The western marsh harrier (''Circus aeruginosus'') is a large
harrier, a
bird of prey
Birds of prey or predatory birds, also known as (although not the same as) raptors, are hypercarnivorous bird species that actively predation, hunt and feed on other vertebrates (mainly mammals, reptiles and smaller birds). In addition to speed ...
from
temperate
In geography, the temperate climates of Earth occur in the middle latitudes (approximately 23.5° to 66.5° N/S of the Equator), which span between the tropics and the polar regions of Earth. These zones generally have wider temperature ran ...
and
subtropical
The subtropical zones or subtropics are geographical zone, geographical and Köppen climate classification, climate zones immediately to the Northern Hemisphere, north and Southern Hemisphere, south of the tropics. Geographically part of the Ge ...
western
Eurasia
Eurasia ( , ) is a continental area on Earth, comprising all of Europe and Asia. According to some geographers, Physical geography, physiographically, Eurasia is a single supercontinent. The concept of Europe and Asia as distinct continents d ...
and adjacent
Africa
Africa is the world's second-largest and second-most populous continent after Asia. At about 30.3 million km2 (11.7 million square miles) including adjacent islands, it covers 20% of Earth's land area and 6% of its total surfac ...
. It is also known as the Eurasian marsh harrier. Formerly, a number of relatives were included in ''C. aeruginosus'', which was then known as "
marsh harrier
The marsh harriers are bird of prey, birds of prey of the harrier (bird), harrier subfamily. They are medium-sized Bird of prey, raptors and the largest and broadest-winged harriers. Most of them are associated with marshland and dense reedbeds. ...
". The related
taxa
In biology, a taxon (back-formation from ''taxonomy''; : taxa) is a group of one or more populations of an organism or organisms seen by taxonomists to form a unit. Although neither is required, a taxon is usually known by a particular name and ...
are now generally considered to be separate species: the
eastern marsh harrier (''C. spilonotus''), the
Papuan harrier
The Papuan harrier (''Circus spilothorax'') is a bird of prey native to New Guinea which belongs to the harrier genus ''Circus''. It was classified as a subspecies of the eastern marsh harrier (''Circus spilonotus'') of eastern Asia.
It is thi ...
(''C. spilothorax'') of eastern Asia and the
Wallacea
Wallacea is a biogeography, biogeographical designation for a group of mainly list of islands of Indonesia, Indonesian islands separated by deep-water straits from the Asian and Australia (continent), Australian continental shelf, continental ...
, the
swamp harrier
The swamp harrier (''Circus approximans''), also known as the Australasian marsh harrier or Australasian harrier, is a large, slim bird of prey widely distributed across Australasia. In New Zealand, it is also known by the Māori name . It arriv ...
(''C. approximans'') of
Australasia
Australasia is a subregion of Oceania, comprising Australia, New Zealand (overlapping with Polynesia), and sometimes including New Guinea and surrounding islands (overlapping with Melanesia). The term is used in a number of different context ...
and the
Madagascar marsh harrier (''C. maillardi'') of the western
Indian Ocean
The Indian Ocean is the third-largest of the world's five oceanic divisions, covering or approximately 20% of the water area of Earth#Surface, Earth's surface. It is bounded by Asia to the north, Africa to the west and Australia (continent), ...
islands.
The western marsh harrier is often divided into two
subspecies
In Taxonomy (biology), biological classification, subspecies (: subspecies) is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (Morphology (biology), morpholog ...
, the widely
migratory ''C. a. aeruginosus'' which is found across most of its range, and ''C. a. harterti'' which is resident all-year in north-west Africa.
Taxonomy
The western marsh harrier was
formally described by the Swedish naturalist
Carl Linnaeus
Carl Linnaeus (23 May 1707 – 10 January 1778), also known after ennoblement in 1761 as Carl von Linné,#Blunt, Blunt (2004), p. 171. was a Swedish biologist and physician who formalised binomial nomenclature, the modern system of naming o ...
in 1758 in the
tenth edition of his ''
Systema Naturae
' (originally in Latin written ' with the Orthographic ligature, ligature æ) is one of the major works of the Sweden, Swedish botanist, zoologist and physician Carl Linnaeus (1707–1778) and introduced the Linnaean taxonomy. Although the syste ...
'' under the
binomial name
In taxonomy, binomial nomenclature ("two-term naming system"), also called binary nomenclature, is a formal system of naming species of living things by giving each a name composed of two parts, both of which use Latin grammatical forms, altho ...
