Eulimidae is a
family
Family (from ) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). It forms the basis for social order. Ideally, families offer predictabili ...
of very small
parasitic
Parasitism is a close relationship between species, where one organism, the parasite, lives (at least some of the time) on or inside another organism, the host, causing it some harm, and is adapted structurally to this way of life. The ent ...
sea snail
Sea snails are slow-moving marine (ocean), marine gastropod Mollusca, molluscs, usually with visible external shells, such as whelk or abalone. They share the Taxonomic classification, taxonomic class Gastropoda with slugs, which are distinguishe ...
s,
marine gastropod
Gastropods (), commonly known as slugs and snails, belong to a large Taxonomy (biology), taxonomic class of invertebrates within the phylum Mollusca called Gastropoda ().
This class comprises snails and slugs from saltwater, freshwater, and fro ...
mollusk
Mollusca is a phylum of protostomic invertebrate animals, whose members are known as molluscs or mollusks (). Around 76,000 extant species of molluscs are recognized, making it the second-largest animal phylum after Arthropoda. The ...
s in the superfamily
Vanikoroidea
Vanikoroidea is a superfamily of sea snails, marine gastropod molluscs in the clade Littorinimorpha. The superfamily Eulimoidea is a synonym of Vanikoroidea.
Families
Families within the superfamily Vanikoroidea include:
*Family Eulimidae Ph ...
.
Description
These small parasitic snails live on (or in some cases in) the bodies of
echinoderms
An echinoderm () is any animal of the phylum Echinodermata (), which includes starfish, brittle stars, sea urchins, sand dollars and sea cucumbers, as well as the sessile sea lilies or "stone lilies". While bilaterally symmetrical as larv ...
such as sea cucumbers, sea urchins, sea stars, etc. All species lack a
radula
The radula (; : radulae or radulas) is an anatomical structure used by mollusks for feeding, sometimes compared to a tongue. It is a minutely toothed, chitinous ribbon, which is typically used for scraping or cutting food before the food enters ...
, in most cases possessing a
proboscis
A proboscis () is an elongated appendage from the head of an animal, either a vertebrate or an invertebrate. In invertebrates, the term usually refers to tubular arthropod mouthparts, mouthparts used for feeding and sucking. In vertebrates, a pr ...
which they extend into their host's
body cavity
A body cavity is any space or compartment, or potential space, in an animal body. Cavities accommodate organs and other structures; cavities as potential spaces contain fluid.
The two largest human body cavities are the ventral body cavity, a ...
in order to feed.
Genera
Genera within the family Eulimidae include:
* ''
Abyssoaclis'' Barros et al., 2003
* ''
Aclis'' S.L. Lovén, 1846
* ''
Acrochalix'' Bouchet & Warén, 1986
* ''
