The European tree frog (''Hyla arborea'') is a small tree
frog
A frog is any member of a diverse and largely semiaquatic group of short-bodied, tailless amphibian vertebrates composing the order (biology), order Anura (coming from the Ancient Greek , literally 'without tail'). Frog species with rough ski ...
. As traditionally defined, it was found throughout much of Europe, Asia and northern Africa,
but based on molecular genetic and other data several populations formerly included in it are now recognized as separate species (for example, ''
H. intermedia'' of Italy and nearby, ''
H. molleri'' of the Iberian Peninsula, ''
H. meridionalis'' of parts of southwestern Europe and northern Africa, and ''
H. orientalis'' of parts of Eastern Europe, Turkey and the Black Sea and Caspian Sea regions), limiting the true European tree frog to Europe from France to Poland and Greece.
Description

European tree frogs are small; males range from in length, and females range from in length. They are slender, with long legs.
Their
dorsal
Dorsal (from Latin ''dorsum'' ‘back’) may refer to:
* Dorsal (anatomy), an anatomical term of location referring to the back or upper side of an organism or parts of an organism
* Dorsal, positioned on top of an aircraft's fuselage
The fus ...
skin is smooth, while their ventral skin is granular. Their dorsal skin can be green, gray, or tan depending on the temperature, humidity, or their mood. Their ventral skin is a whitish color, and the dorsal and ventral skin is separated by a dark brown
lateral
Lateral is a geometric term of location which may also refer to:
Biology and healthcare
* Lateral (anatomy), a term of location meaning "towards the side"
* Lateral cricoarytenoid muscle, an intrinsic muscle of the larynx
* Lateral release ( ...
stripe from the eyes to the groin. Females have white throats, while males have golden brown throats
with large (folded)
vocal sacs. The head of ''H. arborea'' is rounded, the lip drops strongly, the
pupil
The pupil is a hole located in the center of the iris of the eye that allows light to strike the retina.Cassin, B. and Solomon, S. (1990) ''Dictionary of Eye Terminology''. Gainesville, Florida: Triad Publishing Company. It appears black becau ...
has the shape of a horizontal
ellipse
In mathematics, an ellipse is a plane curve surrounding two focus (geometry), focal points, such that for all points on the curve, the sum of the two distances to the focal points is a constant. It generalizes a circle, which is the special ty ...
, and the
tympanum is clearly recognizable. The discs on the frog's toes, which it uses to climb trees and hedges, is a characteristic feature of ''H. arborea'' . Like other frogs, their hind legs are much larger and stronger than the fore legs, enabling the frogs to jump rapidly.
Distribution and habitat
Members of the ''H. arborea''
species complex
In biology, a species complex is a group of closely related organisms that are so similar in appearance and other features that the boundaries between them are often unclear. The taxa in the complex may be able to hybridize readily with each oth ...
are the only representatives of the widespread
tree frog
A tree frog (or treefrog) is any species of frog that spends a major portion of its lifespan in trees, known as an arboreal state. Several lineages of frogs among the Neobatrachia suborder have given rise to treefrogs, although they are not clos ...
family (Hylidae)
indigenous to
mainland Europe
Continental Europe or mainland Europe is the contiguous mainland of Europe, excluding its surrounding islands. It can also be referred to ambiguously as the European continent, – which can conversely mean the whole of Europe – and, by so ...
.
and are found across most of Europe (except Ireland),
[ northwest Africa, and temperate Asia to Japan.] This species complex is native to these countries:
Albania; Armenia; Austria; Azerbaijan; Belarus; Belgium; Bosnia and Herzegovina; Bulgaria; Croatia; Cyprus; the Czech Republic; Denmark; France; Georgia; Germany; Greece; Hungary; Israel (found in the Ayalon Valley); Italy; Latvia; Liechtenstein; Lithuania; Luxembourg; Macedonia, the Republic of; Moldova; Montenegro; the Netherlands; Poland; Portugal; Romania; the Russian Federation; Serbia; Slovakia; Slovenia; Sweden; Switzerland; Turkey; Ukraine.
In the United Kingdom, the species status is contested. It has been introduced, but at least one British population, now thought to be extinct, may have been native. Historical evidence suggests that the species could have possibly perished due to over-collection for medicinal use. Celtic Rewilding have proposed a reintroduction to the UK. It has been reintroduced to Latvia (Courland
Courland is one of the Historical Latvian Lands in western Latvia. Courland's largest city is Liepāja, which is the third largest city in Latvia. The regions of Semigallia and Selonia are sometimes considered as part of Courland as they were ...
).
European tree frogs can be found in marshlands, damp meadows, reed beds, parks, gardens, vineyards, orchards, stream banks, lake shores, or humid or dry forests. They tend to avoid dark or thick forests, and they are able to tolerate some periods of dryness; therefore, sometimes they are found in dry habitat
In ecology, habitat refers to the array of resources, biotic factors that are present in an area, such as to support the survival and reproduction of a particular species. A species' habitat can be seen as the physical manifestation of its ...
s.
Behavior
*Historically, tree frogs were used as barometer
A barometer is a scientific instrument that is used to measure air pressure in a certain environment. Pressure tendency can forecast short term changes in the weather. Many measurements of air pressure are used within surface weather analysis ...
s because they respond to approaching rain by croaking.
*Depending on subspecies
In Taxonomy (biology), biological classification, subspecies (: subspecies) is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (Morphology (biology), morpholog ...