''Falco aeruginosus''. Linnaeus specified the
locality
Locality may refer to:
* Locality, a historical named location or place in Canada
* Locality (association), an association of community regeneration organizations in England
* Locality (linguistics)
* Locality (settlement)
* Suburbs and localitie ...
as Europe but restricted this to Sweden in 1761. The western marsh harrier is now placed in the
genus
Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
''
Circus
A circus is a company of performers who put on diverse entertainment shows that may include clowns, acrobats, trained animals, trapeze acts, musicians, dancers, hoopers, tightrope walkers, jugglers, magicians, ventriloquists, and unicy ...
'' that was introduced by the French naturalist
Bernard Germain de Lacépède
Bernard-Germain-Étienne de La Ville-sur-Illon, comte de Lacépède or La Cépède (; 26 December 17566 October 1825) was a French natural history, naturalist and an active freemason. He is known for his contribution to the Comte de Buffon's g ...
in 1799.
The genus name ''Circus'' is derived from the
Ancient Greek
Ancient Greek (, ; ) includes the forms of the Greek language used in ancient Greece and the classical antiquity, ancient world from around 1500 BC to 300 BC. It is often roughly divided into the following periods: Mycenaean Greek (), Greek ...
''kirkos'', referring to a bird of prey named for its circling flight (''kirkos'', "circle"), probably the
hen harrier
The hen harrier (''Circus cyaneus'') is a bird of prey. It breeds in Palearctic, Eurasia. The term "hen harrier" refers to its former habit of preying on free-ranging fowl.
It bird migration, migrates to more southerly areas in winter. Eurasian ...
. The specific ''aeruginosus'' is
Latin
Latin ( or ) is a classical language belonging to the Italic languages, Italic branch of the Indo-European languages. Latin was originally spoken by the Latins (Italic tribe), Latins in Latium (now known as Lazio), the lower Tiber area aroun ...
for "rusty".
Two subspecies are recognised:
[
* ''C. a. aeruginosus'' (Linnaeus, 1758) – breeding in Europe and the Middle East to central Asia south Europe, Africa, south Asia; wintering in south Europe, Africa and south Asia
* ''C. a. harterti'' Zedlitz, 1914 – resident in northwest Africa
]
Description
The western marsh harrier is in length, has a wingspan of and a weight of in males and in females. It is a large, bulky harrier, larger than other European harriers, with fairly broad wings, and is sexually dimorphic
Sexual dimorphism is the condition where sexes of the same species exhibit different Morphology (biology), morphological characteristics, including characteristics not directly involved in reproduction. The condition occurs in most dioecy, di ...
. The male's plumage
Plumage () is a layer of feathers that covers a bird and the pattern, colour, and arrangement of those feathers. The pattern and colours of plumage differ between species and subspecies and may vary with age classes. Within species, there can b ...
is mostly a cryptic reddish-brown with lighter yellowish streaks, which are particularly prominent on the breast. The head and shoulders are mostly pale greyish-yellowish. The rectrices
Flight feathers (''Pennae volatus'') are the long, stiff, asymmetrically shaped, but symmetrically paired pennaceous feathers on the Bird wing, wings or tail of a bird; those on the wings are called remiges (), singular remex (), while those ...
and the secondary and tertiary remiges
Flight feathers (''Pennae volatus'') are the long, stiff, asymmetrically shaped, but symmetrically paired pennaceous feathers on the wings or tail of a bird; those on the wings are called remiges (), singular remex (), while those on the ta ...
are pure grey, the latter contrasting with the brown forewing and the black primary remiges at the wingtips. The upperside and underside of the wing look similar, though the brown is lighter on the underwing. Whether from the side or below, flying males appear characteristically three-colored brown-grey-black. The legs, feet, irides and the cere
The beak, bill, or Rostrum (anatomy), rostrum is an external anatomical structure found mostly in birds, but also in turtles, non-avian dinosaurs and a few mammals. A beak is used for pecking, wikt:grasp#Verb, grasping, and holding (in wikt:probe ...
of the black bill are yellow.
The female is almost entirely chocolate-brown. The top of the head, the throat and the shoulders are of a conspicuously lighter yellowish colour; this can be clearly delimited and very contrasting, or (particularly in worn plumage) be more washed-out, resembling the male's head colours. But the eye area of the female is always darker, making the light eye stand out, while the male's head is altogether not very contrastingly coloured and the female lacks the grey wing-patch and tail. Juveniles are similar to females, but usually have less yellow, particularly on the shoulders.
