Etheostoma Pottsii
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''Etheostoma'' is a
genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
of small
freshwater fish Freshwater fish are fish species that spend some or all of their lives in bodies of fresh water such as rivers, lakes, ponds and inland wetlands, where the salinity is less than 1.05%. These environments differ from marine habitats in many wa ...
in the
family Family (from ) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). It forms the basis for social order. Ideally, families offer predictabili ...
Percidae The Percidae are a family of ray-finned fish, part of the order Perciformes, which are found in fresh and brackish waters of the Northern Hemisphere. The majority are Nearctic, but there are also Palearctic species. The family contains nearly 250 ...
, and within the sub-family ''
Etheostomatinae Etheostomatinae is a species-rich subfamily of freshwater ray-finned fish, the members of which are commonly known as the darters. The subfamily is part of the family Percidae which also includes the perches, ruffes and pikeperches. The family ...
'', native to North America. Most are restricted to the
United States The United States of America (USA), also known as the United States (U.S.) or America, is a country primarily located in North America. It is a federal republic of 50 U.S. state, states and a federal capital district, Washington, D.C. The 48 ...
, but species are also found in
Canada Canada is a country in North America. Its Provinces and territories of Canada, ten provinces and three territories extend from the Atlantic Ocean to the Pacific Ocean and northward into the Arctic Ocean, making it the world's List of coun ...
and
Mexico Mexico, officially the United Mexican States, is a country in North America. It is the northernmost country in Latin America, and borders the United States to the north, and Guatemala and Belize to the southeast; while having maritime boundar ...
. They are commonly known as
darters The darters, anhingas, or snakebirds are mainly tropical waterbirds in the family Anhingidae, which contains a single genus, ''Anhinga''. There are four living species, three of which are very common and widespread while the fourth is rarer and c ...
, although the term "darter" is shared by several other genera. Many can produce alarm
pheromone A pheromone () is a secreted or excreted chemical factor that triggers a social response in members of the same species. Pheromones are chemicals capable of acting like hormones outside the body of the secreting individual, to affect the behavio ...
s that serve to warn nearby fish in case of an attack.Smith, R.J.F. (1992): Alarm signals in fishes. ''Reviews in Fish Biology and Fisheries, 2 (1): 33–63.''


Species

The 157 recognized species in this genus are: * '' Etheostoma acuticeps'' R. M. Bailey, 1959 (sharphead darter) * '' Etheostoma akatulo''
Layman In religious organizations, the laity () — individually a layperson, layman or laywoman — consists of all members who are not part of the clergy, usually including any non-ordained members of religious orders, e.g. a nun or a lay brother. ...
& Mayden, 2009
(bluemask darter) * '' Etheostoma artesiae'' ( O. P. Hay, 1881) (redspot darter) * '' Etheostoma asprigene'' ( S. A. Forbes, 1878) (mud darter) * '' Etheostoma atripinne'' (
D. S. Jordan David Starr Jordan (January 19, 1851 – September 19, 1931) was the founding president of Stanford University, serving from 1891 to 1913. He was an ichthyologist during his research career. Prior to serving as president of Stanford Universi ...
, 1877)
(Cumberland snubnose darter) * '' Etheostoma australe'' D. S. Jordan, 1889 (Conchos darter) * '' Etheostoma autumnale'' Mayden, 2010 (autumn darter) * '' Etheostoma baileyi''
Page Page most commonly refers to: * Page (paper), one side of a leaf of paper, as in a book Page, PAGE, pages, or paging may also refer to: Roles * Page (assistance occupation), a professional occupation * Page (servant), traditionally a young m ...
& Burr, 1982
(emerald darter) * '' Etheostoma barbouri'' Kuehne & J. W. Small, 1971 (teardrop darter) * '' Etheostoma barrenense'' Burr & Page, 1982 (splendid darter) * '' Etheostoma basilare'' Page, Hardman &
Near NEAR or Near may refer to: People * Thomas J. Near, US evolutionary ichthyologist * Near, a developer who created the higan emulator Science, mathematics, technology, biology, and medicine * National Emergency Alarm Repeater (NEAR), a form ...
