European Dipper
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The white-throated dipper (''Cinclus cinclus''), also known as the European dipper or just dipper, is an aquatic
passerine A passerine () is any bird of the order Passeriformes (; from Latin 'sparrow' and '-shaped') which includes more than half of all bird species. Sometimes known as perching birds, passerines generally have an anisodactyl arrangement of their ...
bird Birds are a group of warm-blooded vertebrates constituting the class (biology), class Aves (), characterised by feathers, toothless beaked jaws, the Oviparity, laying of Eggshell, hard-shelled eggs, a high Metabolism, metabolic rate, a fou ...
found in
Europe Europe is a continent located entirely in the Northern Hemisphere and mostly in the Eastern Hemisphere. It is bordered by the Arctic Ocean to the north, the Atlantic Ocean to the west, the Mediterranean Sea to the south, and Asia to the east ...
,
Middle East The Middle East (term originally coined in English language) is a geopolitical region encompassing the Arabian Peninsula, the Levant, Turkey, Egypt, Iran, and Iraq. The term came into widespread usage by the United Kingdom and western Eur ...
,
Central Asia Central Asia is a region of Asia consisting of Kazakhstan, Kyrgyzstan, Tajikistan, Turkmenistan, and Uzbekistan. The countries as a group are also colloquially referred to as the "-stans" as all have names ending with the Persian language, Pers ...
and the
Indian Subcontinent The Indian subcontinent is a physiographic region of Asia below the Himalayas which projects into the Indian Ocean between the Bay of Bengal to the east and the Arabian Sea to the west. It is now divided between Bangladesh, India, and Pakista ...
. The
species A species () is often defined as the largest group of organisms in which any two individuals of the appropriate sexes or mating types can produce fertile offspring, typically by sexual reproduction. It is the basic unit of Taxonomy (biology), ...
is divided into several
subspecies In Taxonomy (biology), biological classification, subspecies (: subspecies) is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (Morphology (biology), morpholog ...
, based primarily on colour differences, particularly of the pectoral band.


Taxonomy and systematics

The white-throated dipper was described in 1758 by the Swedish naturalist
Carl Linnaeus Carl Linnaeus (23 May 1707 – 10 January 1778), also known after ennoblement in 1761 as Carl von Linné,#Blunt, Blunt (2004), p. 171. was a Swedish biologist and physician who formalised binomial nomenclature, the modern system of naming o ...
in the tenth edition of his ''
Systema Naturae ' (originally in Latin written ' with the Orthographic ligature, ligature æ) is one of the major works of the Sweden, Swedish botanist, zoologist and physician Carl Linnaeus (1707–1778) and introduced the Linnaean taxonomy. Although the syste ...
'' under the
binomial name In taxonomy, binomial nomenclature ("two-term naming system"), also called binary nomenclature, is a formal system of naming species of living things by giving each a name composed of two parts, both of which use Latin grammatical forms, altho ...
''Sturnus cinclus''. The current genus ''
Cinclus Dippers are members of the genus ''Cinclus'' in the bird family Cinclidae, so-called because of their bobbing or dipping movements. They are unique among passerines for their ability to dive and swim underwater. Taxonomy The genus ''Cinclus'' ...
'' was introduced by the German naturalist
Moritz Balthasar Borkhausen Moritz Balthasar Borkhausen (3 December 1760, Giessen – 30 November 1806, Darmstadt) was a German naturalist and forester. He took part in the production of ' by Johann Conrad Susemihl. He received his education in Giessen, and in 1796 started ...
in 1797. The name ''cinclus'' is from the
Ancient Greek Ancient Greek (, ; ) includes the forms of the Greek language used in ancient Greece and the classical antiquity, ancient world from around 1500 BC to 300 BC. It is often roughly divided into the following periods: Mycenaean Greek (), Greek ...
word that was used to describe small tail-wagging birds that resided near water. Of the five species now placed in the genus, a molecular genetic study has shown that the white-throated dipper is most closely related to the other Eurasian species, the
brown dipper The brown dipper (''Cinclus pallasii''), also known as Pallas's dipper, Asian dipper or the Asiatic dipper, is an aquatic songbird found in the mountains of the east Palearctic. It is a thrush-like bird with a cocked tail. Its plumage is chocolat ...
(''Cinclus pallasii''). There are 14
subspecies In Taxonomy (biology), biological classification, subspecies (: subspecies) is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (Morphology (biology), morpholog ...
of which one is now extinct (with ): * ''C. c. hibernicus''
Hartert Ernst Johann Otto Hartert (29 October 1859 – 11 November 1933) was a widely published German people, German ornithologist. Life and career Hartert was born in the Free and Hanseatic City of Hamburg on 29 October 1859. In July 1891, he married ...
, 1910 – ''Irish dipper'' – Ireland and west Scotland * ''C. c. gularis'' ( Latham, 1801) – ''British dipper'' – Scotland (except west), north, central and west England and Wales * ''C. c. cinclus'' (
Linnaeus Carl Linnaeus (23 May 1707 – 10 January 1778), also known after ennoblement in 1761 as Carl von Linné,#Blunt, Blunt (2004), p. 171. was a Swedish biologist and physician who formalised binomial nomenclature, the modern system of naming o ...
, 1758) – ''Northern white-throated dipper'' – north Europe, west and central France and north Spain and Portugal, Corsica and Sardinia * ''C. c. aquaticus'' ( Bechstein, 1797) – ''Central European dipper'' – central and south Europe * ''C. c. olympicus'' Madarász, 1903 – Cyprus * ''C. c. minor'' Tristram, 1870 – northwest Africa * ''C. c. rufiventris'' Tristram, 1885 – west Syria and Lebanon * ''C. c. uralensis'' Serebrovski, 1927 –
Ural Mountains The Ural Mountains ( ),; , ; , or simply the Urals, are a mountain range in Eurasia that runs north–south mostly through Russia, from the coast of the Arctic Ocean to the river Ural (river), Ural and northwestern Kazakhstan.
* ''C. c. caucasicus'' Madarász, 1903 – Turkey to the Caucasus, north Iran and north Iraq * ''C. c. persicus'' Witherby, 1906 – southwest Iran * ''C. c. leucogaster'' Bonaparte, 1850 – south-central Russia, northwest China south to Afghanistan and north Pakistan * ''C. c. baicalensis'' Dresser, 1892 – south-central and southeast Siberia * ''C. c. cashmeriensis'' Gould, 1860 – west and central Himalayas * ''C. c. przewalskii'' Bianchi, 1905 – east Himalayas, south Tibet and west China