There is a rare hypermelanic morph with largely dark plumage. It is most often found in the east of the species' range. Juveniles of this morph may look entirely black in flight.
Distribution and habitat
This species has a wide breeding range from Europe and northwestern Africa to Central Asia and the northern parts of the Middle East. It breeds in almost every country of Europe but is absent from mountainous regions and subarctic
The subarctic zone is a region in the Northern Hemisphere immediately south of the true Arctic, north of hemiboreal regions and covering much of Alaska, Canada, Iceland, the north of Fennoscandia, Northwestern Russia, Siberia, and the Cair ...
Scandinavia
Scandinavia is a subregion#Europe, subregion of northern Europe, with strong historical, cultural, and linguistic ties between its constituent peoples. ''Scandinavia'' most commonly refers to Denmark, Norway, and Sweden. It can sometimes also ...
. It is rare but increasing in Great Britain
Great Britain is an island in the North Atlantic Ocean off the north-west coast of continental Europe, consisting of the countries England, Scotland, and Wales. With an area of , it is the largest of the British Isles, the List of European ...
where it has spread as far as eastern Scotland.[ In the Middle East there are populations in ]Turkey
Turkey, officially the Republic of Türkiye, is a country mainly located in Anatolia in West Asia, with a relatively small part called East Thrace in Southeast Europe. It borders the Black Sea to the north; Georgia (country), Georgia, Armen ...
, Iraq
Iraq, officially the Republic of Iraq, is a country in West Asia. It is bordered by Saudi Arabia to Iraq–Saudi Arabia border, the south, Turkey to Iraq–Turkey border, the north, Iran to Iran–Iraq border, the east, the Persian Gulf and ...
, and Iran
Iran, officially the Islamic Republic of Iran (IRI) and also known as Persia, is a country in West Asia. It borders Iraq to the west, Turkey, Azerbaijan, and Armenia to the northwest, the Caspian Sea to the north, Turkmenistan to the nort ...
, while in Central Asia the range extends eastwards as far as north-west China
China, officially the People's Republic of China (PRC), is a country in East Asia. With population of China, a population exceeding 1.4 billion, it is the list of countries by population (United Nations), second-most populous country after ...
, Mongolia
Mongolia is a landlocked country in East Asia, bordered by Russia to the north and China to the south and southeast. It covers an area of , with a population of 3.5 million, making it the world's List of countries and dependencies by po ...
, and the Lake Baikal
Lake Baikal is a rift lake and the deepest lake in the world. It is situated in southern Siberia, Russia between the Federal subjects of Russia, federal subjects of Irkutsk Oblast, Irkutsk Oblasts of Russia, Oblast to the northwest and the Repu ...
region of Siberia
Siberia ( ; , ) is an extensive geographical region comprising all of North Asia, from the Ural Mountains in the west to the Pacific Ocean in the east. It has formed a part of the sovereign territory of Russia and its predecessor states ...
.
Most populations of the western marsh harrier are migratory or dispersive. Some birds winter in milder regions of southern and western Europe, while others migrate to the Sahel
The Sahel region (; ), or Sahelian acacia savanna, is a Biogeography, biogeographical region in Africa. It is the Ecotone, transition zone between the more humid Sudanian savannas to its south and the drier Sahara to the north. The Sahel has a ...
, Nile
The Nile (also known as the Nile River or River Nile) is a major north-flowing river in northeastern Africa. It flows into the Mediterranean Sea. The Nile is the longest river in Africa. It has historically been considered the List of river sy ...
basin and Great Lakes
The Great Lakes, also called the Great Lakes of North America, are a series of large interconnected freshwater lakes spanning the Canada–United States border. The five lakes are Lake Superior, Superior, Lake Michigan, Michigan, Lake Huron, H ...
region in Africa
Africa is the world's second-largest and second-most populous continent after Asia. At about 30.3 million km2 (11.7 million square miles) including adjacent islands, it covers 20% of Earth's land area and 6% of its total surfac ...
, or to Arabia
The Arabian Peninsula (, , or , , ) or Arabia, is a peninsula in West Asia, situated north-east of Africa on the Arabian plate. At , comparable in size to India, the Arabian Peninsula is the largest peninsula in the world.
Geographically, the ...
