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The smooth snake (''Coronella austriaca'')Street D (1979). ''The Reptiles of Northern and Central Europe''. London: B.T. Batsford Ltd. 268 pp. . is a
species A species () is often defined as the largest group of organisms in which any two individuals of the appropriate sexes or mating types can produce fertile offspring, typically by sexual reproduction. It is the basic unit of Taxonomy (biology), ...
of non- venomous snake in the
family Family (from ) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). It forms the basis for social order. Ideally, families offer predictabili ...
Colubridae Colubridae (, commonly known as colubrids , from , 'snake') is a family of snakes. With 249 genera, it is the largest snake family. The earliest fossil species of the family date back to the Late Eocene epoch, with earlier origins suspected. C ...
. The species is found in northern and central Europe, but also as far east as northern Iran. The
Reptile Database The Reptile Database is a scientific database that collects taxonomic information on all living reptile species (i.e. no fossil species such as dinosaur Dinosaurs are a diverse group of reptiles of the clade Dinosauria. They first appeared ...
recognizes two
subspecies In Taxonomy (biology), biological classification, subspecies (: subspecies) is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (Morphology (biology), morpholog ...
as being valid, including the
nominotypical subspecies In biological classification, subspecies (: subspecies) is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics ( morphology), but that can successfully interbreed. ...
described here.


Description

Both sexes of ''C. austriaca'' grow to an average total length (including tail) of about to . Two specimens measuring have been recorded in Sweden, as well as one in Russia that was . The head has a rostral scale that is at least as deep as it is wide, creating a triangular indentation between the internasal scales (rarely separating them). The top of the head is covered with nine large plates. The nasal scale is often divided. There is one (rarely two) preoculars and two postoculars. The temporals number 2+2 or 2+3 (rarely 1+2). There are seven (rarely eight) upper labials, of which the third and fourth or fourth and fifth border the eye. In the middle part of the body there are 19 (rarely 17 or 21) rows of dorsal scales. In contrast with many other snakes found in the region, these scales are flat (not keeled). This gives the snake a smooth texture to the touch, from which it gets its common name. The ventral scales number 150–164 in males and 162–200 in females. The anal scale is divided (rarely single) and the subcaudal scales are paired. Males have 54–70 subcaudal scales and females 40–76. The colour pattern consists of a brown, grey or reddish ground colourSteward JW (1971). ''The Snakes of Europe''. Cranbury, New Jersey: Associated University Press (Fairleigh Dickinson University Press). 238 pp. LCCCN 77-163307. . with two rows of small, rather indistinct dark spots running down the back towards the tail. In some cases, each pair of spots may be united toward the neck area, forming a series of cross-bars over the back. There is also a very indistinct series of dark spots running along each of the flanks. These four series of spots along the body overlay four parallel, rather shadowy stripes that also run down the back and flanks. On the top of the head is a dark marking which is often in the shape of a crown, giving rise to the generic name ''Coronella'' (which means coronet). A relatively thick dark stripe extends from each nostril, through the eye, and along the side of the head to a little beyond the neck. The upper labials are whitish, greyish-white or light brown, sometimes with darker spots. The tongue is reddish brown or dark red.


Biology

The smooth snake feeds on smaller animals, especially other reptiles. It subdues larger prey by constriction, although unlike true constrictors it does not kill by this method. Smooth snakes are ovoviviparous: the juveniles hatch out of eggs internally and are born live. In Britain it is restricted to heathland habitats.


Geographic range

''Coronella austriaca'' is found from the south of England through France and the
Low Countries The Low Countries (; ), historically also known as the Netherlands (), is a coastal lowland region in Northwestern Europe forming the lower Drainage basin, basin of the Rhine–Meuse–Scheldt delta and consisting today of the three modern "Bene ...
to northern Spain and Portugal, Germany, Norway and Sweden (as far north as latitude 63°), Latvia, Lithuania, Estonia, Switzerland, Austria, Italy and
Sicily Sicily (Italian language, Italian and ), officially the Sicilian Region (), is an island in the central Mediterranean Sea, south of the Italian Peninsula in continental Europe and is one of the 20 regions of Italy, regions of Italy. With 4. ...
(but not in
Corsica Corsica ( , , ; ; ) is an island in the Mediterranean Sea and one of the Regions of France, 18 regions of France. It is the List of islands in the Mediterranean#By area, fourth-largest island in the Mediterranean and lies southeast of the Metro ...
or
Sardinia Sardinia ( ; ; ) is the Mediterranean islands#By area, second-largest island in the Mediterranean Sea, after Sicily, and one of the Regions of Italy, twenty regions of Italy. It is located west of the Italian Peninsula, north of Tunisia an ...
), the western
Balkans The Balkans ( , ), corresponding partially with the Balkan Peninsula, is a geographical area in southeastern Europe with various geographical and historical definitions. The region takes its name from the Balkan Mountains that stretch throug ...
and Greece, and European Russia as far north as latitude 57°. In Asia, it is found from Turkey to
Azerbaijan Azerbaijan, officially the Republic of Azerbaijan, is a Boundaries between the continents, transcontinental and landlocked country at the boundary of West Asia and Eastern Europe. It is a part of the South Caucasus region and is bounded by ...
,
Georgia Georgia most commonly refers to: * Georgia (country), a country in the South Caucasus * Georgia (U.S. state), a state in the southeastern United States Georgia may also refer to: People and fictional characters * Georgia (name), a list of pe ...
,
Armenia Armenia, officially the Republic of Armenia, is a landlocked country in the Armenian Highlands of West Asia. It is a part of the Caucasus region and is bordered by Turkey to the west, Georgia (country), Georgia to the north and Azerbaijan to ...
and northern Iran. It is absent in Denmark despite the species being found just south of the German border as well as in southern Sweden. The species is only known from Denmark from 6 specimens, all found between 1870 and 1914. In Finland, the species is found only in
Ã…land Ã…land ( , ; ) is an Federacy, autonomous and Demilitarized zone, demilitarised region of Finland. Receiving its autonomy by a 1920 decision of the League of Nations, it is the smallest region of Finland by both area () and population (30,54 ...
, and it is not common there.


