''Eumetabolodon'' is an

extinct Extinction is the termination of an organism by the death of its Endling, last member. A taxon may become Functional extinction, functionally extinct before the death of its last member if it loses the capacity to Reproduction, reproduce and ...

genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...

of procolophonine

procolophonid Procolophonidae is an extinct family of small, lizard-like parareptiles known from the Late Permian to Late Triassic that were distributed across Pangaea, having been reported from Europe, North America, China, South Africa, South America, Antarc ...

parareptile Parareptilia ("near-reptiles") is an extinct group of Basal (phylogenetics), basal Sauropsida, sauropsids ("Reptile, reptiles"), traditionally considered the sister taxon to Eureptilia (the group that likely contains all living reptiles and birds ...

from early and middle

Triassic The Triassic ( ; sometimes symbolized 🝈) is a geologic period and system which spans 50.5 million years from the end of the Permian Period 251.902 million years ago ( Mya), to the beginning of the Jurassic Period 201.4 Mya. The Triassic is t ...

(

Induan The Induan is the first age of the Early Triassic epoch in the geologic timescale, or the lowest stage of the Lower Triassic series in chronostratigraphy. It spans the time between 251.9 Ma and 249.9 Ma (million years ago). The Induan is so ...

to earliest

Anisian In the geologic timescale, the Anisian is the lower stage (stratigraphy), stage or earliest geologic age, age of the Middle Triassic series (stratigraphy), series or geologic epoch, epoch and lasted from million years ago until million years ag ...

stages) deposits of

Nei Mongol Inner Mongolia, officially the Inner Mongolia Autonomous Region, is an Autonomous regions of China, autonomous region of China. Its border includes two-thirds of the length of China's China–Mongolia border, border with the country of Mongolia. ...

, northern

China China, officially the People's Republic of China (PRC), is a country in East Asia. With population of China, a population exceeding 1.4 billion, it is the list of countries by population (United Nations), second-most populous country after ...

. Two species of ''Eumetabolodon'' were named by J. L. Li in

1983 1983 saw both the official beginning of the Internet and the first mobile cellular telephone call. Events January * January 1 – The migration of the ARPANET to TCP/IP is officially completed (this is considered to be the beginning of the ...

and the

type species In International_Code_of_Zoological_Nomenclature, zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the spe ...

is ''Eumetabolodon bathycephalus''.

Discovery

The type species is known from the

holotype A holotype (Latin: ''holotypus'') is a single physical example (or illustration) of an organism used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of s ...

IVPP The Institute of Vertebrate Paleontology and Paleoanthropology (IVPP; ) of China is a research institution and collections repository for fossils, including many dinosaur and pterosaur specimens (many from the Yixian Formation). As its name sugges ...

V6064, a nearly complete

skull The skull, or cranium, is typically a bony enclosure around the brain of a vertebrate. In some fish, and amphibians, the skull is of cartilage. The skull is at the head end of the vertebrate. In the human, the skull comprises two prominent ...

which was collected in the Zhuengeerqi locality, from the upper Heshanggou Formation (

Olenekian In the geologic timescale, the Olenekian is an age (geology), age in the Early Triassic epoch (geology), epoch; in chronostratigraphy, it is a stage (stratigraphy), stage in the Lower Triassic series (stratigraphy), series. It spans the time betw ...

and earliest

stages),

Ordos Basin The Ordos Plateau, also known as the Ordos Basin or simply the Ordos, is a highland sedimentary basin in parts of most Northern China with an elevation of , and consisting mostly of land enclosed by the Ordos Loop, a large northerly rectangular ...

. A second specimen, IVPP V6070 was collected in the Zhuengeerqi locality, from the

Ermaying Formation The Ermaying Formation is a geological formation of Anisian (Middle Triassic) age in north-central China. It is found across much of the Ordos Basin, at outcrops within the provinces of Shaanxi, Shanxi, and Inner Mongolia. It is composed of up to ...

(upper Olenekian stage), Ordos Basin. Additional 17 specimens, IVPP V6065-69, V6166 (1&2), V6167, V6168 (1&2), V6169-75, were collected in the Gucheng locality, from the upper Heshanggou Formation, Ordos Basin. ''"E." dongshengensis'' is known only from the holotype IVPP V6073, a partial skull including right frontal,

prefrontal Prefrontal may refer to: *Prefrontal bone, a skull bone in some tetrapods *Prefrontal cortex, a region of the brain of a mammal *Prefrontal scales The prefrontal scales on snakes and other reptiles are the scales adjacent and anterior to the fr ...

, lacrimal,

maxilla In vertebrates, the maxilla (: maxillae ) is the upper fixed (not fixed in Neopterygii) bone of the jaw formed from the fusion of two maxillary bones. In humans, the upper jaw includes the hard palate in the front of the mouth. The two maxil ...

with seven teeth and the right ramus of

lower jaw In jawed vertebrates, the mandible (from the Latin ''mandibula'', 'for chewing'), lower jaw, or jawbone is a bone that makes up the lowerand typically more mobilecomponent of the mouth (the upper jaw being known as the maxilla). The jawbone i ...

. It was collected in the Wusilangou locality near

Dongsheng Dongsheng District ( Mongolian: ; zh, s=东胜区; alternate spelling English: Koshang; Turkic: Košang) is a District and the seat of Ordos City, Inner Mongolia, China. It has a district population of 574,242. The district is predominantly Ha ...

(also in Nei Mongol), from the

Early Triassic The Early Triassic is the first of three epochs of the Triassic Period of the geologic timescale. It spans the time between 251.9 Ma and Ma (million years ago). Rocks from this epoch are collectively known as the Lower Triassic Series, which ...

Shihtienfeng Formation.

Phylogeny

Recent

cladistic Cladistics ( ; from Ancient Greek 'branch') is an approach to biological classification in which organisms are categorized in groups ("clades") based on hypotheses of most recent common ancestry. The evidence for hypothesized relationships is ...

analyses suggested that ''Eumetabolodon'' is a

paraphyletic Paraphyly is a taxonomic term describing a grouping that consists of the grouping's last common ancestor and some but not all of its descendant lineages. The grouping is said to be paraphyletic ''with respect to'' the excluded subgroups. In co ...

genus; the type species represents a relatively derived procolophonine, whereas ''"E." dongshengensis'' is a more basal procolophonid of the subfamily Theledectinae. In 2020 a separate genus ''

Youngetta ''Youngetta'' is an extinct genus of procolophonid from the Early Triassic of China China, officially the People's Republic of China (PRC), is a country in East Asia. With population of China, a population exceeding 1.4 billion, it is ...

'' was named. The

cladogram A cladogram (from Greek language, Greek ''clados'' "branch" and ''gramma'' "character") is a diagram used in cladistics to show relations among organisms. A cladogram is not, however, an Phylogenetic tree, evolutionary tree because it does not s ...

below follows the topology from a 2012 analysis by Cisneros.

References

{{Taxonbar, from=Q5409570 Procolophoninae Early Triassic reptiles of Asia Fossil taxa described in 1983 Middle Triassic reptiles of Asia Anisian life Prehistoric reptile genera