Amamibalcis'' Kuroda & Habe, 1950
* ''
Annulobalcis'' Habe, 1965
* ''
Apicalia''
A. Adams, 1862
* ''
Arcuella'' G. Nevill & H. Nevill, 1874
* ''
Asterolamia'' Warén, 1980
* ''
Asterophila'' Randall & Heath, 1912
* ''
Auriculigerina''
Dautzenberg, 1925
* ''
Austrorissopsis'' Grant-Mackie & Chapman-Smith, 1971
* ''
Awanuia''
Powell, 1927
* ''
Bacula''
H. Adams &
A. Adams, 1863
* ''
Batheulima'' F. Nordsieck, 1968
* ''
Bathycrinicola'' Bouchet & Warén, 1986
* ''
Bulimeulima'' Bouchet & Warén, 1986
* ''
Campylorhaphion'' Bouchet & Warén, 1986
* ''
Chileutomia'' Tate &
Cossman, 1898
* ''
Clypeastericola'' Warén, 1994
* ''
Concavibalcis'' Warén, 1980
* ''
Costaclis''
Bartsch, 1947
* ''
Crinolamia'' Bouchet & Warén, 1979
* ''
Crinophtheiros'' Bouchet & Warén, 1986
* ''
Curveulima''
Laseron, 1955
* ''
Cyclonidea'' C. F. Laseron, 1956
* ''
Diacolax'' Mandahl-Barth, 1946
* ''
Discaclis'' Moolenbeek & Warén, 1987
* ''
Echineulima'' Lützen & Nielsen, 1975
* ''
Echiuroidicola'' Warén, 1980
* ''
Enteroxenos''
Bonnevie, 1902
* ''
Entocolax'' Voigt, 1888
* ''
Entoconcha'' J. Muller, 1852
* ''
Ersilia''
Monterosato, 1872
* ''
Eulima''
Risso, 1826
* ''
Eulimacrostoma'' Leonardo Santos de Souza, Alexandre Dias Pimenta, 2019
Leonardo Santos de Souza, Alexandre Dias Pimenta; Eulimacrostoma gen. nov., a new genus of Eulimidae (Gastropoda, Caenogastropoda) with description of a new species and reevaluation of other western Atlantic species; Zoosystematics and Evolution 95(2) 405-415
/ref>
* '' Eulimetta'' Warén, 1992
* '' Eulimostraca'' Bartsch, 1917
* '' Eulitoma'' Laseron, 1955
* '' Fuscapex'' Warén, 1981
* '' Fusceulima'' Laseron, 1955
* '' Gasterosiphon'' Koehler & Vaney, 1905
* '' Goodingia'' Lützen, 1972
* '' Goriella'' Moolenbeek, 2008
* '' Goubinia'' Dautzenberg, 1923
* '' Haliella'' Monterosato, 1878
* '' Halielloides'' Bouchet & Warén, 1986
* '' Hebeulima'' Laseron, 1955
* '' Hemiaclis'' G. O. Sars, 1878
* '' Hemiliostraca'' Pilsbry, 1917
* '' Hoenselaaria'' Moolenbeek, 2009
* '' Hoplopteron'' P. Fischer, 1876
* '' Hypermastus'' Pilsbry, 1899
* '' Larochella'' A.W.B. Powell, 1927
* '' Margineulima'' Cossmann, 1888
* '' Megadenus'' Rosén, 1910
* '' Melanella'' Bowdich, 1822
* '' Menon'' Hedley, 1900
* '' Microeulima'' Waren, 1992
* '' Molpadicola'' Grusov, 1957
* '' Monogamus'' Lützen, 1976
* '' Mucronalia'' A. Adams, 1860
* '' Nanobalcis'' Waren & Mifsud, 1990
* '' Niso'' Risso, 1826
* '' Oceanida'' de Folin, 1870
* '' Ophieulima'' Warén & Sibuet, 1981
* '' Ophioarachnicola'' Warén, 1980
* '' Ophiolamia'' Warén & Carney, 1981
* '' Paedophoropus'' Ivanov, 1933
* '' Palisadia'' Laseron, 1956
* '' Paramegadenus'' Humphreys & Lützen, 1972
* '' Parastilbe'' Cossman, 1900
* '' Parvioris'' Warén, 1981
* '' Peasistilifer'' Warén, 1980
* '' Pelseneeria'' Koehler & Vaney, 1908
* '' Pictobalcis'' Laseron, 1955
* '' Pisolamia'' Bouchet & Lützen, 1976
* '' Polygireulima'' Sacco, 1892