, the Indian subcontinent
The Indian subcontinent is a physiographic region of Asia below the Himalayas which projects into the Indian Ocean between the Bay of Bengal to the east and the Arabian Sea to the west. It is now divided between Bangladesh, India, and Pakista ...
, and Myanmar
Myanmar, officially the Republic of the Union of Myanmar; and also referred to as Burma (the official English name until 1989), is a country in northwest Southeast Asia. It is the largest country by area in Mainland Southeast Asia and has ...
. The all-year resident subspecies
In Taxonomy (biology), biological classification, subspecies (: subspecies) is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (Morphology (biology), morpholog ...
''harterti'' inhabits Morocco
Morocco, officially the Kingdom of Morocco, is a country in the Maghreb region of North Africa. It has coastlines on the Mediterranean Sea to the north and the Atlantic Ocean to the west, and has land borders with Algeria to Algeria–Morocc ...
, Algeria
Algeria, officially the People's Democratic Republic of Algeria, is a country in the Maghreb region of North Africa. It is bordered to Algeria–Tunisia border, the northeast by Tunisia; to Algeria–Libya border, the east by Libya; to Alger ...
, and Tunisia
Tunisia, officially the Republic of Tunisia, is a country in the Maghreb region of North Africa. It is bordered by Algeria to the west and southwest, Libya to the southeast, and the Mediterranean Sea to the north and east. Tunisia also shares m ...
.
Vagrants
Vagrancy is the condition of wandering homelessness without regular employment or income. Vagrants usually live in poverty and support themselves by travelling while engaging in begging, scavenging, or petty theft. In Western countries, ...
have reached Iceland
Iceland is a Nordic countries, Nordic island country between the Atlantic Ocean, North Atlantic and Arctic Oceans, on the Mid-Atlantic Ridge between North America and Europe. It is culturally and politically linked with Europe and is the regi ...
, the Azores
The Azores ( , , ; , ), officially the Autonomous Region of the Azores (), is one of the two autonomous regions of Portugal (along with Madeira). It is an archipelago composed of nine volcanic islands in the Macaronesia region of the North Atl ...
, Malaysia
Malaysia is a country in Southeast Asia. Featuring the Tanjung Piai, southernmost point of continental Eurasia, it is a federation, federal constitutional monarchy consisting of States and federal territories of Malaysia, 13 states and thre ...
, and Sumatra
Sumatra () is one of the Sunda Islands of western Indonesia. It is the largest island that is fully within Indonesian territory, as well as the list of islands by area, sixth-largest island in the world at 482,286.55 km2 (182,812 mi. ...
. The first documented (but unconfirmed) record for the Americas was one bird reportedly photographed on 4 December 1994 at Chincoteague National Wildlife Refuge
The Chincoteague National Wildlife Refuge is a wildlife preserve operated by the United States Fish and Wildlife Service, U.S. Fish & Wildlife Service. It is primarily located on the Virginia half of Assateague Island with portions (only about ...
in Accomack County, Virginia
Accomack County is a United States county that, together with Northampton County, constitutes the Eastern Shore region of the Commonwealth of Virginia. These two counties also form the southern portion of the Delmarva Peninsula, which is bo ...
. Subsequently, there were confirmed records from Guadeloupe
Guadeloupe is an Overseas departments and regions of France, overseas department and region of France in the Caribbean. It consists of six inhabited islands—Basse-Terre Island, Basse-Terre, Grande-Terre, Guadeloupe, Grande-Terre, Marie-Galant ...
(winter of 2002/2003), from Laguna Cartagena National Wildlife Refuge
The Laguna Cartagena National Wildlife Refuge ( Spanish: ''Refugio de Vida Silvestre de Laguna Cartagena'') is a 1043-acre National Wildlife Refuge in Lajas, Puerto Rico. It is part of the Caribbean Islands National Wildlife Refuge Complex.
...
in Puerto Rico
; abbreviated PR), officially the Commonwealth of Puerto Rico, is a Government of Puerto Rico, self-governing Caribbean Geography of Puerto Rico, archipelago and island organized as an Territories of the United States, unincorporated territo ...
(early 2004 and January/February 2006)[ and in ]Bermuda
Bermuda is a British Overseas Territories, British Overseas Territory in the Atlantic Ocean, North Atlantic Ocean. The closest land outside the territory is in the American state of North Carolina, about to the west-northwest.
Bermuda is an ...
