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''Euphorbia celastroides'' is a
flowering plant Flowering plants are plants that bear flowers and fruits, and form the clade Angiospermae (). The term angiosperm is derived from the Ancient Greek, Greek words (; 'container, vessel') and (; 'seed'), meaning that the seeds are enclosed with ...
in the family
Euphorbiaceae Euphorbiaceae (), the spurge family, is a large family of flowering plants. In English, they are also commonly called euphorbias, which is also the name of Euphorbia, the type genus of the family. Most spurges, such as ''Euphorbia paralias'', ar ...
. It is referred to by the common name akoko by
Hawaiians Native Hawaiians (also known as Indigenous Hawaiians, Kānaka Maoli, Aboriginal Hawaiians, or simply Hawaiians; , , , and ) are the Indigenous Polynesian people of the Hawaiian Islands. Hawaiʻi was settled at least 800 years ago by Polynesians ...
, and is a species of
spurge ''Euphorbia'' is a large and diverse genus of flowering plants, commonly called spurge, in the family Euphorbiaceae. Euphorbias range from tiny annual plants to large and long-lived trees, with perhaps the tallest being '' Euphorbia ampliphyl ...
closely related to the
poinsettia The poinsettia (; ''Euphorbia pulcherrima'') is a commercially important flowering plant species of the diverse spurge family Euphorbiaceae. Indigenous to Mexico and Central America, the poinsettia was first described by Europeans in 1834. It ...
. This species develops into a round-shape shrub. This species is
endemic Endemism is the state of a species being found only in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also foun ...
to the
Hawaiian Islands The Hawaiian Islands () are an archipelago of eight major volcanic islands, several atolls, and numerous smaller islets in the Pacific Ocean, North Pacific Ocean, extending some from the Hawaii (island), island of Hawaii in the south to nort ...
.


Description

''Euphorbia celastroides'' grows as a medium-sized
shrub A shrub or bush is a small to medium-sized perennial woody plant. Unlike herbaceous plants, shrubs have persistent woody stems above the ground. Shrubs can be either deciduous or evergreen. They are distinguished from trees by their multiple ...
or small tree reaching in height.Hawaiian flora at botany.si.edu
/ref> To grow properly, this species requires temperatures of and light shade. This plant develops in a fashion similar to a
shrub A shrub or bush is a small to medium-sized perennial woody plant. Unlike herbaceous plants, shrubs have persistent woody stems above the ground. Shrubs can be either deciduous or evergreen. They are distinguished from trees by their multiple ...
. In the summer, it assumes a red-violet colouring. It does not lose its leaves in the winter, due to the warm climate of its range. Female flowers have a three-part
pistil Gynoecium (; ; : gynoecia) is most commonly used as a collective term for the parts of a flower that produce ovules and ultimately develop into the fruit and seeds. The gynoecium is the innermost whorl (botany), whorl of a flower; it consists ...
over a three-part
ovary The ovary () is a gonad in the female reproductive system that produces ova; when released, an ovum travels through the fallopian tube/ oviduct into the uterus. There is an ovary on the left and the right side of the body. The ovaries are end ...
, usually producing three (or sometimes more) seeds.''E. celastroides''
at ZipCodeZoo
This species is tolerant of heat and drought. They are susceptible to
fungal diseases Pathogenic fungi are fungi A fungus (: fungi , , , or ; or funguses) is any member of the group of eukaryotic organisms that includes microorganisms such as yeasts and mold (fungus), molds, as well as the more familiar mushrooms. These orga ...
. Its cyathia may be located in short or open-branched
cymes In botany, an inflorescence is a group or cluster of flowers arranged on a plant's stem that is composed of a main branch or a system of branches. An inflorescence is categorized on the basis of the arrangement of flowers on a main axis ( ped ...
, or remain ungrouped in leaf
axil A leaf (: leaves) is a principal appendage of the stem of a vascular plant, usually borne laterally above ground and specialized for photosynthesis. Leaves are collectively called foliage, as in "autumn foliage", while the leaves, stem, fl ...
s. The leaves are distichous (grow in two vertical rows) and may have a
glaucous ''Glaucous'' (, ) is used to describe the pale grey or bluish-green appearance of the surfaces of some plants, as well as in the names of birds, such as the glaucous gull (''Larus hyperboreus''), glaucous-winged gull (''Larus glaucescens''), ...
coating. This plant produces a green or brown, rounded fruit 2 to 4 mm long, containing grey-brown seeds 0.5 to 2.5 mm long.


Distribution and habitat

Most varieties of this species can only be found in the
Hawaiian Islands The Hawaiian Islands () are an archipelago of eight major volcanic islands, several atolls, and numerous smaller islets in the Pacific Ocean, North Pacific Ocean, extending some from the Hawaii (island), island of Hawaii in the south to nort ...
. ''E. celastroides'' is tolerant of drought and grows in dry areas, inland as well coastal. This species is endemic to the polihale and kanaio regions of
Kauai Kauai (), anglicized as Kauai ( or ), is one of the main Hawaiian Islands. It has an area of 562.3 square miles (1,456.4 km2), making it the fourth-largest of the islands and the 21st-largest island in the United States. Kauai lies 73 m ...
and
Maui Maui (; Hawaiian language, Hawaiian: ) is the second largest island in the Hawaiian archipelago, at 727.2 square miles (1,883 km2). It is the List of islands of the United States by area, 17th-largest in the United States. Maui is one of ...
.CHAMAESYCE CELASTROIDES COASTAL DRY SHRUBLAND
at hawaii.edu


Conservation

''Euphorbia celastroides'' has not yet been evaluated by the
IUCN The International Union for Conservation of Nature (IUCN) is an international organization working in the field of nature conservation and sustainable use of natural resources. Founded in 1948, IUCN has become the global authority on the status ...
. However, due to its endemic nature, it is very vulnerable to human threats. Two examples of such threats are four-wheeled vehicles (which crush the plant) and
introduced species An introduced species, alien species, exotic species, adventive species, immigrant species, foreign species, non-indigenous species, or non-native species is a species living outside its native distributional range, but which has arrived ther ...
(which compete for resources).


Varieties

This plant has many varieties. These varieties include:Euphorbia on SageBud
/ref> *Var. ''amplectens'' *Var. ''halawana'' *Var. ''hanapepensis'' *Var. ''haupuana'' *Var. ''humbertii'' *Var. ''ingrata'' *Var. ''kaenana'' *Var. ''kohalana'' *Var. ''laehiensis'' *Var. ''lorifolia'' *Var. ''mauiensis'' *Var. ''nelsonii'' *Var. ''nematopoda'' *Var. ''niuensis'' *Var. ''pseudoniuensis'' *Var. ''saxicola'' *Var. ''typica'' *Var. ''waikoluensis''


References

{{Taxonbar, from=Q5410407 celastroides Endemic flora of Hawaii Taxa named by Pierre Edmond Boissier