, 2003
(corrugated darter) * ''Etheostoma bellator'' Royal Dallas Suttkus, Suttkus & R. M. Bailey, 1993 (warrior darter) * ''Etheostoma bellum'' Timothy Zorach, Zorach, 1968 (orangefin darter) * ''Etheostoma bison'' Patrick A. Ceas, Ceas & Page, 1997 (buffalo darter) * ''Etheostoma blennioides'' Constantine Samuel Rafinesque, Rafinesque, 1819 (greenside darter) * ''Etheostoma blennius'' Charles Henry Gilbert, C. H. Gilbert & Joseph Swain (academic), Swain, 1887 (blenny darter) * ''Etheostoma boschungi'' Benjamin R. Wall Jr., Wall & James David Williams, J. D. Williams, 1974 (slackwater darter) * ''Etheostoma brevirostrum'' Suttkus & David A. Etnier, Etnier, 1991 (holiday darter) * ''Etheostoma brevispinum'' (Robert Ervin Coker, Coker, 1926) (Carolina fantail darter) * ''Etheostoma burri'' Ceas & Page, 1997 (brook darter) * ''Etheostoma caeruleum'' David Humphreys Storer, D. H. Storer, 1845 (rainbow darter) * ''Etheostoma camurum'' (Edward Drinker Cope, Cope, 1870) (bluebreast darter) * ''Etheostoma cervus'' Steven L. Powers, Powers & Mayden, 2003 (Chickasaw darter) * ''Etheostoma chermocki'' Herbert T. Boschung, Boschung, Mayden & Joseph R. Tomelleri, Tomelleri, 1992 (vermilion darter) * ''Etheostoma chienense'' Page & Ceas, 1992 (relict darter) * ''Etheostoma chlorobranchium'' Zorach, 1972 (greenfin darter) * ''Etheostoma chlorosomum'' (O. P. Hay, 1881) (bluntnose darter) * ''Etheostoma chuckwachatte'' Mayden & Robert M. Wood, R. M. Wood, 1993 (lipstick darter) * ''Etheostoma cinereum'' D. H. Storer, 1845 (ashy darter) * ''Etheostoma clinton'' Mayden &
Layman In religious organizations, the laity () — individually a layperson, layman or laywoman — consists of all members who are not part of the clergy, usually including any non-ordained members of religious orders, e.g. a nun or a lay brother. ...
, 2012
(beaded darter) * ''Etheostoma collettei'' Ray S. Birdsong, Birdsong & Leslie William Knapp, L. W. Knapp, 1969 (creole darter) * ''Etheostoma collis'' (Carl Leavitt Hubbs, C. L. Hubbs & Mott Dwight Cannon, Cannon, 1935) (Carolina darter) * ''Etheostoma colorosum'' Suttkus & R. M. Bailey, 1993 (coastal darter) * ''Etheostoma coosae'' (Henry Weed Fowler, Fowler, 1945) (Coosa darter) * ''Etheostoma corona'' Page & Ceas, 1992 (crown darter) * ''Etheostoma cragini'' C. H. Gilbert, 1885 (Arkansas darter) * ''Etheostoma crossopterum'' Marvin Eugene Braasch, Braasch & Mayden, 1985 (fringed darter) * ''Etheostoma davisoni'' O. P. Hay, 1885 (Choctawhatchee darter) * ''Etheostoma denoncourti'' Jay Richard Stauffer Jr., Stauffer & Ellen S. van Snik Gray, van Snik, 1997 (golden darter) * ''Etheostoma derivativum'' Page, Hardman & Near, 2003 (stone darter) * ''Etheostoma ditrema'' John S. Ramsey, Ramsey & Suttkus, 1965 (coldwater darter) * ''Etheostoma douglasi'' R. M. Wood & Mayden, 1993 (Tuskaloosa darter) * ''Etheostoma duryi'' James Alexander Henshall, Henshall, 1889 (black darter) * ''Etheostoma edwini'' (Carl Leavitt Hubbs, C. L. Hubbs & Mott Dwight Cannon, Cannon, 1935) (brown darter) * ''Etheostoma erythrozonum'' John F. Switzer, Switzer & R. M. Wood, 2009 (Meramec saddled darter) * ''Etheostoma etnieri'' Raymond William Bouchard, Bouchard, 1977 (cherry darter) * ''Etheostoma etowahae'' R. M. Wood & Mayden, 1993 (Etowah darter) * ''Etheostoma euzonum'' (C. L. Hubbs & John D. Black, J. D. Black, 1940) (Arkansas saddled darter) * ''Etheostoma exile'' (Charles Frédéric Girard, Girard, 1859) (Iowa darter) * ''Etheostoma faulkneri'' Sterling & Warren, 2020 (Yoknapatawpha darter) * ''Etheostoma flabellare'' Rafinesque, 1819 (fantail darter) * ''Etheostoma flavum'' Etnier & R. M. Bailey, 1989 (saffron darter) * ''Etheostoma fonticola'' (D. S. Jordan & C. H. Gilbert, 1886) (fountain darter) * ''Etheostoma forbesi'' Page & Ceas, 1992 (barrens darter) * ''Etheostoma fragi'' Donald A. Distler, Distler, 1968 (strawberry darter) * ''Etheostoma fricksium'' Samuel Frederick Hildebrand, Hildebrand, 1923 (savannah darter) * ''Etheostoma fusiforme'' (Girard, 1854) (swamp darter) * ''Etheostoma gore'' Mayden &
Layman In religious organizations, the laity () — individually a layperson, layman or laywoman — consists of all members who are not part of the clergy, usually including any non-ordained members of religious orders, e.g. a nun or a lay brother. ...
