Eutrema Penlandii
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''Eutrema penlandii'' is a rare species of flowering plant in the
mustard family Brassicaceae () or (the older but equally valid) Cruciferae () is a medium-sized and economically important family of flowering plants commonly known as the mustards, the crucifers, or the cabbage family. Most are herbaceous plants, while some ar ...
known by the common names Mosquito Range mustard, Penland's alpine fen mustard and Penland's eutrema. It is
endemic Endemism is the state of a species being found only in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also foun ...
to
Colorado Colorado is a U.S. state, state in the Western United States. It is one of the Mountain states, sharing the Four Corners region with Arizona, New Mexico, and Utah. It is also bordered by Wyoming to the north, Nebraska to the northeast, Kansas ...
in the United States, where it is known only from a small strip of unique mountain habitat measuring 40 kilometers long. It is threatened by
mining Mining is the Resource extraction, extraction of valuable geological materials and minerals from the surface of the Earth. Mining is required to obtain most materials that cannot be grown through agriculture, agricultural processes, or feasib ...
and associated environmental changes in this part of the
Continental Divide A continental divide is a drainage divide on a continent such that the drainage basin on one side of the divide feeds into one ocean or sea, and the basin on the other side either feeds into a different ocean or sea, or else is endorheic, not ...
in central Colorado. The plant was federally listed as a threatened species of the US in 1993.USFWS
The plant ''Eutrema penlandii'' (Penland alpine fen mustard) determined to be a threatened species.
''Federal Register'' July 28, 1993.


Distribution

This plant is the only '' Eutrema'' in the lower 48 states and only one of two that occur in North America; its nearest close relative is '' Eutrema edwardii'', a plant of the
Arctic The Arctic (; . ) is the polar regions of Earth, polar region of Earth that surrounds the North Pole, lying within the Arctic Circle. The Arctic region, from the IERS Reference Meridian travelling east, consists of parts of northern Norway ( ...
. ''E. penlandii'' is a small
alpine climate Alpine climate is the typical climate for elevations above the tree line, where trees fail to grow due to cold. This climate is also referred to as a mountain climate or highland climate. Definition There are multiple definitions of alpine cli ...
species, possibly a
relict A relict is a surviving remnant of a natural phenomenon. Biology A relict (or relic) is an organism that at an earlier time was abundant in a large area but now occurs at only one or a few small areas. Geology and geomorphology In geology, a r ...
from the last
ice age An ice age is a long period of reduction in the temperature of Earth's surface and atmosphere, resulting in the presence or expansion of continental and polar ice sheets and alpine glaciers. Earth's climate alternates between ice ages, and g ...
which persists in the high, cold
fen A fen is a type of peat-accumulating wetland fed by mineral-rich ground or surface water. It is one of the main types of wetland along with marshes, swamps, and bogs. Bogs and fens, both peat-forming ecosystems, are also known as mires ...
s of the
Rocky Mountains The Rocky Mountains, also known as the Rockies, are a major mountain range and the largest mountain system in North America. The Rocky Mountains stretch in great-circle distance, straight-line distance from the northernmost part of Western Can ...
. Today the plant only occurs in the
Mosquito Range The Mosquito Range (elevation approximately 14,000 ft) is a high mountain range in the Rocky Mountains of central Colorado in the United States. The peaks of the range form a ridge running north–south for roughly 40 mi (64 km) fr ...
, a ridge of mountains in the Rockies.Mohlenbrock, R. H
This land: a guide to central national forests.
University of California Press The University of California Press, otherwise known as UC Press, is a publishing house associated with the University of California that engages in academic publishing. It was founded in 1893 to publish scholarly and scientific works by faculty ...
. 2006. page 47.
It was first collected by the botanist C. William Penland in 1935 on Hoosier Ridge in the Mosquito Range, and it is one of several rare alpine plants native to the area, such as the globe gilia (''Gilia globularis'').


