The watercress darter (''Etheostoma nuchale'') is a species of freshwater

ray-finned fish Actinopterygii (; ), members of which are known as ray-finned fish or actinopterygians, is a class of bony fish that comprise over 50% of living vertebrate species. They are so called because of their lightly built fins made of webbings of sk ...

, a darter from the

subfamily In biological classification, a subfamily (Latin: ', plural ') is an auxiliary (intermediate) taxonomic rank, next below family but more inclusive than genus. Standard nomenclature rules end botanical subfamily names with "-oideae", and zo ...

Etheostomatinae Etheostomatinae is a species-rich subfamily of freshwater ray-finned fish, the members of which are commonly known as the darters. The subfamily is part of the family Percidae which also includes the perches, ruffes and pikeperches. The family ...

, part of the

family Family (from ) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). It forms the basis for social order. Ideally, families offer predictabili ...

Percidae The Percidae are a family of ray-finned fish, part of the order Perciformes, which are found in fresh and brackish waters of the Northern Hemisphere. The majority are Nearctic, but there are also Palearctic species. The family contains nearly 250 ...

, which also contains the

perch Perch is a common name for freshwater fish from the genus ''Perca'', which belongs to the family Percidae of the large order Perciformes. The name comes from , meaning the type species of this genus, the European perch (''P. fluviatilis'') ...

es, ruffes and pikeperches. It is

endemic Endemism is the state of a species being found only in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also foun ...

to the eastern

United States The United States of America (USA), also known as the United States (U.S.) or America, is a country primarily located in North America. It is a federal republic of 50 U.S. state, states and a federal capital district, Washington, D.C. The 48 ...

where it is only known from the

Black Warrior River The Black Warrior River is a waterway in west-central Alabama in the southeastern United States. The river rises in the extreme southern edges of the Appalachian Highlands and flows 178 miles (286 km) to the Tombigbee River, of which the ...

drainage basin A drainage basin is an area of land in which all flowing surface water converges to a single point, such as a river mouth, or flows into another body of water, such as a lake or ocean. A basin is separated from adjacent basins by a perimeter, ...

near

Birmingham, Alabama Birmingham ( ) is a city in the north central region of Alabama, United States. It is the county seat of Jefferson County, Alabama, Jefferson County. The population was 200,733 at the 2020 United States census, 2020 census, making it the List ...

Description

This species can reach a length of TL though most only reach a length of around .

Distribution and habitat

The watercress darter is only known to exist in four specific bodies of water, two in the Watercress Darter National Wildlife Refuge and two at the Seven Springs in the Powderly neighborhood of Birmingham. The Seven Springs populations were discovered in 2003. The species can be found in the

watercress Watercress or yellowcress (''Nasturtium officinale'') is a species of aquatic flowering plant in the cabbage family, Brassicaceae. Watercress is a rapidly growing perennial plant native to Eurasia. It is one of the oldest known leaf vegetabl ...

zone of springs where the water is slow-moving. They are usually found in dense mats of watercress or other aquatic vegetation, where they rest on the leaves and stems well above the stream bed. They feed on

snail A snail is a shelled gastropod. The name is most often applied to land snails, terrestrial molluscs, terrestrial pulmonate gastropod molluscs. However, the common name ''snail'' is also used for most of the members of the molluscan class Gas ...

crustacean Crustaceans (from Latin meaning: "those with shells" or "crusted ones") are invertebrate animals that constitute one group of arthropods that are traditionally a part of the subphylum Crustacea (), a large, diverse group of mainly aquatic arthrop ...

s, and

insect Insects (from Latin ') are Hexapoda, hexapod invertebrates of the class (biology), class Insecta. They are the largest group within the arthropod phylum. Insects have a chitinous exoskeleton, a three-part body (Insect morphology#Head, head, ...

larva A larva (; : larvae ) is a distinct juvenile form many animals undergo before metamorphosis into their next life stage. Animals with indirect development such as insects, some arachnids, amphibians, or cnidarians typically have a larval phase ...

e in slow-moving water.

Conservation and status

The species is endangered and is protected in areas such as the Watercress Darter National Wildlife Refuge in Jefferson County. The

IUCN The International Union for Conservation of Nature (IUCN) is an international organization working in the field of nature conservation and sustainable use of natural resources. Founded in 1948, IUCN has become the global authority on the status ...

lists it as being "

Endangered An endangered species is a species that is very likely to become extinct in the near future, either worldwide or in a particular political jurisdiction. Endangered species may be at risk due to factors such as habitat loss, poaching, inv ...

" due to its small range, small total population size and vulnerability to deterioration in the quality of the water in the springs in which it occurs. On September 22, 2008, more than 12,000 watercress darters were killed when workers from the city of Birmingham removed a

beaver Beavers (genus ''Castor'') are large, semiaquatic rodents of the Northern Hemisphere. There are two existing species: the North American beaver (''Castor canadensis'') and the Eurasian beaver (''C. fiber''). Beavers are the second-large ...

dam at Roebuck Springs, emptying a pond that was the darters' habitat. An alternative solution would have been to install a

flow device Flow may refer to: Science and technology * Fluid flow, the motion of a gas or liquid * Flow (geomorphology), a type of mass wasting or slope movement in geomorphology * Flow (mathematics), a group action of the real numbers on a set * Flow (psyc ...

in the dam to regulate the pond height. The pond was emptied because it often flooded a nearby

tennis court A tennis court is the venue where the sport of tennis is played. It is a firm rectangular surface with a low net stretched across the centre. The same surface can be used to play both Types of tennis match, doubles and singles matches. A variet ...

. The city emptied the beaver pond without the permission of the US Fish and Wildlife Service (FWS). The FWS has ordered the dam to be restored and the pond refilled in an effort to save the remaining darters. The

Alabama Department of Conservation and Natural Resources The Alabama Department of Conservation and Natural Resources (ADCNR) is the state agency responsible for the conservation and management of Alabama's natural resources including state parks, state lands, wildlife and aquatic resources. ADCNR al ...

is suing the City of Birmingham for destroying endangered species and their wetland habitat.

References

External links

Village Creek in Roebuck Springs, Alabama - Video by Glynn Wilson

Outdoor Alabama - Watercress Darter
* {{Taxonbar, from=Q3756681 Etheostoma Endemic fish of the United States Endemic fauna of Alabama Fish described in 1965 ESA endangered species