* '' Prostilifer'' Warén, 1980
* '' Pseudosabinella'' McLean, 1995
* '' Pulicicochlea'' Ponder & Gooding, 1978
* '' Punctifera'' Warén, 1981
*'' Pyramidelloides'' Nevill 1884
* '' Rectilabrum'' Bouchet & Warén, 1986
* '' Robillardia'' E.A. Smith, 1889
* † '' Rostreulima'' Cossmann, 1913
* '' Ruapukea'' R.K. Dell, 1952
* '' Sabinella'' Monterosato, 1890
* '' Sanciaella'' Moolenbeek & Hoenselaar, 2010
* '' Scalaribalcis'' Warén, 1980
* '' Scalaronoba'' A.W.B. Powell, 1927
* '' Scalenostoma'' Deshayes, 1863
* '' Selma'' A. Adams, 1863
* † '' Semistylifer'' Cossmann, 1921
* '' Severnsia'' Geiger, 2016
* '' Sticteulima'' Laseron, 1955
* '' Stilapex'' Iredale, 1925
* '' Stilifer'' Broderip n Broderip & Sowerby 1832
* '' Subniso'' McLean, 2000
* '' Teretianax'' Iredale, 1918
* '' Thaleia'' Waren, 1979
* '' Thyca'' H. Adams & A. Adams, 1854
* '' Thyonicola'' Mandahl-Barth, 1941
* '' Trochostilifer'' Warén, 1980
* '' Tropiometricola'' Warén, 1981
* '' Turveria'' Berry, 1956
* '' Umbilibalcis'' Bouchet & Waren, 1986
* '' Vitreobalcis'' Warén, 1980
* '' Vitreolina'' Monterosato, 1884
;Genera brought into synonymy:
* ''Acicularia'' Monterosato, 1884: synonym of '' Polygireulima'' Sacco, 1892
* Athleenia Bartsch, 1946: synonym of '' Oceanida'' de Folin, 1870
* ''Balcis'' Leach, 1847: synonym of '' Melanella'' Bowdich, 1822
* ''Bessomia'' Berry, 1959: synonym of '' Thyca'' H. Adams & A. Adams, 1854
* '' Bonellia'' Deshayes, 1838: synonym of '' Niso'' Risso, 1826
* ''Chryseulima'' Laseron, 1955: synonym of '' Apicalia'' A. Adams, 1862
* ''Comenteroxenos'' Tikasingh, 1961: synonym of '' Enteroxenos'' Bonnevie, 1902
* ''Cuspeulima'' Laseron, 1955: synonym of '' Eulima'' Risso, 1826
* ''Entosiphon'' Koehler & Vaney, 1903: synonym of '' Gasterosiphon'' Koehler & Vaney, 1905
* ''Eulimaustra'' Laseron, 1955: synonym of '' Melanella'' Bowdich, 1822
* Eulimitra Laseron, 1955: synonym of '' Hemiliostraca'' Pilsbry, 1917
* ''Eulimoda'' Laseron, 1955: synonym of '' Sabinella'' Monterosato, 1890
* ''Granulithyca'' Habe, 1976: synonym of '' Thyca'' H. Adams & A. Adams, 1854
* ''Halliella'' Monterosato, 1878: synonym of '' Haliella'' Monterosato, 1878
* ''Helicosyrinx'' Baur, 1864: synonym of '' Entoconcha'' J. Müller, 1852
* ''Hersilia'' Monterosato, 1884: synonym of '' Ersilia'' Monterosato, 1872
* ''Hoplopteropsis'' de Morgan, 1916: synonym of '' Oceanida'' de Folin, 1870
* ''Hyperlia'' Pilsbry, 1918: synonym of '' Scalenostoma'' Deshayes, 1863
* ''Kiramodulus'' Kuroda, 1949: synonym of '' Thyca'' H. Adams & A. Adams, 1854
* ''Lambertia'' Souverbie, 1869: synonym of '' Stilapex'' Iredale, 1925
* ''Leiostraca'' H. Adams & A. Adams, 1853: synonym of '' Eulima'' Risso, 1826
* ''Lentigobalcis'' Habe, 1961: synonym of '' Sticteulima'' Laseron, 1955
* ''Luetzenia'' Rehder, 1980: synonym of '' Robillardia'' E.A. Smith, 1889
* ''Luetzenia'' Warén, 1980: synonym of '' Echineulima'' Lützen & Nielsen, 1975