, 2012
(Cumberland darter) * ''Etheostoma gracile'' (Girard, 1859) (slough darter) * ''Etheostoma grahami'' (Girard, 1859) (Rio Grande darter) * ''Etheostoma gutselli'' (Hildebrand, 1932) (Tuckasegee darter) * ''Etheostoma histrio'' D. S. Jordan & C. H. Gilbert, 1887 (harlequin darter) * ''Etheostoma hopkinsi'' (Fowler, 1945) (Christmas darter) * ''Etheostoma inscriptum'' (D. S. Jordan & Alembert Winthrop Brayton, Brayton, 1878) (turquoise darter) * ''Etheostoma jessiae'' (D. S. Jordan & Brayton, 1878) (blueside darter) * ''Etheostoma jimmycarter'' Mayden & Layman, 2012 (bluegrass darter) * ''Etheostoma jordani'' C. H. Gilbert, 1891 (greenbreast darter) * ''Etheostoma juliae'' Seth Eugene Meek, Meek, 1891 (yoke darter) * ''Etheostoma kanawhae'' (Edward Cowden Raney, Raney, 1941) (Kanawha darter) * ''Etheostoma kantuckeense'' Ceas & Page, 1997 (Highland Rim darter) * ''Etheostoma kennicotti'' (Frederic Ward Putnam, Putnam, 1863) (stripetail darter) * ''Etheostoma lachneri'' Suttkus & R. M. Bailey, 1994 (Tombigbee darter) * ''Etheostoma lawrencei'' Ceas & Burr, 2002 (headwater darter) * ''Etheostoma lemniscatum'' Rebecca E. Blanton Johansen, Blanton, 2008 (tuxedo darter) * ''Etheostoma lepidum'' (S. F. Baird & Girard, 1853) (greenthroat darter) * ''Etheostoma longimanum'' D. S. Jordan, 1888 (longfin darter) * ''Etheostoma lugoi'' Steven Mark Norris, S. M. Norris & Wendell L. Minckley, W. L. Minckley, 1997 (tufa darter) * ''Etheostoma luteovinctum'' C. H. Gilbert & Swain, 1887 (redband darter) * ''Etheostoma lynceum'' O. P. Hay, 1885 (brighteye darter) * ''Etheostoma maculatum'' Jared Potter Kirtland, Kirtland, 1840 (spotted darter) * ''Etheostoma mariae'' (Fowler, 1947) (pinewoods darter) * ''Etheostoma marmorpinnum'' Blanton & Robert E. Jenkins, R. E. Jenkins, 2008 (marbled darter) * ''Etheostoma maydeni'' Powers & Bernard R. Kuhajda, Kuhajda, 2012 (redlips darter) * ''Etheostoma microperca'' D. S. Jordan & C. H. Gilbert, 1888 (least darter) * ''Etheostoma mihileze'' Mayden, 2010 (sunburst darter) * ''Etheostoma moorei'' Raney & Suttkus, 1964 (yellowcheek darter) * ''Etheostoma nebra'' Near & Matthew R. Thomas, M. R. Thomas, 2015 (buck darter) Near, T.J. & Thomas, M.R. (2015): A New Barcheek Darter Species from Buck Creek (Cumberland River System), Kentucky (Percidae: Etheostomatinae: ''Catonotus'': ''Oopareia''). ''Bulletin of the Peabody Museum of Natural History, 56 (2): 127–146.'' * ''Etheostoma neopterum'' William Mike Howell, W. M. Howell & Guido Dingerkus, Dingerkus, 1978 (lollipop darter) * ''Etheostoma nianguae'' C. H. Gilbert & Seth Eugene Meek, Meek, 1887 (Niangua darter) * ''Etheostoma nigripinne'' Braasch & Mayden, 1985 (blackfin darter) * ''Etheostoma nigrum'' Rafinesque, 1820 (johnny darter) * ''Etheostoma nuchale'' William Mike Howell, W. M. Howell & Richard Dale Caldwell, R. D. Caldwell, 1965 (watercress darter) * ''Etheostoma obama'' Mayden & Layman, 2012 (spangled darter) * ''Etheostoma obeyense'' Philip Henry Kirsch, Kirsch, 1892 (barcheek darter) * ''Etheostoma occidentale'' Powers & Mayden, 2007 (West Rim darter) * ''Etheostoma okaloosae'' (Fowler, 1941) (Okaloosa darter) * ''Etheostoma olivaceum'' Marvin Eugene Braasch, Braasch & Page, 1979 (sooty darter) * ''Etheostoma olmstedi'' D. H. Storer, 1842 (tessellated darter) * ''Etheostoma oophylax'' Patrick A. Ceas, Ceas &
Page Page most commonly refers to: * Page (paper), one side of a leaf of paper, as in a book Page, PAGE, pages, or paging may also refer to: Roles * Page (assistance occupation), a professional occupation * Page (servant), traditionally a young m ...
, 1992
(guardian darter) * ''Etheostoma orientale'' Powers & Mayden, 2007 (East Rim darter) * ''Etheostoma osburni'' (Carl Leavitt Hubbs, C. L. Hubbs & Milton Bernhard Trautman, Trautman, 1932) (candy darter) * ''Etheostoma pallididorsum'' Donald A. Distler, Distler & Artie L. Metcalf, Metcalf, 1962 (paleback darter) * ''Etheostoma parvipinne'' C. H. Gilbert & Swain, 1887 (goldstripe darter) * ''Etheostoma percnurum'' Robert E. Jenkins, R. E. Jenkins, 1994 (duskytail darter) * ''Etheostoma perlongum'' C. L. Hubbs & Edward Cowden Raney, Raney, 1946) (Waccamaw darter) * ''Etheostoma phytophilum'' Henry L. Bart Jr., Bart & Michael S. Taylor, M. S. Taylor, 1999 (rush darter) * ''Etheostoma planasaxatile'' Powers & Mayden, 2007 (duck darter) * ''Etheostoma podostemone'' D. S. Jordan & O. P. Jenkins, 1889 (riverweed darter) * ''Etheostoma pottsii'' (Girard, 1859) (Chihuahua darter) * ''Etheostoma proeliare'' (O. P. Hay, 1881) (cypress darter) * ''Etheostoma pseudovulatum'' Page & Ceas, 1992 (egg-mimic darter) * ''Etheostoma punctulatum'' (Louis Agassiz, Agassiz, 1854) (stippled darter) * ''Etheostoma pyrrhogaster'' R. M. Bailey & Etnier, 1988 (firebelly darter) * ''Etheostoma radiosum'' (C. L. Hubbs & John D. Black (ichthyologist), J. D. Black, 1941) (orangebelly darter) * ''Etheostoma rafinesquei'' Burr & Page, 1982 (Kentucky darter) * ''Etheostoma ramseyi'' Suttkus & R. M. Bailey, 1994 (Alabama darter) * ''Etheostoma raneyi'' Suttkus & Bart, 1994 (Yazoo darter) * ''Etheostoma rubrum'' Raney & Suttkus, 1966 (bayou darter) * ''Etheostoma rufilineatum'' (Cope, 1870) (redline darter) * ''Etheostoma rupestre'' C. H. Gilbert & Swain, 1887 (rock darter) * ''Etheostoma sagitta'' (D. S. Jordan & Swain, 1883) (arrow darter) * ''Etheostoma saludae'' (C. L. Hubbs & Cannon, 1935) (saluda darter) * ''Etheostoma scotti'' Bruce Henry Bauer, Bauer, Etnier & Noel M. Burkhead, Burkhead, 1995 (Cherokee darter) * ''Etheostoma segrex'' Steven Mark Norris, S. M. Norris & Wendell L. Minckley, W. L. Minckley, 1997 (Río Salado darter) * ''Etheostoma sellare'' (Lewis Radcliffe, Radcliffe & William Welsh Welsh, W. W. Welsh, 1913) (Maryland darter) * ''Etheostoma sequatchiense'' Burr, 1979 (Sequatchie