The Escucha Formation is a geological

formation Formation may refer to: Linguistics * Back-formation, the process of creating a new lexeme by removing or affixes * Word formation, the creation of a new word by adding affixes Mathematics and science * Cave formation or speleothem, a secondary ...

La Rioja La Rioja () is an autonomous community and province in Spain, in the north of the Iberian Peninsula. Its capital is Logroño. Other cities and towns in the province include Calahorra, Arnedo, Alfaro, Haro, Santo Domingo de la Calzada, a ...

and

Teruel Teruel () is a city in Aragon, located in eastern Spain, and is also the capital of Teruel Province. It has a population of 35,675 in 2014 making it the least populated provincial capital in the country. It is noted for its harsh climate, with ...

provinces of northeastern

Spain , image_flag = Bandera de España.svg , image_coat = Escudo de España (mazonado).svg , national_motto = '' Plus ultra'' ( Latin)(English: "Further Beyond") , national_anthem = (English: "Royal March") , ...

whose strata date back to the late

Aptian The Aptian is an age in the geologic timescale or a stage in the stratigraphic column. It is a subdivision of the Early or Lower Cretaceous Epoch or Series and encompasses the time from 121.4 ± 1.0 Ma to 113.0 ± 1.0 Ma (million years ag ...

to middle

Albian The Albian is both an age of the geologic timescale and a stage in the stratigraphic column. It is the youngest or uppermost subdivision of the Early/Lower Cretaceous Epoch/ Series. Its approximate time range is 113.0 ± 1.0 Ma to 100.5 ± ...

stages of the

Early Cretaceous The Early Cretaceous (geochronological name) or the Lower Cretaceous ( chronostratigraphic name), is the earlier or lower of the two major divisions of the Cretaceous. It is usually considered to stretch from 145 Ma to 100.5 Ma. Geology Pr ...

. Dinosaur remains are among the fossils that have been recovered from the formation.Weishampel, et al., 2004 The approximately thick formation underlies the

Utrillas Formation Utrillas is a municipality located in the province of Teruel, Aragon Aragon ( , ; Spanish and an, Aragón ; ca, Aragó ) is an autonomous community in Spain, coextensive with the medieval Kingdom of Aragon. In northeastern Spain, the Ar ...

and overlies Castrillo de la Reina, Benassal &

Oliete Formation Oliete is a municipality located in the province of Teruel, Aragon, Spain. According to the 2017 census (INE), the municipality has a population of 357 inhabitants. Attractions The municipality boasts many attractions. These can be divided in ...

s. The Escucha Formation comprises

siltstone Siltstone, also known as aleurolite, is a clastic sedimentary rock that is composed mostly of silt. It is a form of mudrock with a low clay mineral content, which can be distinguished from shale by its lack of fissility.Blatt ''et al.'' 1980, ...

s, mudstones,

sandstone Sandstone is a clastic sedimentary rock composed mainly of sand-sized (0.0625 to 2 mm) silicate grains. Sandstones comprise about 20–25% of all sedimentary rocks. Most sandstone is composed of quartz or feldspar (both silicates ...

coal Coal is a combustible black or brownish-black sedimentary rock, formed as stratum, rock strata called coal seams. Coal is mostly carbon with variable amounts of other Chemical element, elements, chiefly hydrogen, sulfur, oxygen, and nitrogen ...

s and

amber Amber is fossilized tree resin that has been appreciated for its color and natural beauty since Neolithic times. Much valued from antiquity to the present as a gemstone, amber is made into a variety of decorative objects."Amber" (2004). In M ...

, in which several fossil insects were found. The formation was deposited in a variety of continental to paralic (

deltaic A river delta is a landform shaped like a triangle, created by deposition of sediment that is carried by a river and enters slower-moving or stagnant water. This occurs where a river enters an ocean, sea, estuary, lake, reservoir, or (more rare ...

)

environments Environment most often refers to: __NOTOC__ * Natural environment, all living and non-living things occurring naturally * Biophysical environment, the physical and biological factors along with their chemical interactions that affect an organism or ...

.Peyrot et al., 2007

Fossil content

The Escucha Formation has provided the following fossils, among others:Escucha Formation
at

Fossilworks Fossilworks is a portal which provides query, download, and analysis tools to facilitate access to the Paleobiology Database, a large relational database assembled by hundreds of paleontologists from around the world. History Fossilworks was cre ...

.org

Dinosaurs

;Other dinosaurs * Allosauroidea indet. * Iguanodontia indet. * Titanosauriformes indet. ;Reptiles * '' Aragochersis lignitesta''Pérez García et al., 2020 * '' Toremys cassiopeia'' * '' Trachydermochelys sp.'' * Testudines indet. * '' Anteophthalmosuchus escuchae'' * Goniopholididae indet. * '' Hulkepholis plotos'' ;Fish * Chondrichthyes indet. * Osteichthyes indet.