, temperature, humidity, and the frog's 'mood', skin colour ranges from bright to olive green, grey, brown and yellow.
*European tree frogs eat a variety of small arthropod
Arthropods ( ) are invertebrates in the phylum Arthropoda. They possess an arthropod exoskeleton, exoskeleton with a cuticle made of chitin, often Mineralization (biology), mineralised with calcium carbonate, a body with differentiated (Metam ...
s, such as spiders, flies, beetles, butterflies, and smooth caterpillars. Their ability to take long leaps allows them to catch fast-flying insects, which make up most of their diets.
*They hibernate
Hibernation is a state of minimal activity and metabolic reduction entered by some animal species. Hibernation is a seasonal heterothermy characterized by low body-temperature, slow breathing and heart-rate, and low metabolic rate. It is most ...
in walls, cellars, under rocks, under clumps of vegetation, or buried in leaf piles or manure piles.[Wells, K. D. ‘‘The Ecology of Behavior of Amphibians.’’ The University of Chicago, 2007.]
Reproduction
European tree frogs reproduce in stagnant bodies of water, such as lakes, ponds, swamps, reservoirs, and sometimes puddles, from late March to June. They croak in the breeding season, even when migrating to their mating pools or ponds. Males will often change breeding ponds, even within the same breeding season.[Lardner, B. (2000). Morphological and life history responses to predators in larvae of seven anurans. ''Oikos'', 88(1): 169–180.] After a spring rain, the males will call females from low vegetation or shallow ponds. About 800 to 1000 eggs are laid in clumps the size of a walnut. Individual eggs are about 1.5 mm in diameter. After 10–14 days, the eggs hatch. Then, after three months, tadpole
A tadpole or polliwog (also spelled pollywog) is the Larva, larval stage in the biological life cycle of an amphibian. Most tadpoles are fully Aquatic animal, aquatic, though some species of amphibians have tadpoles that are terrestrial animal, ...
s metamorphose into frogs. Metamorphosis
Metamorphosis is a biological process by which an animal physically develops including birth transformation or hatching, involving a conspicuous and relatively abrupt change in the animal's body structure through cell growth and different ...
usually peaks from late July to early August. They are able to live for up to 15 years.
Conservation status
According to the IUCN Red List of Endangered Species, ''H. arborea'' is “listed as Least Concern in view of its wide distribution, presumed large population, and because it is unlikely to be declining fast enough to qualify for listing in a more threatened category.” However, according to the IUCN, the population trend of ''H. arborea'' is decreasing.
Some of the main threats to European tree frogs include habitat fragmentation
Habitat fragmentation describes the emergence of discontinuities (fragmentation) in an organism's preferred environment (habitat), causing population fragmentation and ecosystem decay. Causes of habitat fragmentation include geological proces ...
and destruction, pollution of wetlands, predation from fish, capture for the pet trade, and climate change
Present-day climate change includes both global warming—the ongoing increase in Global surface temperature, global average temperature—and its wider effects on Earth's climate system. Climate variability and change, Climate change in ...
. Besides these main threats, other possible reasons for the decline in their populations include increased UVB radiation
Ultraviolet radiation, also known as simply UV, is electromagnetic radiation of wavelengths of 10–400 nanometers, shorter than that of visible light, but longer than X-rays. UV radiation is present in sunlight and constitutes about 10% of th ...
and local and far-ranging pesticides, fertilizers, and pollutants. Trout have been observed preying on European tree frogs, and in Europe, trout introduced into a pond result in a significant decline in their population.
While ''H. arborea'' is sensitive to habitat fragmentation, habitat restoration
Ecological restoration, or ecosystem restoration, is the process of assisting the recovery of an ecosystem that has been degraded, damaged, destroyed or transformed. It is distinct from conservation in that it attempts to retroactively repair ...
(beginning in the 1980s) has been successful to increase populations. Besides habitat restoration, other attempts to increase population have included building of new breeding ponds, creation of “habitat corridors to connect breeding sites”, and reintroductions. This has been successful in Sweden, Latvia, and Denmark. Habitat protection has been shown to be the most important approach to conserving European tree frog populations.
Gallery
File:P7101931.JPG
File:Hyla arborea (Marek Szczepanek).jpg
File:Tree frog in Romania.JPG
File:Hyla arborea01.jpg
File:Laubfrosch-wiki.jpg
File:Romanian European Tree Frog.JPG
File:Hyla arborea var. molleri (Hello folks).jpg, Formerly included in the European tree frog, the form ''molleri'' is now recognized as its own species, the Iberian tree frog
File:HylaArboreaTadpole+Metam.jpg, ''Hyla arborea'' tadpole
A tadpole or polliwog (also spelled pollywog) is the Larva, larval stage in the biological life cycle of an amphibian. Most tadpoles are fully Aquatic animal, aquatic, though some species of amphibians have tadpoles that are terrestrial animal, ...
and metamorph
File:HylaArboreaLarva.JPG, Tadpole with hind legs
File:HylaArboreaSpawnHatching2.JPG, Newborn tadpoles hatching from frogspawn
File:Boomkikkers bij Witteveen - SoundCloud - luc de bruijn.ogg, European tree frogs in
References
{{Taxonbar, from=Q2142057
Hyla
Amphibians of Europe
Least concern biota of Europe
Articles containing video clips
Amphibians described in 1758
Taxa named by Carl Linnaeus
gl:Estroza