* ''Neovolusia'' Emerson, 1965: synonym of '' Niso'' Risso, 1826
* ''Paedophorus'' Ivanov, 1933: synonym of '' Paedophoropus'' Ivanov, 1933
* ''Parastilifer'' A.V. Ivanov, 1952: synonym of '' Pelseneeria'' Koehler & Vaney, 1908
* ''Parenteroxenos'' A.V. Ivanov, 1945: synonym of '' Thyonicola'' Mandahl-Barth, 1941
* ''Rosenia'' Nierstrasz, 1913: synonym of '' Pelseneeria'' Koehler & Vaney, 1908
* ''Spiroclimax'' Mörch, 1875: synonym of '' Oceanida'' de Folin, 1870
* ''Stilimella'' Laseron, 1955: synonym of '' Scalenostoma'' Deshayes, 1863
* ''Strombiformis'' Da Costa, 1778: synonym of '' Eulima'' Risso, 1826
* ''Stylapex'': synonym of '' Stilapex'' Iredale, 1925
* ''Stylifer'' Cossmann, 1921: synonym of '' Stilifer'' Broderip n Broderip & Sowerby I 1832
* ''Stylina'' Fleming, 1828: synonym of '' Pelseneeria'' Koehler & Vaney, 1908
* ''Subeulima'' Souverbie, 1875: synonym of '' Bacula'' H. Adams & A. Adams, 1863
* ''Subularia'' Monterosato, 1884: synonym of '' Eulima'' Risso, 1826
* ''Teretianax'' Iredale, 1918: synonym of '' Pyramidelloides'' G. Nevill, 1885
* Subfamily Thycinae: synonym of Eulimidae
* ''Turtonia'' Rosén, 1910: synonym of '' Pelseneeria'' Koehler & Vaney, 1908
* ''Venustilifer'' Powell, 1939: synonym of '' Pelseneeria'' Koehler & Vaney, 1908
* ''Volusia'' A. Adams, 1851: synonym of '' Niso'' Risso, 1826
Image:Thyca ectoconcha sur face orale de L. multifora.jpg, '' Thyca ectoconcha'' parasiting a sea star '' Linckia multifora''.
Image:Pyramidelloides mirandus 01.jpg, '' Pyramidelloides mirandus''
Image:Eulimidae sp. sur le trivium d'un juvénile H. verrucosa.jpg, Eulimids on a juvenile sea cucumber '' Holothuria verrucosa''.
References
Further reading
* Tryon G. W. 1886. ''Manual of Conchology
Volume 8
Naticidae, Capyptraeidae, Turritellidae, Vermetidae, Caecidae, Eulimidae, Turbonillidae, Pyramidellidae''. 461 pp., 79 plates
page 258
plate 68
Bartsch, P. (1947). A monograph of the West Atlantic mollusks of the family Aclididae. Smithsonian Miscellaneous Collections. 106 (20): 1-29, 6 pls.
* Warrèn Anders, 1984, ''A generic revision of the family Eulimidae (Gastropoda, Prosobranchia)'', The Journal of Molluscan Studies, Supplement 13 1-96
* Vaught, K.C. (1989). ''A classification of the living Mollusca''. American Malacologists: Melbourne, FL (USA). . XII, 195 pp.
*
Takano, T.; Kano, Y. (2014). Molecular phylogenetic investigations of the relationships of the echinoderm-parasite family Eulimidae within Hypsogastropoda (Mollusca). Molecular Phylogenetics and Evolution. 79: 258-269. Note: States that Eulimidae are a member of superfamily Vanikoroidea, on the basis of sequence data. This placement implies synonymy of the superfamilies Eulimoidea and Vanikoroidea
External links
Marine Identification Portal: Eulimidae
*
Miocene Gastropods and Biostratigraphy of the Kern River Area, California; United States Geological Survey Professional Paper 642
{{Authority control
Vanikoroidea