Description

The white-throated dipper is about long, rotund and short tailed. The head of the adult (''gularis'' and ''aquaticus'') is brown, the back slate-grey mottled with black, looking black from a distance, and the wings and tail are brown. The throat and upper breast are white, followed by a band of warm chestnut which merges into black on the belly and flanks. The bill is almost black, the legs and irides brown. ''C. c. cinclus'' has a black belly band. The young are greyish brown and have no chestnut band.


Voice

The male has a sweet
wren Wrens are a family, Troglodytidae, of small brown passerine birds. The family includes 96 species and is divided into 19 genera. All species are restricted to the New World except for the Eurasian wren that is widely distributed in the Old Worl ...
-like song. During courtship the male sings whilst he runs and postures, exhibiting his snowy breast, and when displaying he will take long and high flights, like those of the
common kingfisher The common kingfisher (''Alcedo atthis''), also known as the Eurasian kingfisher and river kingfisher, is a small kingfisher with seven subspecies recognized within its wide distribution across Eurasia and North Africa. It is resident in much of ...
, accompanied by sharp metallic calls ''clink, clink'', different from the normal ''zil''.


Behaviour and ecology

The white-throated dipper is closely associated with swiftly running rivers and streams or the lakes into which they fall. It often perches bobbing spasmodically with its short tail uplifted on the rocks round which the water swirls and tumbles. It acquired its name from these sudden dips, not from its diving habit, though it dives as well as walks into the water. It flies rapidly and straight, its short wings whirring swiftly and without pauses or glides, calling a shrill ''zil, zil, zil''. It will then either drop on the water and dive or plunge in with a small splash. From a perch it will walk into the water and deliberately submerge, but there is no truth in the assertion that it can defy the laws of
specific gravity Relative density, also called specific gravity, is a dimensionless quantity defined as the ratio of the density (mass of a unit volume) of a substance to the density of a given reference material. Specific gravity for solids and liquids is nea ...
and walk along the bottom. Undoubtedly when entering the water it grips with its strong feet, but the method of progression beneath the surface is by swimming, using the wings effectively for flying under water. It holds itself down by muscular exertion, with its head well down and its body oblique, its course beneath the surface often revealed by a line of rising bubbles. In this way it secures its food, usually aquatic
invertebrate Invertebrates are animals that neither develop nor retain a vertebral column (commonly known as a ''spine'' or ''backbone''), which evolved from the notochord. It is a paraphyletic grouping including all animals excluding the chordata, chordate s ...
s including caddis worms and other aquatic insect
larva A larva (; : larvae ) is a distinct juvenile form many animals undergo before metamorphosis into their next life stage. Animals with indirect development such as insects, some arachnids, amphibians, or cnidarians typically have a larval phase ...
e,
beetle Beetles are insects that form the Taxonomic rank, order Coleoptera (), in the superorder Holometabola. Their front pair of wings are hardened into wing-cases, elytra, distinguishing them from most other insects. The Coleoptera, with about 40 ...
s, '' Limnaea'', ''
Ancylus ''Ancylus'' is a genus of very small, freshwater, air-breathing limpets. They are aquatic pulmonate gastropod mollusks in the tribe Ancylini within the family Planorbidae, the ram's horn snails and their allies.Bouchet, P.; Rosenberg, G. (20 ...
'' and other freshwater
molluscs Mollusca is a phylum of protostome, protostomic invertebrate animals, whose members are known as molluscs or mollusks (). Around 76,000 extant taxon, extant species of molluscs are recognized, making it the second-largest animal phylum ...