(December 2015).[
]
Like the other marsh harriers, it is strongly associated with wetland
A wetland is a distinct semi-aquatic ecosystem whose groundcovers are flooded or saturated in water, either permanently, for years or decades, or only seasonally. Flooding results in oxygen-poor ( anoxic) processes taking place, especially ...
areas, especially those rich in common reed (''Phragmites australis''). It can also be met with in a variety of other open habitats, such as farmland
Agricultural land is typically land ''devoted to'' agriculture, the systematic and controlled use of other forms of lifeparticularly the rearing of livestock and production of cropsto produce food for humans. It is generally synonymous with bot ...
and grassland
A grassland is an area where the vegetation is dominance (ecology), dominated by grasses (Poaceae). However, sedge (Cyperaceae) and rush (Juncaceae) can also be found along with variable proportions of legumes such as clover, and other Herbaceo ...
, particularly where these border marshland. It is a territorial
A territory is an area of land, sea, or space, belonging or connected to a particular country, person, or animal.
In international politics, a territory is usually a geographic area which has not been granted the powers of self-government, ...
bird in the breeding season, and even in winter it seems less social than other harriers, which often gather in large flocks.[ But this is probably simply due to ]habitat
In ecology, habitat refers to the array of resources, biotic factors that are present in an area, such as to support the survival and reproduction of a particular species. A species' habitat can be seen as the physical manifestation of its ...
preferences, as the marsh harriers are completely allopatric
Allopatric speciation () – also referred to as geographic speciation, vicariant speciation, or its earlier name the dumbbell model – is a mode of speciation that occurs when biological populations become geographically isolated from ...
while several of ''C. aeruginosus'' grassland and steppe
In physical geography, a steppe () is an ecoregion characterized by grassland plains without closed forests except near rivers and lakes.
Steppe biomes may include:
* the montane grasslands and shrublands biome
* the tropical and subtropica ...
relatives winter in the same regions and assemble at food sources such as locust
Locusts (derived from the Latin ''locusta'', locust or lobster) are various species of short-horned grasshoppers in the family Acrididae that have a swarming phase. These insects are usually solitary, but under certain circumstances they b ...
outbreaks. Still, in Keoladeo National Park
Keoladeo National Park, or Keoladeo Ghana National Park, is a national park in Rajasthan, India. The national park hosts thousands of native, resident and migratory birds, especially during the winter season, when many different species fly to t ...
of Rajasthan
Rajasthan (; Literal translation, lit. 'Land of Kings') is a States and union territories of India, state in northwestern India. It covers or 10.4 per cent of India's total geographical area. It is the List of states and union territories of ...
(India
India, officially the Republic of India, is a country in South Asia. It is the List of countries and dependencies by area, seventh-largest country by area; the List of countries by population (United Nations), most populous country since ...
) around 100 Eurasian marsh harriers are observed to roost together each November/December; they assemble in tall grassland dominated by ''Desmostachya bipinnata
''Desmostachya bipinnata'', commonly known as halfa grass, big cordgrass, and salt reed-grass, is an Old World perennial grass, long known and used in human history. The grass is tall, tufted, leafy, perennial grass, branching from the base, erec ...
'' and vetiver
''Chrysopogon zizanioides'', commonly known as vetiver and khus, is a perennial bunchgrass of the family Poaceae.
Vetiver is most closely related to sorghum while sharing many morphological characteristics with other fragrant grasses, such as ...
(''Chrysopogon zizanioides''), but where this is too disturbed by human activity they will use floating carpets of common water hyacinth (''Eichhornia crassipes'') instead – the choice of such roost sites may be to give early warning of predators, which will conspicuously rustle through the plants if they try to sneak upon the resting birds[
]
Behaviour and ecology
Breeding
The start of the breeding season varies from mid-March to early May. Western marsh harrier males often pair with two and occasionally three females. Pair bond
In biology, a pair bond is the strong affinity that develops in some species between a mating pair, often leading to the production and rearing of young and potentially a lifelong bond. Pair-bonding is a term coined in the 1940s that is frequently ...
s usually last for a single breeding season, but some pairs remain together for several years.
The ground nest is made of sticks, reeds and grasses. It is usually built in a reedbed
A reedbed or reed bed is a natural habitat found in floodplains, waterlogged depressions and
estuaries. Reedbeds are part of a succession from young reeds colonising open water or wet ground through a gradation of increasingly dry ground. As ...
