposits near

Altay City Altay or Aletai is a county-level city in Altay Prefecture within Ili Kazakh Autonomous Prefecture, in far Northern Xinjiang, China. The city centre is located on the banks of Kelan River. Administrative divisions Altay City is divided into ...

Xinjiang Xinjiang,; , SASM/GNC romanization, SASM/GNC: Chinese postal romanization, previously romanized as Sinkiang, officially the Xinjiang Uygur Autonomous Region (XUAR), is an Autonomous regions of China, autonomous region of the China, People' ...

, China. '' Exbucklandia miocenica'' is a Miocene species from

Yunnan Yunnan; is an inland Provinces of China, province in Southwestern China. The province spans approximately and has a population of 47.2 million (as of 2020). The capital of the province is Kunming. The province borders the Chinese provinces ...

, China. '' Exbucklandia tengchongensis'' is known from fossils recovered from a

diatomite Diatomaceous earth ( ), also known as diatomite ( ), celite, or kieselguhr, is a naturally occurring, soft, siliceous sedimentary rock that can be crumbled into a fine white to off-white powder. It has a particle size ranging from more than 3 ...

mine in

Tengchong County Tengchong () is a county-level city of Baoshan City, western Yunnan province, People's Republic of China. It is well known for its volcanic activity. The city is named after the town of Tengchong which serves as its political center, previously k ...

in Yunnan, China. It dates from the

Pliocene The Pliocene ( ; also Pleiocene) is the epoch (geology), epoch in the geologic time scale that extends from 5.33 to 2.58cycad Cycads are seed plants that typically have a stout and woody (ligneous) trunk (botany), trunk with a crown (botany), crown of large, hard, stiff, evergreen and (usually) pinnate leaves. The species are dioecious, that is, individual plants o ...

in the "Tentamen" part of ''Flora der Vorwelt,'' the

classic A classic is an outstanding example of a particular style; something of Masterpiece, lasting worth or with a timeless quality; of the first or Literary merit, highest quality, class, or rank – something that Exemplification, exemplifies its ...

paleobotanical work by

Kaspar Maria von Sternberg Count Kaspar Maria von Sternberg (also: ''Caspar Maria'', ''Count Sternberg'', , ; 6 January 1761 – 20 December 1838) was a Bohemian aristocrat, theologian, mineralogist, geognost, entomologist and botanist. He is known as the "Father of Paleo ...

. The name has been attributed to

Carl Borivoj Presl Carl Borivoj Presl (; 17 February 1794 – 2 October 1852) was a Czech botanist. Biography Presl lived his entire life in Prague, and was a professor of botany at the University of Prague (1833–52). In 1836,

Robert Brown Robert Brown may refer to: Robert Brown (born 1965), British Director, Animator and author Entertainers and artists * Washboard Sam or Robert Brown (1910–1966), American musician and singer * Robert W. Brown (1917–2009), American printmaker ...

, unaware that the name had already been taken, published the name ''Bucklandia populnea'' for the living species now known as ''Exbucklandia populnea'' Robert Brown. 1836. ''Asiatic Researches'' 19(1):95. to honor the

Reverend The Reverend (abbreviated as The Revd, The Rev'd or The Rev) is an honorific style (form of address), style given to certain (primarily Western Christian, Western) Christian clergy and Christian minister, ministers. There are sometimes differen ...

William Buckland William Buckland Doctor of Divinity, DD, Royal Society, FRS (12 March 1784 – 14 August 1856) was an English theologian, geologist and paleontology, palaeontologist. His work in the early 1820s proved that Kirkdale Cave in North Yorkshire h ...

, an English

geologist A geologist is a scientist who studies the structure, composition, and History of Earth, history of Earth. Geologists incorporate techniques from physics, chemistry, biology, mathematics, and geography to perform research in the Field research, ...

and

paleontologist Paleontology, also spelled as palaeontology or palæontology, is the scientific study of the life of the past, mainly but not exclusively through the study of fossils. Paleontologists use fossils as a means to classify organisms, measure geolo ...