Description

This plant is a perennial herb growing about six inches tall and bearing clusters of cream-white flowers in the summer. The alpine habitat of the plant is a wet
tundra In physical geography, a tundra () is a type of biome where tree growth is hindered by frigid temperatures and short growing seasons. There are three regions and associated types of tundra: #Arctic, Arctic, Alpine tundra, Alpine, and #Antarctic ...
kept constantly fed with ample water by the surrounding
snowfield A snow field, snowfield or neve is an accumulation of permanent snow and ice, typically found above the snow line, normally in mountainous and glacial terrain. Glaciers originate in snowfields. The lower end of a glacier is usually free from ...
s. Wind melts the snow which runs into the tundra and turns it to a fen, a
wetland A wetland is a distinct semi-aquatic ecosystem whose groundcovers are flooded or saturated in water, either permanently, for years or decades, or only seasonally. Flooding results in oxygen-poor ( anoxic) processes taking place, especially ...
on
limestone Limestone is a type of carbonate rock, carbonate sedimentary rock which is the main source of the material Lime (material), lime. It is composed mostly of the minerals calcite and aragonite, which are different Polymorphism (materials science) ...
substrates. The plant grows in rocky nooks that shelter it from the wind and keep the
moss Mosses are small, non-vascular plant, non-vascular flowerless plants in the taxonomic phylum, division Bryophyta (, ) ''sensu stricto''. Bryophyta (''sensu lato'', Wilhelm Philippe Schimper, Schimp. 1879) may also refer to the parent group bryo ...
y
peat Peat is an accumulation of partially Decomposition, decayed vegetation or organic matter. It is unique to natural areas called peatlands, bogs, mires, Moorland, moors, or muskegs. ''Sphagnum'' moss, also called peat moss, is one of the most ...
soil saturated, but not flooded. The
elevation The elevation of a geographic location (geography), ''location'' is its height above or below a fixed reference point, most commonly a reference geoid, a mathematical model of the Earth's sea level as an equipotential gravitational equipotenti ...
is at least 3700 meters. The habitat is within a section of the Continental Divide that trends east-west in opposition to the general north–south direction of most of the ranges, and the plants grow on south- and east-facing slopes. There are about eleven occurrences, with populations at Hoosier Ridge, Mount Silverheels, the Mosquito Pass, the Hilltop Mine,
Mount Blue Sky Mount Blue Sky (formerly Mount Evans) is the highest peak in the Mount Evans Wilderness in the Front Range of the Rocky Mountains of North America. The prominent fourteener is located southwest by south ( bearing 214°) of Idaho Springs in ...
, and Loveland Mountain. The total area occupied by the plant is probably around .


Status


Threats

Mining Mining is the Resource extraction, extraction of valuable geological materials and minerals from the surface of the Earth. Mining is required to obtain most materials that cannot be grown through agriculture, agricultural processes, or feasib ...
is a common industry in the high mountain peaks occupied by the plant, and mining companies and stakeholders did not agree with federal protection plans for the species and its habitat. Active claims for
gold Gold is a chemical element; it has chemical symbol Au (from Latin ) and atomic number 79. In its pure form, it is a brightness, bright, slightly orange-yellow, dense, soft, malleable, and ductile metal. Chemically, gold is a transition metal ...
and trace metals are still in place, though mining activity is currently low because of the status of the market. Activities associated with mining, such as
ditch A ditch is a small to moderate trench created to channel water. A ditch can be used for drainage, to drain water from low-lying areas, alongside roadways or fields, or to channel water from a more distant source for plant irrigation. Ditches ...
ing, diking, and trail maintenance, alter the
hydrology Hydrology () is the scientific study of the movement, distribution, and management of water on Earth and other planets, including the water cycle, water resources, and drainage basin sustainability. A practitioner of hydrology is called a hydro ...
of the landscape, and this is a major threat to the habitat that the plant depends on. There is a specific water balance maintained by the
snowmelt In hydrology, snowmelt is surface runoff produced from melting snow. It can also be used to describe the period or season during which such runoff is produced. Water produced by snowmelt is an important part of the annual water cycle in many part ...
from the high ridges which keeps the soil quite wet but not flooded, and changing the path of flowing water could disrupt it. The plant almost always grows within half a meter of actively flowing meltwater. Mining activity could also lower the pH of the normally
alkaline soils Alkali, or alkaline, soils are clay soils with high pH (greater than 8.5), a poor soil structure and a low infiltration capacity. Often they have a hard calcareous layer at 0.5 to 1 metre depth. Alkali soils owe their unfavorable physico ...
. Other threats include recreational activity including
hiking A hike is a long, vigorous walk, usually on trails or footpaths in the countryside. Walking for pleasure developed in Europe during the eighteenth century. Long hikes as part of a religious pilgrimage have existed for a much longer time. "Hi ...
,
mountain biking Mountain biking (MTB) is a sport of riding bicycles off-road, often over rough terrain, usually using specially designed mountain bikes. Mountain bikes share similarities with other bikes but incorporate features designed to enhance durability ...
, and the use of
off-road vehicle An off-road vehicle (ORV), also known as an off-highway vehicle (OHV), overland vehicle or adventure vehicle, is a type of transportation specifically engineered to navigate unpaved roads and surfaces. These include trails, forest roads, and ...
s, horses, and
burro The donkey or ass is a domesticated equine. It derives from the African wild ass, ''Equus africanus'', and may be classified either as a subspecies thereof, ''Equus africanus asinus'', or as a separate species, ''Equus asinus''. It was domes ...
s.


References


External links


USDA Plants ProfileCenter for Plant Conservation
{{Taxonbar, from=Q5414565 penlandii Flora of Colorado Alpine flora