, but the species will also nest in arable fields. There are between three and eight eggs in a normal clutch
A clutch is a mechanical device that allows an output shaft to be disconnected from a rotating input shaft. The clutch's input shaft is typically attached to a motor, while the clutch's output shaft is connected to the mechanism that does th ...
. The eggs are oval in shape and white in colour, with a bluish or greenish tinge when recently laid. The eggs are incubated for 31–38 days and the young birds fledge
Fledging is the stage in a flying animal's life between egg, hatching or birth and becoming capable of flight.
This term is most frequently applied to birds, but is also used for bats. For altricial birds, those that spend more time in vulnera ...
after 30–40 days.
Food and feeding
It hunts in typical harrier fashion, gliding low over flat open ground on its search for prey, with its wings held in a shallow V-shape and often with dangling legs.
It feeds on small mammals, small birds, insects, reptiles, and frogs.[
]
Status and conservation
The western marsh harrier declined in many areas between the 19th and the late 20th centuries due to persecution, habitat destruction
Habitat destruction (also termed habitat loss or habitat reduction) occurs when a natural habitat is no longer able to support its native species. The organisms once living there have either moved elsewhere, or are dead, leading to a decrease ...
and excessive pesticide
Pesticides are substances that are used to control pests. They include herbicides, insecticides, nematicides, fungicides, and many others (see table). The most common of these are herbicides, which account for approximately 50% of all p ...
use. It is now a protected species in many countries. In Great Britain
Great Britain is an island in the North Atlantic Ocean off the north-west coast of continental Europe, consisting of the countries England, Scotland, and Wales. With an area of , it is the largest of the British Isles, the List of European ...
, the population was likely extinct by the end of the 19th century. A single pair in Horsey, Norfolk bred in 1911, but by 2006, the Rare Breeding Birds Panel had recorded at least 265 females rearing 453 young. It made a comeback in Ireland
Ireland (, ; ; Ulster Scots dialect, Ulster-Scots: ) is an island in the North Atlantic Ocean, in Northwestern Europe. Geopolitically, the island is divided between the Republic of Ireland (officially Names of the Irish state, named Irelan ...
as well, where it had become extinct in 1918.
It still faces a number of threats, including the shooting of birds migrating through the Mediterranean
The Mediterranean Sea ( ) is a sea connected to the Atlantic Ocean, surrounded by the Mediterranean basin and almost completely enclosed by land: on the east by the Levant in West Asia, on the north by Anatolia in West Asia and Southern ...
region. They are vulnerable to disturbance during the breeding season and also liable to lead shot
Shot is a collective term for small spheres or pellets, often made of lead. These have been projected from slings since ancient times and were the original projectiles for shotguns and are still fired primarily from shotguns and grenade launch ...
poisoning. Still, the threats to this bird have been largely averted and it is today classified as Species of Least Concern
A least-concern species is a species that has been evaluated and categorized by the International Union for Conservation of Nature (IUCN) as not being a focus of wildlife conservation because the specific species is still plentiful in the wil ...
by the IUCN
The International Union for Conservation of Nature (IUCN) is an international organization working in the field of nature conservation and sustainable use of natural resources. Founded in 1948, IUCN has become the global authority on the status ...
.[
]
References
Sources
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Further reading
*
*
*
External links
Marsh harrier at RSPB's Birds by Name
Western Marsh Harrier or European Marsh Harrier species text in The Atlas of Southern African Birds
a
Oiseaux.net
Ageing and sexing (PDF; 5.7 MB) by Javier Blasco-Zumeta & Gerd-Michael Heinze
*
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*
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{{Authority control
western marsh harrier
The western marsh harrier (''Circus aeruginosus'') is a large harrier (bird), harrier, a bird of prey from temperate and subtropical western Palearctic, Eurasia and adjacent Africa. It is also known as the Eurasian marsh harrier. Formerly, a numb ...
Birds of prey of Europe
Birds of Africa
western marsh harrier
The western marsh harrier (''Circus aeruginosus'') is a large harrier (bird), harrier, a bird of prey from temperate and subtropical western Palearctic, Eurasia and adjacent Africa. It is also known as the Eurasian marsh harrier. Formerly, a numb ...
western marsh harrier
The western marsh harrier (''Circus aeruginosus'') is a large harrier (bird), harrier, a bird of prey from temperate and subtropical western Palearctic, Eurasia and adjacent Africa. It is also known as the Eurasian marsh harrier. Formerly, a numb ...