Insects

;Other arthropods * '' Alavia neli'' * '' Archaelagonops alavensis'' * ''

Mesozygiella dunlopi ''Mesozygiella'' is an extinct genus of orb-weaving spider (family Araneidae), with one known species, ''Mesozygiella dunlopi'', dating from the Early Cretaceous, making it the earliest orb-weaver yet discovered. Two male specimens of the specie ...

'' * '' Alavesiaphis margaritae'' * '' Hispanocader lisae'' * '' Alavametra popovi'' * '' Iberovelia quisquilia'' * '' Glaesivelia pulcherrima'' * '' Gymnopollisthrips maior'' * '' G. minor'' * '' Manicapsocidus enigmaticus'' * '' Archaeatropos alavensis'' * '' Preempheria antiqua'' * '' Empheropsocus arilloi'' * '' E. margineglabrus'' * '' Morazatermes krishnai'' * '' Cantabritermes simplex'' * '' Autrigonoforceps iberica'' * '' Hispanelcana alavensis'' * '' H. arilloi'' * '' H. lopezvallei'' * '' Libanophron sugaar'' * '' Hippocoon basajauni'' * '' Burmaphron iratxoak'' * '' B. jentilak'' * '' B. sorginak'' * '' Tagsmiphron olentzero'' * '' Elasmophron mari'' * '' Ectenobythus iberiensis'' * '' Liztor pilosus'' * '' Cretepyris martini'' * '' Zophepyris alavaensis'' * '' Ampulicomorpha perialla'' * '' Microcostaphron parvus'' * '' Archephedrus stolamissus'' * '' Protorhyssalopsis perrichoti'' * '' Spathiopteryx alavarommopsis'' * '' Serphites lamiak'' * '' Aposerphites angustus'' * '' Archaeromma hispanicum'' * '' Galloromma alavaensis'' * '' Alavaromma orchamum'' * '' Tithonoscelio resinalis'' * '' Bruescelio platycephalus'' * '' Electroteleiopsis hebdomas'' * '' Alavascelio delvallei'' * '' Amissascelio temporarius'' * '' Perimoscelio tyrbastes'' * '' Perimoscelio confector'' * ''

Proterosceliopsis masneri ''Proterosceliopsis'' is an extinct genus of platygastroid parasitic wasp, known from the Mid-Cretaceous of Eurasia. The genus was first described in 2014 from the Albian amber of the Escucha Formation. In 2019 additional species were described ...

'' * '' Juxtascelio interitus'' * '' Iberopria perialla'' * ''

Iberoevania roblesi ''Iberoevania'' is an extinct genus of evaniid which existed in what is now Spain during the early Cretaceous period. It was named by Enrique Peñalver, Jaime Ortega-Blanco, André Nel and Xavier Delclòs in 2010 File:2010 Events Collage New ...

'' * ''

Cretevania alonsoi ''Cretevania'' is an extinct genus of Evaniidae, which lived in what is now China, Burma, England, Lebanon, Mongolia, Russia and Spain during the Cretaceous period. the genus was described by Rasnitsyn in 1975, and the type species is ''Creteva ...

'' * '' Iberomaimetsha nihtmara'' * '' Iberomaimetsha rasnitsyni'' * '' Valaa delclosi'' * '' Megalava truncata'' * '' Eosyntexis parva'' * '' Cretasonoma corinformibus'' * '' Penarhytus tenebris'' * ''

Prosolierius parvus ''Prosolierius'' is an extinct genus of rove beetle which existed in what is now Lebanon, Myanmar and Spain during the Barremian The Barremian is an age in the geologic timescale (or a chronostratigraphic stage) between 129.4 ± 1.5 Ma (mill ...

'' * '' Cretakarenni hispanicus'' * '' Rhizophtoma longus'' * '' Darwinylus marcosi'' * '' Mediumiuga sinespinis'' * '' Archiaustroconops alavensis'' * ''

Leptoconops (Leptoconops) zherikhini ''Leptoconops'' (black gnat) is a midge genus in the family Ceratopogonidae. It has a mostly tropical or subtropical distribution worldwide, but some species occur as far north as Moscow region in Russia and the Yukon Territory in Canada. ...

'' * '' Gerontodacus skalskii'' * '' Chimeromyia alava'' * '' Chimeromyina concilia'' * '' Euliphora grimaldii'' * '' Alavesia subiasi'' * '' Tethepomima holomma'' * '' Tethepomyia buruhandi'' * ''

Lysistrata emerita ''Lysistrata'' ( or ; Attic Greek: , ''Lysistrátē'', "Army Disbander") is an ancient Greek comedy by Aristophanes, originally performed in classical Athens in 411 BC. It is a comic account of a woman's extraordinary mission to end the Peloponne ...

