''Espadarana audax'' is a species of

frog A frog is any member of a diverse and largely semiaquatic group of short-bodied, tailless amphibian vertebrates composing the order (biology), order Anura (coming from the Ancient Greek , literally 'without tail'). Frog species with rough ski ...

in the family Centrolenidae. It is found on the Amazonian versant of

Andes The Andes ( ), Andes Mountains or Andean Mountain Range (; ) are the List of longest mountain chains on Earth, longest continental mountain range in the world, forming a continuous highland along the western edge of South America. The range ...

in northeast Peru ( San Martín Region), eastern

Ecuador Ecuador, officially the Republic of Ecuador, is a country in northwestern South America, bordered by Colombia on the north, Peru on the east and south, and the Pacific Ocean on the west. It also includes the Galápagos Province which contain ...

(

Napo Province Napo () is a province in Ecuador. Its capital is Tena. The province contains the Napo River. The province is low developed without much industrial presence. The thick rainforest is home to many natives that remain isolated by preference, descend ...

), and southern

Colombia Colombia, officially the Republic of Colombia, is a country primarily located in South America with Insular region of Colombia, insular regions in North America. The Colombian mainland is bordered by the Caribbean Sea to the north, Venezuel ...

( Cauca and

Putumayo Department Putumayo () is a departments of Colombia, department of Southern Colombia. It is in the south-west of the country, bordering Ecuador and Peru. Its capital is Mocoa. The word ''putumayo'' comes from the Quechua languages. The verb ''p'utuy'' ...

s).

Taxonomy

The generic placement of what was originally described as ''Centrolenella audax'' in 1973 has long been uncertain, but in 2014 it was placed in the genus ''Espadarana''. At the same time, ''Centrolene fernandoi'' was brought into

synonymy A synonym is a word, morpheme, or phrase that means precisely or nearly the same as another word, morpheme, or phrase in a given language. For example, in the English language, the words ''begin'', ''start'', ''commence'', and ''initiate'' are a ...

with ''Espadarana audax''.

Description

Adult males measure in snout–vent length. The snout is truncate or round in lateral view. The texture of

dorsal Dorsal (from Latin ''dorsum'' ‘back’) may refer to: * Dorsal (anatomy), an anatomical term of location referring to the back or upper side of an organism or parts of an organism * Dorsal, positioned on top of an aircraft's fuselage The fus ...

skin is shagreen and includes spinules and white warts. The toes have some webbing.

Habitat and conservation

The species' natural

habitat In ecology, habitat refers to the array of resources, biotic factors that are present in an area, such as to support the survival and reproduction of a particular species. A species' habitat can be seen as the physical manifestation of its ...

s are moist

montane forest Montane ecosystems are found on the slopes of mountains. The alpine climate in these regions strongly affects the ecosystem because temperatures lapse rate, fall as elevation increases, causing the ecosystem to stratify. This stratification is ...

s at elevations of

above sea level Height above mean sea level is a measure of a location's vertical distance (height, elevation or altitude) in reference to a vertical datum based on a historic mean sea level. In geodesy, it is formalized as orthometric height. The zero level ...

. Threats to this species, assessed at the time when ''Centrolene fernandoi'' was still recognized as a separate species, include habitat loss (deforestation from agriculture and logging) and

chytridiomycosis Chytridiomycosis ( ) is an infectious disease in amphibians, caused by the chytrid fungi '' Batrachochytrium dendrobatidis'' and '' Batrachochytrium salamandrivorans''. Chytridiomycosis has been linked to dramatic population declines or extinc ...

References

{{Taxonbar, from=Q28055346 Espadarana Amphibians of the Andes Amphibians of Colombia Amphibians of Ecuador Frogs of Peru Amphibians described in 1973 Taxa named by John Douglas Lynch Taxa named by William Edward Duellman Taxonomy articles created by Polbot