darter) * ''Etheostoma serrifer'' (C. L. Hubbs & Cannon, 1935) (sawcheek darter) * ''Etheostoma simoterum'' (Cope, 1868) (snubnose darter) * ''Etheostoma sitikuense'' Blanton, 2008 (Citico darter) * ''Etheostoma smithi'' Page & Braasch, 1976 (slabrock darter) * ''Etheostoma spectabile'' (Agassiz, 1854) (orangethroat darter) * ''Etheostoma spilotum'' C. H. Gilbert, 1887 (Kentucky arrow darter, Cumberland Plateau darter) * ''Etheostoma squamiceps'' D. S. Jordan, 1877 (spottail darter) * ''Etheostoma stigmaeum'' (D. S. Jordan, 1877) (speckled darter) * ''Etheostoma striatulum'' Page & Braasch, 1977 (striated darter) * ''Etheostoma susanae'' (D. S. Jordan & Swain, 1883) (Cumberland darter) * ''Etheostoma swaini'' (D. S. Jordan, 1884) (Gulf darter) * ''Etheostoma swannanoa'' D. S. Jordan & Barton Warren Evermann, Evermann, 1889 (Swannanoa darter) * ''Etheostoma tallapoosae'' Suttkus & Etnier, 1991 (Tallapoosa darter) * ''Etheostoma tecumsehi'' Ceas & Page, 1997 (Shawnee darter) * ''Etheostoma teddyroosevelt'' Mayden & Layman, 2012 (highland darter) * ''Etheostoma tennesseense'' Powers & Mayden, 2007 (Tennessee darter) * ''Etheostoma tetrazonum'' (C. L. Hubbs & J. D. Black, 1940) (Missouri saddled darter) * ''Etheostoma thalassinum'' (D. S. Jordan & Brayton, 1878) (seagreen darter) * ''Etheostoma thompsoni'' Suttkus, Bart & Etnier, 2012 (gumbo darter) * ''Etheostoma tippecanoe'' D. S. Jordan & Evermann, 1890 (Tippecanoe darter) * ''Etheostoma trisella'' R. M. Bailey & W. J. Richards, 1963 (trispot darter) * ''Etheostoma tuscumbia'' C. H. Gilbert & Swain, 1887 (Tuscumbia darter) * ''Etheostoma uniporum'' Donald A. Distler, Distler, 1968 (current darter) * ''Etheostoma variatum'' Kirtland, 1838 (variegated darter) * ''Etheostoma virgatum'' (D. S. Jordan, 1880) (striped darter) * ''Etheostoma vitreum'' (Cope, 1870) (glassy darter) * ''Etheostoma vulneratum'' (Cope, 1870) (wounded darter) * ''Etheostoma wapiti'' Etnier & James David Williams, J. D. Williams, 1989 (boulder darter) * ''Etheostoma whipplei'' (Girard, 1859) (redfin darter) * ''Etheostoma xanthovum'' Julia E. Wood, Richard C. Harrington, Zachariah D. Alley, Matthew R. Thomas, Jeffrey W. Simmons and Thomas J. Near. 2023Julia E. Wood, Richard C. Harrington, Zachariah D. Alley, Matthew R. Thomas, Jeffrey W. Simmons and Thomas J. Near. 2023. A New Species of Spottail Darter Endemic to the Clarks River in Kentucky and Tennessee (Percidae: Etheostomatinae: Etheostoma). Bulletin of the Peabody Museum of Natural History. 64(1); 11–37. DOI: 10.3374/014.064.0102 * ''Etheostoma zonale'' (Cope, 1868) (banded darter) * ''Etheostoma zonifer'' C. L. Hubbs & Cannon, 1935) (backwater darter) * ''Etheostoma zonistium'' R. M. Bailey & Etnier, 1988 (bandfin darter)


References

{{Authority control Etheostoma, Etheostomatinae Freshwater fish of North America, * Taxa named by Constantine Samuel Rafinesque Freshwater fish genera