, and also
fish A fish (: fish or fishes) is an aquatic animal, aquatic, Anamniotes, anamniotic, gill-bearing vertebrate animal with swimming fish fin, fins and craniate, a hard skull, but lacking limb (anatomy), limbs with digit (anatomy), digits. Fish can ...
and small
amphibian Amphibians are ectothermic, anamniote, anamniotic, tetrapod, four-limbed vertebrate animals that constitute the class (biology), class Amphibia. In its broadest sense, it is a paraphyletic group encompassing all Tetrapod, tetrapods, but excl ...
s. A favourite food is the small crustacean ''
Gammarus ''Gammarus'' is an amphipod crustacean genus in the family Gammaridae. It contains more than 200 described species, making it one of the most species-rich genera of crustaceans. Different species have different optimal conditions, particularly ...
'', an
amphipod Amphipoda () is an order of malacostracan crustaceans with no carapace and generally with laterally compressed bodies. Amphipods () range in size from and are mostly detritivores or scavengers. There are more than 10,700 amphipod species cur ...
shrimp. It also walks and runs on the banks and rocks seeking terrestrial invertebrates. Dippers may be preyed on by predatory fish such as
brown trout The brown trout (''Salmo trutta'') is a species of salmonid ray-finned fish and the most widely distributed species of the genus ''Salmo'', endemic to most of Europe, West Asia and parts of North Africa, and has been widely introduced globally ...
although only one case has been recorded for this species unlike in
American dipper The American dipper (''Cinclus mexicanus''), also known as a water ouzel, is a semiaquatic bird species native to western North America. Description It is a stocky grey bird with a head sometimes tinged with brown, and white feathers on the ey ...
. The winter habits of the dipper vary considerably and apparently individually. When the swift hill streams are frozen, it is forced to descend to the lowlands and even visit the coasts, but some will remain if there is any open water.


Breeding

The white-throated dippers first breed when they are one year old. They are monogamous and defend a territory. The nest is almost invariably built either very near or above water. It is often placed on a rocky ledge or in a cavity. Human-made structures such as bridges are also used. The nest consists of a dome-shaped structure made of moss, grass stems and leaves with a side entrance within which is an inner cup made of stems, rootlets and hair. Both sexes build the main larger structure, but the female builds the inner cup. The eggs are laid daily. The clutch can contain from 1-8 eggs but usually 4–5. The eggs are smooth and glossy white and are with a calculated weight of . They are incubated by the female beginning after the last or sometimes the penultimate egg has been laid. The male will bring food to the incubating female. The eggs hatch after around 16 days and then both parents feed the altricial and nidicolous nestlings. For the first 12–13 days they are brooded by the female. Both parents remove the faecal sacs for the first 9 days. The chicks fledge at around 22 days of age, but the parents continue to feed their young for another week but feeding can continue for 18 days. If the female has started a second clutch, then only the male parent feeds the fledglings. One or two broods are reared, usually in the same nest. When disturbed, the young that hardly feathered will at once drop into the water and dive. The maximum recorded age of a white-throated dipper from ring-recovery data is 10 years and 7 months for a bird ringed in Finland. Within the United Kingdom and Ireland the maximum age is 8 years and 9 months for a bird ringed and recovered in
County Laois County Laois ( ; ) is a county in Ireland. It is part of the Eastern and Midland Region and in the province of Leinster. It was known as Queen's County from 1556 to 1922. The modern county takes its name from Loígis, a medieval kingdom. Hist ...
, Ireland.