Subspecies


See also

* List of reptiles of Europe


References


Further reading

* Arnold EN, Burton JA (1978). ''A Field Guide to the Reptiles and Amphibians of Britain and Europe''. (With 351 illustrations, 257 in colour by D.W. Ovenden). London: Collins. 272 pp. + Plates 1-40. . (''Coronella austriaca'', pp. 204–205 + Plate 38, figure 1 + Map 117). * Boulenger GA (1894). ''Catalogue of the Snakes in the British Museum (Natural History). Volume II., Containing the Conclusion of the Colubridæ Aglyphæ.'' London: Trustees of the British Museum (Natural History). (Taylor and Francis, printers). xi + 382 pp. + Plates I-XX. (''Coronella austriaca'', pp. 191–193). * Laurenti JN (1768). ''Specimen medicum, exhibens synopsin reptilium emendatam cum experimentis circa venena et antidota reptilium austriacorum.'' Vienna: "Joan. Thom. Nob. de Trattnern". 214 pp. + Plates I-V. (''Coronella austriaca'', new species, p. 84 + Plate V, figure 1). (in Latin). * Malkmus R (1995). "Coronella austriaca acutirostris ''subspec. nov. aus dem Nordwesten der Iberischen Halbinsel (Reptilia: Serpentes: Colubridae)'' ". ''Zoologische Abhandlungen, Museum für Tierkunde Dresden'' 48 (3): 265-278. (''Coronella austriaca acutirostris'', new subspecies). (in German).


Gallery

Image:Rézsikló (Coronella austriaca) feje.jpg, Head of ''C. austriaca''. Image:Rézsikló (Coronella austriaca)3.jpg, ''C. austriaca'' in
Hungary Hungary is a landlocked country in Central Europe. Spanning much of the Pannonian Basin, Carpathian Basin, it is bordered by Slovakia to the north, Ukraine to the northeast, Romania to the east and southeast, Serbia to the south, Croatia and ...
. File:Coronella austriaca s.jpg, ''C. austriaca'' in
Bosnia and Herzegovina Bosnia and Herzegovina, sometimes known as Bosnia-Herzegovina and informally as Bosnia, is a country in Southeast Europe. Situated on the Balkans, Balkan Peninsula, it borders Serbia to the east, Montenegro to the southeast, and Croatia to th ...
. Image:coronella_austriaca.jpg, ''C. austriaca'' curled up. Image:CoronellaAustriaca2.jpg, Subadult ''C. austriaca'' in northern
Germany Germany, officially the Federal Republic of Germany, is a country in Central Europe. It lies between the Baltic Sea and the North Sea to the north and the Alps to the south. Its sixteen States of Germany, constituent states have a total popu ...
.


External links


Smooth snake (''Coronella austriaca'')
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Surrey Amphibian and Reptile Group (SARG)
Accessed 5 June 2008.

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Amphibians and Reptiles of Europe
Accessed 7 October 2006.
Smooth snake (''Coronella austriaca'')
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ARKive
Accessed 7 October 2006.

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Reptiles and Amphibians of the UK
Accessed 7 October 2006.

at ttps://web.archive.org/web/20070808155828/http://www.reptilia-amphibia.net/ Reptiles & Amphibians of France Accessed 30 October 2006.
''Coronella austriaca''
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Accessed 30 March 2007.
Smooth Snake
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Blog posting with two pictures of ''Coronella austriaca''
Accessed 18 July 2007.
Smooth snake (''Coronella austriaca'')
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Dissertation abstract about smooth snakes mimicry of vipers being a protective behavior, scaring off some predators.
Accessed 15 July 2014
IUCN Red List
{{Authority control Coronella Snakes of Asia Reptiles of Europe Reptiles of Azerbaijan Reptiles of Russia Fauna of the United Kingdom Reptiles described in 1768 Taxa named by Josephus Nicolaus Laurenti