''Etayoa'' is an

ungulate Ungulates ( ) are members of the diverse clade Euungulata ("true ungulates"), which primarily consists of large mammals with Hoof, hooves. Once part of the clade "Ungulata" along with the clade Paenungulata, "Ungulata" has since been determined ...

of the family Carodniidae in the order

Xenungulata Xenungulata ("strange ungulates") is an order of extinct and primitive South American hoofed mammals that lived from the Late Paleocene to Early Eocene ( Itaboraian to Casamayoran in the SALMA classification). Fossils of the order are known f ...

that lived during the

Early Eocene In the geologic timescale the Ypresian is the oldest age (geology), age or lowest stage (stratigraphy), stratigraphic stage of the Eocene. It spans the time between , is preceded by the Thanetian Age (part of the Paleocene) and is followed by th ...

(~ 55 Ma) in northern

South America South America is a continent entirely in the Western Hemisphere and mostly in the Southern Hemisphere, with a considerably smaller portion in the Northern Hemisphere. It can also be described as the southern Subregion#Americas, subregion o ...

Etymology

The genus of the type species ''Etayoa bacatensis'' was named by palaeontologist Carlos Villarroel after Fernando Etayo Serna, who contributed extensively to the

paleontology Paleontology, also spelled as palaeontology or palæontology, is the scientific study of the life of the past, mainly but not exclusively through the study of fossils. Paleontologists use fossils as a means to classify organisms, measure ge ...

and

stratigraphy Stratigraphy is a branch of geology concerned with the study of rock layers (strata) and layering (stratification). It is primarily used in the study of sedimentary and layered volcanic rocks. Stratigraphy has three related subfields: lithost ...

research in Colombia. The species epithet ''bacatensis'' refers to

Bacatá Bacatá (Chibcha: or ) is the name given to the main settlement of the Muisca Confederation on the Bogotá savanna. It mostly refers to an area, rather than an individual village, although the name is also found in texts referring to the modern ...

,Villarroel, 1987 the name in Muysccubun for the main settlement of the southern

Muisca Confederation The Muisca Confederation was a loose confederation of different Muisca rulers (''zaques'', ''zipas'', ''iraca'', and ''tundama'') in the central Andes, Andean highlands of what is today Colombia before the Spanish conquest of the Americas, Spanis ...

; the name of which has been used for the current Colombian capital

Bogotá Bogotá (, also , , ), officially Bogotá, Distrito Capital, abbreviated Bogotá, D.C., and formerly known as Santa Fe de Bogotá (; ) during the Spanish Imperial period and between 1991 and 2000, is the capital city, capital and largest city ...

, founded in a different location than the original Bacatá.

Description

The

type species In International_Code_of_Zoological_Nomenclature, zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the spe ...

fossil A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserve ...

specimen consists of a partial

mandible In jawed vertebrates, the mandible (from the Latin ''mandibula'', 'for chewing'), lower jaw, or jawbone is a bone that makes up the lowerand typically more mobilecomponent of the mouth (the upper jaw being known as the maxilla). The jawbone i ...

with teeth, found in the

Bogotá Formation The Bogotá Formation (, E1-2b, Tpb, Pgb) is a geological formation of the Eastern Hills and Bogotá savanna on the Altiplano Cundiboyacense, Eastern Ranges of the Colombian Andes. The predominantly shale and siltstone formation, with sandstone ...

in the locality

Ciudad Bolívar Ciudad Bolívar (; Spanish for "Bolivar City"), formerly known as Angostura and St. Thomas de Guyana, is the capital of Venezuela's southeastern Bolívar, Venezuela, Bolívar State. It lies at the spot where the Orinoco River narrows to about ...

Colombia Colombia, officially the Republic of Colombia, is a country primarily located in South America with Insular region of Colombia, insular regions in North America. The Colombian mainland is bordered by the Caribbean Sea to the north, Venezuel ...

.1987
Description of ''Etayoa bacatensis''
-

Paleobiology Database The Paleobiology Database (PBDB) is an online resource for information on the distribution and classification of fossil animals, plants, and microorganisms. History The Paleobiology Database originated in the NCEAS-funded Phanerozoic Marine Pale ...

The estimated size of the ungulate is the size of a dog. 2011
Un xenungulado del Paleoceno de la sabana de Bogotá
- Paleontología en Colombia

Paleoclimate and environment

The finds of iguanians, including the fossil record of hoplocercines, and boine, caenophidian, and ungaliophiine snakes in the Bogotá Formation indicate a tropical forest environment, present just before the Early Eocene Climatic Optimum (EECO).Head et al., 2012 The abundant

paleosol In Earth science, geoscience, paleosol (''palaeosol'' in Great Britain and Australia) is an ancient soil that formed in the past. The definition of the term in geology and paleontology is slightly different from its use in soil science. In geo ...

s of the Bogotá Formation show an increase in chemical weathering across the Paleocene-Eocene (P-E) transition; the

Paleocene–Eocene Thermal Maximum The Paleocene–Eocene thermal maximum (PETM), alternatively ”Eocene thermal maximum 1 (ETM1)“ and formerly known as the "Initial Eocene" or “Late Paleocene thermal maximum", was a geologically brief time interval characterized by a ...

.Morón et al., 2013

References

Bibliography

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Etymology

Description

Paleoclimate and environment

References

Bibliography

Further reading