Dippers and humans

The first detailed description of the white-throated dipper, dating from c.1183, is that of
Gerald of Wales Gerald of Wales (; ; ; ) was a Cambro-Norman priest and historian. As a royal clerk to the king and two archbishops, he travelled widely and wrote extensively. He studied and taught in France and visited Rome several times, meeting the Pope. He ...
(''Giraldus Cambrensis)'', the twelfth-century cleric, historian and traveller, in his book ''
Topographia Hibernica ''Topographia Hibernica'' (Latin for ''Topography of Ireland''), also known as ''Topographia Hiberniae'', is an account of the landscape and people of Ireland written by Gerald of Wales around 1188, soon after the Norman invasion of Ireland. ...
'', an account of his travels through
Ireland Ireland (, ; ; Ulster Scots dialect, Ulster-Scots: ) is an island in the North Atlantic Ocean, in Northwestern Europe. Geopolitically, the island is divided between the Republic of Ireland (officially Names of the Irish state, named Irelan ...
in 1183–86. Gerald, a keen observer of wildlife, describes the dipper accurately, but with his notorious tendency to believe anything he was told, which so often detracts from the value of his work, states that it was an aberrant variety of the
common kingfisher The common kingfisher (''Alcedo atthis''), also known as the Eurasian kingfisher and river kingfisher, is a small kingfisher with seven subspecies recognized within its wide distribution across Eurasia and North Africa. It is resident in much of ...
. The true kingfisher, according to Gerald, did not occur in Ireland in the 1180s, although it was widespread there by the eighteenth century. The white-throated dipper is
Norway Norway, officially the Kingdom of Norway, is a Nordic countries, Nordic country located on the Scandinavian Peninsula in Northern Europe. The remote Arctic island of Jan Mayen and the archipelago of Svalbard also form part of the Kingdom of ...
's
national bird This is a list of national birds, including official birds of overseas territories and other states described as nations. Most species in the list are officially designated. Some species hold only an "unofficial" status. The column is marked a ...
.


Gallery

File:Cinclus cinclus -Brandon Creek, County Kerry, Ireland-8.jpg, At Brandon Creek, County Kerry, Ireland File:White-throated Dipper from Tawang river, near Jung, Arunachal Pradesh India 26032019.jpg, From Jung town in
Arunachal Pradesh Arunachal Pradesh (; ) is a States and union territories of India, state in northeast India. It was formed from the North-East Frontier Agency (NEFA) region, and India declared it as a state on 20 February 1987. Itanagar is its capital and la ...
in eastern Himalayas
India India, officially the Republic of India, is a country in South Asia. It is the List of countries and dependencies by area, seventh-largest country by area; the List of countries by population (United Nations), most populous country since ...
File:Cinclus cinclus MWNH 1850.JPG, Eggs, Collection
Museum Wiesbaden The Museum Wiesbaden is a two-branch museum of Art museum, art and Natural history museum, natural history in the Hesse, Hessian capital of Wiesbaden, Germany. It is one of the three Hessian State museums, in addition to the museums in Hessian ...
,
Germany Germany, officially the Federal Republic of Germany, is a country in Central Europe. It lies between the Baltic Sea and the North Sea to the north and the Alps to the south. Its sixteen States of Germany, constituent states have a total popu ...
File:White-throated Dipper young one begging for food.jpg, White-throated Dipper young one begging for food, at Sumdo, Ladakh, India File:White-throated_Dipper_swimming.jpg, White-throated Dipper swimming in the river pišnica,
slovenia Slovenia, officially the Republic of Slovenia, is a country in Central Europe. It borders Italy to the west, Austria to the north, Hungary to the northeast, Croatia to the south and southeast, and a short (46.6 km) coastline within the Adriati ...


References


Sources

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External links

* *
White-throated Dipper videos, photos & sounds
on the Internet Bird Collection
Ageing and sexing by Javier Blasco-Zumeta & Gerd-Michael Heinze
{{Authority control
white-throated dipper The white-throated dipper (''Cinclus cinclus''), also known as the European dipper or just dipper, is an aquatic passerine bird found in Europe, Middle East, Central Asia and the Indian Subcontinent. The species is divided into several subspecies ...
Birds of Europe Birds of Central Asia
white-throated dipper The white-throated dipper (''Cinclus cinclus''), also known as the European dipper or just dipper, is an aquatic passerine bird found in Europe, Middle East, Central Asia and the Indian Subcontinent. The species is divided into several subspecies ...
Taxa named by Carl Linnaeus