.Umberto Quattrocchi. 2000. ''CRC World Dictionary of Plant Names'' volume II. CRC Press: Boca Raton; New York; Washington,DC;, USA. London, UK. (vol. II). (see ''External links'' below). In the case of ''Bucklandia'', Brown did not meet the requirements of

valid publication In botanical nomenclature, a validly published name is a name that meets the requirements in the ''International Code of Nomenclature for algae, fungi, and plants'' (''ICN'') for valid publication. Valid publication of a name represents the minim ...

, but the name was validated by William Griffith, posthumously, in 1847.William Griffith. 1847. ''Journals of Travels in Assam Burma Bootan Affghanistan and the Neighbouring Countries'' (sic). Bishop's College Press: Calcutta. (reprinted 2001. Munshiram Manoharlal Publishers, New Delhi). In 1924,

Paul Henri Lecomte Paul Henri Lecomte (8 January 1856, in Saint-Nabord, Vosges – 12 June 1934, in Paris) was a French botanist. In 1884, after attaining a number of degrees, Lecomte became a professor at Lycée Saint-Louis in Paris. In addition to his teaching dut ...

named a second species, ''Bucklandia tonkinensis''.Paul Henri Lecomte. 1924. ''Bulletin du Muséum d'Histoire Naturelle. Paris''. 30:392. For more than 100 years, it remained unnoticed that two genera of plants had received the name ''Bucklandia''. This conflict was finally resolved by Roland W. Brown in 1946. At that time, Brown wrote, "Inasmuch as the 1825 cycadeoid name has priority, the witchhazel genus requires a new name. For this purpose, I propose ''Exbucklandia'', the derivation of which is obvious." Unaware of Brown's paper, Cornelis G.G.J. van Steenis sought to resolve the conflict of names in 1952 by replacing ''Bucklandia'' with ''Symingtonia''.Cornelis G.G.J. van Steenis. 1952. ''Acta Botanica Neerlandica'' 1:443-444. He published a notice of this oversight in 1954, acknowledging the priority of ''Exbucklandia'' over ''Symingtonia''.Cornelis G.G.J. van Steenis. 1954. page 595. In: "Miscellaneous Botanical Notes VI". ''Blumea'' 7(3):595-598. In that paper, van Steenis transferred the second species to ''Exbucklandia'', because Roland Brown had not explicitly done so in 1946. Thus van Steenis originated the

combination In mathematics, a combination is a selection of items from a set that has distinct members, such that the order of selection does not matter (unlike permutations). For example, given three fruits, say an apple, an orange and a pear, there are ...

''Exbucklandia tonkinensis'', but not validly, because he failed to

cite To cite is to quote or mention a source. CITE or Cite may refer to: * Cite (cycling team), Italy * Cite (magazine), ''Cite'' (magazine), an American architecture quarterly * CITE-FM, a Canadian radio station * Center for Innovation Testing and Ev ...

the publication of the

basionym In the scientific name of organisms, basionym or basyonym means the original name on which a new name is based; the author citation of the new name should include the authors of the basionym in parentheses. The term "basionym" is used in both botan ...

as the

ICBN The ''International Code of Nomenclature for algae, fungi, and plants'' (ICN or ICNafp) is the set of rules and recommendations dealing with the formal botanical names that are given to plants, fungi and a few other groups of organisms, all tho ...

has required since 1953. In 1959, Hung-Ta Chang validly made the combination ''Exbucklandia tonkinensis''.Hung-Ta Chang. 1959. ''Acta Scientiarum Naturalium Universitatis Sunyatseni''. 1959(2). In that same paper, Chang described and named the third species, ''Exbucklandia longipetala''. A taxonomic history and a description of each of the three species can be found in ''Flora of China'' volume 9.

Cultivation

Species

Description

Affinities

Evolution

References

External links