'' * '' Hegalari minor'' * '' Hegalari antzinako'' * '' Alavamanota hispanica'' * '' Allocotocera xavieri'' * '' Cretohaplusia ortunoi'' * ''

Eltxo cretaceus ''Eltxo'' is a genus of wood midges in the family Cecidomyiidae. The single described species, ''Eltxo cretaceus'', is only known from amber inclusions from the Lower Cretaceous Lower may refer to: *Lower (surname) *Lower Township, New Jersey ...

'' * ''

Helius (Helius) alavensis In ancient Greek religion and mythology, Helios (; grc, , , Sun; Homeric Greek: ) is the god and personification of the Sun (Solar deity). His name is also Latinized as Helius, and he is often given the epithets Hyperion ("the one above ...

'' * ''

Helius (Helius) spiralis In ancient Greek religion and mythology, Helios (; grc, , , Sun; Homeric Greek: ) is the god and personification of the Sun (Solar deity). His name is also Latinized as Helius, and he is often given the epithets Hyperion ("the one above") ...

'' * '' Espanoderus barbarae'' * '' Alavaraphidia imperterrita'' * ''

Baissoptera cretaceoelectra ''Baissoptera'' is an extinct genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy ...

'' * '' Cretokatianna bucculenta'' * '' Sphyrotheciscus senectus'' * '' Archeallacma dolichopoda'' * '' Katiannasminthurus xenopygus'' * '' Pseudosminthurides stoechus'' * '' Burmisotoma spinulifera'' * '' Protoisotoma autrigoniensis'' * '' Proisotoma communis'' * '' Proleptochelia tenuissima'' * '' Electrotanais monolithus'' * '' Alavatanais carabe'' * '' Alavatanais margulisae'' * '' Eurotanais terminator'' * '' Spinomegops arcanus'' * '' Strieremaeus minguezae'' * '' Iberofoveopsis miguelesi'' * '' Hispanothrips utrillensis'' * '' Aragonitermes teruelensis'' * '' Aragonimantis aenigma'' * '' Mymaropsis turolensis'' * '' Serphites silban'' * '' Galloromma turolensis'' * '' Alavaromma orchamum'' * '' Cretevania rubusensis'' * '' Cretevania alcalai'' * ''

Cretevania montoyai ''Cretevania'' is an extinct genus of Evaniidae, which lived in what is now China, Burma, England, Lebanon, Mongolia, Russia and Spain during the Cretaceous period. the genus was described by Rasnitsyn in 1975, and the type species is ''Creteva ...

'' * ''

Actenobius magneoculus ''Actenobius magneoculus'' is an extinct species of anobiid beetle in the family Anobiidae. References Anobiidae Articles created by Qbugbot Beetles described in 2015 {{Paleo-beetle-stub ...

'' * ''

Arra legalovi Arra or ARRA may refer to: Places * Arra, Paschim Bardhaman, census town in West Bengal * Arra, Purba Bardhaman, West Bengal * Arra, Purulia, census town in West Bengal * Arra, Chakwal, a village in Pakistan * Arra, now part of Owney and Arra, Coun ...

'' * ''

'' * '' Atriculicoides sanjusti'' * '' Atriculicoides hispanicus'' * '' Gerontodacus skalskii'' * ''

Microphorites utrillensis ''Microphorites'' is an extinct genus of flies in the family Dolichopodidae. Species * †''Microphorites deploegi'' Nel, Perrichot, Daugeron & Néraudeau, 2004 Charentese amber, France, Cenomanian * †''Microphorites erikai'' Bramuzzo & Nel, ...

'' * ''

Litoleptis fossilis ''Litoleptis'' is a genus of snipe flies of the family Rhagionidae. Species *'' Litoleptis alaskensis'' Chillcott, 1963 - Alaska *'' Litoleptis araeostylus'' Greenwalt, 2019 - Montana *'' Litoleptis asterellaphile'' Imada & Kato, 2016 - Japan *' ...

'' * '' Burmazelmira grimaldii'' * '' Aragomantispa lacerata'' * '' Spinomegops aragonensis'' * '' Ametroproctus valeriae'' * '' Cretaceobodes martinezae'' * '' Trhypochthonius lopezvallei'' ;Invertebrates * Ostracoda indet. * Bivalvia indet. * Gastropoda indet. ;Flora *

Pinophyta Conifers are a group of cone-bearing seed plants, a subset of gymnosperms. Scientifically, they make up the division Pinophyta (), also known as Coniferophyta () or Coniferae. The division contains a single extant class, Pinopsida. All ex ...

* Angiospermae indet.

Correlation

References

Bibliography

* * * * * * {{ISBN, 0-520-24209-2

Fossil content

Dinosaurs

Insects

Correlation

See also

References

Bibliography

Further reading